Helina Robineau-Desvoidy, 1830

Sorokina, Vera S. & Pont, Adrian C., 2010, An annotated catalogue of the Muscidae (Diptera) of Siberia 2597, Zootaxa 2597 (1), pp. 1-87 : 34-40

Genus Helina Robineau-Desvoidy, 1830

Helina Robineau-Desvoidy, 1830: 493 . Type-species: Helina euphemioidea Robineau-Desvoidy, 1830 (= Anthomyia pertusa Meigen, 1826 ), by designation of Coquillett (1901: 137).

Helina annosa ( Zetterstedt, 1838)

Anthomyza annosa Zetterstedt, 1838: 663 . Type-locality: Sweden, Norrbotten, Johannis Ro. Lectotype ♀, MZLU (designated by Pont, in press).

Siberian records: south part of Tyumenskaya Oblast’, southern taiga ( Veselkin, 1966: 788); valley of River Yenisey, forest steppe: Krasnoyarskiy Kray: Minusinsk, Shushenskoe, Respublika Tyva: Kyzyl, Cherbi and taiga: Krasnoyarsk, Yeniseysk ( Sychevskaya, 1974: 129); West and East Siberia ( Pont, 1986a: 137).

General distribution: Holarctic Region, and extending into northern parts of the Oriental Region.

Helina arctata Collin, 1953

Helina arctata Collin, 1953: 173 . Type-localities: United Kingdom, England, Cambridgeshire (Chippenham Fen and Woodditton Wood), Suffolk (Barton Mills), Norfolk (South Walsham, and Caistor near Yarmouth), and Essex (Leigh). Syntypes 6♂ 4♀, OUMNH, and 1♂, BMNH ( Pont, 1995a: 35).

Siberian records: Novosibirskaya Oblast’, environs of Karasuk ( Sorokina, 2006a: 228).

General distribution: Palaearctic Region, from Europe east to West Siberia.

Helina bohemani ( Ringdahl, 1916)

Mydaea bohemani Ringdahl, 1916: 235 . Type-localities: Sweden, Jämtland, Åreskutan , and Lapland. Syntypes 1♂, NHRS, and 1♀, MZLU (all seen).

Siberian records: Novosibirskaya Oblast’, environs of Karasuk ( Sorokina, 2006a: 228); West Siberia ( Pont, 1986a: 138).

General distribution: Holarctic Region; in the Palaearctic, from northern Scandinavia, Russia (north-west European Russia and West Siberia) and Tajikistan.

Helina celsa ( Harris, 1780)

Musca celsa Harris, 1780: 125 , plate 36 fig. 53. Type-locality: not stated (South-east England). Type-series lost ( Pont & Michelsen, 1982: 30).

Musca quadrimaculata Fallén, 1823b: 63 . (Junior primary homonym of Musca quadrimaculata Swederus, 1787 , and Musca quadrimaculata Fabricius, 1787 .) Type-locality: Sweden, Skåne province. Lectotype ♂, NHRS (designated by Pont, 1984b: 292).

Siberian records: south part of Tyumenskaya Oblast’, forest steppe and subtaiga ( Veselkin, 1966: 784); West and East Siberia ( Pont, 1986a: 138).

General distribution: Palaearctic Region, from Europe east to China.

Helina cinerella (van der Wulp, 1867)

Aricia cinerella van der Wulp, 1867: 150 . Type-locality: not stated (from introduction: USA, Wisconsin). Holotype / syntypes ♂, RMNH.

Siberian records: Novosibirskaya Oblast’, environs of Karasuk, forest-steppe ( Sorokina, 2006a: 228, 2008: 163).

General distribution: Holarctic Region; in the Palaearctic, from Europe to the Russian Far East.

Helina daicles ( Walker, 1849)

Anthomyia daicles Walker, 1849: 959 . Type-localities: Sweden, Norrbotten, Haparanda, and Norway, Hammerfest and Söröya Island. Syntypes 1♂ 4♀, BMNH ( Stein, 1901: 194; seen).

Siberian records: West Siberia ( Pont, 1986a: 140).

General distribution: Northern Palaearctic Region, from Norway, Sweden, Finland and Russia (north-west European Russia and West Siberia).

Helina decipiens Mihályi, 1974

Helina decipiens Mihályi, 1974: 160 . Type-locality: Hungary, Budapest, Farkasvölgy. Holotype ♂, HNHM. Siberian records: East Siberia ( Pont, 1986a: 140).

General distribution: Palaearctic Region, from Europe east to East Siberia.

Helina depuncta (Fallén, 1825)

Musca depuncta Fallén, 1825a: 80 . Type-locality: not stated ( Sweden, probably Skåne province). Lectotype ♂, NHRS (designated by Pont, 1984b: 283).

Siberian records: Urals ( Hennig, 1957: 177; Zimin & El’berg, 1970: 532); south part of Tyumenskaya Oblast’, subtaiga ( Veselkin, 1966: 788); West Siberia ( Pont, 1986a: 141).

General distribution: Palaearctic Region, from Europe east to the Russian Far East.

Helina evecta ( Harris, 1780)

Musca evecta Harris, 1780: 125 , plate 36 fig. 55. Type-locality: not stated (South-east England). Type-series lost ( Pont & Michelsen, 1982: 33).

Mydina laetifica Robineau-Desvoidy, 1830: 500 . Type-locality: France, Paris. Holotype / syntypes ♂ ♀, formerly in Robineau-Desvoidy collection, now destroyed (not MNHN).

Siberian records: South Urals, Mt. Iremel’ ( Malozemov, 1989: 37); Novosibirskaya Oblast’, Karasukskiy Rayon, forest-steppe ( Sorokina, 2008: 163); near Biysk and Altay Mts ( Sychevskaya, 1978: 82); Respublika Buryatiya, near Sosnovka, Barguzin Reservation (Lavčiev, 1971: 216); valley of River Yenisey, forest steppe: Krasnoyarskiy Kray (Minusinsk, Shushenskoe), Respublika Tyva (Kyzyl, Cherbi) ( Sychevskaya, 1974: 129); West and East Siberia ( Pont, 1986a: 141).

General distribution: Holarctic, and also parts of the Afrotropical (Sokotra, Yemen), Oriental ( Pakistan, India, Sri Lanka) and Neotropical ( Mexico, Venezuela) Regions.

Helina flavisquama ( Zetterstedt, 1849)

Aricia flavisquama Zetterstedt, 1849: 3287 . Type-locality: Sweden, Lule Lappmark , Kvikkjokk. Holotype ♂, MZLU (Pont, in press).

Siberian records: Respublika Sakha [Yakutiya]: Yakutsk, Khomustakh nasleg, Tommot ( Sychevskaya, 1972: 147); valley of River Yenisey, Dudinka ( Sychevskaya, 1974: 131 – error for Helina fulvisquama ?); East Siberia ( Pont, 1986a: 142).

General distribution: Holarctic Region; in the Palaearctic, from Europe east to the Russian Far East.

Helina fulvisquama ( Zetterstedt, 1845)

Aricia fulvisquama Zetterstedt, 1845: 1491 . Type-locality: Sweden, Lule Lappmark , Tjamotis near Kvikkjokk. Lectotype ♂, NHRS (designated by Pont, in press).

Phaonia (Euphemia) arctica Schnabl in Becker et al., 1915: 50. Type-locality: Russia, “Karskaja Tundra” [= Yamalo-Nenetskiy AO, east slope of Polar Urals]. Holotype ♀, ZIN (seen by Pont, 2004: 79).

Siberian records: Pripolar Urals, Mt. Neroika ( Malozemov, 1992: 92, 93); Yamalo-Nenetskiy AO, east slope of Polar Urals [as “Karskaja Tundra”] (Schnabl in Becker et al., 1915: 50, type-locality of arctica ); valley of River Yenisey, forest tundra: Dudinka, Noril’sk ( Sychevskaya, 1974: 129); West and East Siberia ( Pont, 1986a: 142).

General distribution: Holarctic Region; in the Palaearctic, from montane and northern Europe east to the Russian Far East.

Comment. Hennig (1963c: 897) referred to 4♀ syntypes of P.arctica , but Schnabl described this species from a single ♀ which was seen by Pont (2004: 79) though he did not refer to it as the holotype.

The holotype was collected on 21 July 1909. According to the expedition itinerary by Backlund (1911: 74), the expedition made stop 26 on 21 July, by Mt Minisey, 180 m.

Helina impuncta ( Fallén, 1824)

Musca impuncta Fallén, 1824: 70 . Type-locality: not stated ( Sweden, probably Skåne province). Lectotype ♂, NHRS (designated by Pont, 1984b: 286).

Siberian records: south part of Tyumenskaya Oblast’, subtaiga ( Veselkin, 1966: 788); West Siberia ( Pont, 1986a: 143).

General distribution: Palaearctic Region, from Europe east to Japan.

Helina sp. near intermedia (Villeneuve, 1899)

Siberian record: Novosibirskaya Oblast’ ( Sorokina, 2006a: 228).

Helina latitarsis Ringhahl, 1924

Helina latitarsis Ringdahl, 1924b: 54 . (As “ latitarsis n. nom. pro nivalis Stein nec Zett.”) Type-localities: Germany (Genthin, Rügen Island, Ürdingen), Poland (Treptow = Trzebiatów), Austria (Innsbruck, Hochschwab), Czech Republic ( Bohemia), Italy (Trafoi) in Stein (1916: 65, as nivalis); Sweden (Skåne, Torne Lappmark, Småland, Öland). Syntypes 3♂ 2♀, ZMHU ( Pont & Werner, 2006: 61), 13♂ 4♀, MZLU (seen), and 3♂, NHRS (seen).

Siberian records: Novosibirskaya Oblast’, Karasukskiy Rayon, forest-steppe ( Sorokina, 2008: 163).

General distribution: Palaearctic Region, from Europe east to China.

Helina laxifrons ( Zetterstedt, 1860)

Aricia laxifrons Zetterstedt, 1860: 6200 . Type-locality: Sweden, “Bottnia boreali” (Torne Lappmark). Lectotype ♀, NHRS (designated by Pont, in press).

Siberian records: Novosibirskaya Oblast’ ( Sorokina, 2006a: 228); Altay Mts ( Sychevskaya, 1978: 82); West Siberia ( Pont, 1986a: 144).

General distribution: Holarctic Region; in the Palaearctic, from Europe east to the Russian Far East.

Helina longicornis ( Zetterstedt, 1838)

Anthomyza longicornis Zetterstedt, 1838: 678 . Type-locality: Sweden, Torne Lappmark , Vittangi. Lectotype ♂, MZLU (designated by Pont, in press).

Siberian records: Pripolar Urals, Mt. Neroika ( Malozemov, 1992: 92, 93); Irkutskaya Oblast’, Ust’-Kut on River Lena ( Hennig, 1963c: 896); East Siberia ( Pont, 1986a: 1434).

General distribution: Holarctic Region; in the Palaearctic, from northern Europe to the Russian Far East.

Helina luteisquama ( Zetterstedt, 1845)

Aricia luteisquama Zetterstedt, 1845: 1492 . Type-locality: Sweden, Lule Lappmark , Tjamotis near Kvikkjock. Holotype ♂, NHRS (Pont, in press).

Siberian records: Polar Urals (Schnabl in Becker et al., 1915: 37; Hennig, 1958: 195); West Siberia ( Pont, 1986a: 145).

General distribution: Holarctic Region; in the Palaearctic, from Norway, Sweden, Finland and Russia (north-west European Russia, Siberia and Far East).

Helina maculipennis ( Zetterstedt, 1845)

Aricia maculipennis Zetterstedt, 1845: 1475 . Type-locality: Sweden, Dalarna. Lectotype ♂, NHRS (designated by Pont, in press).

Siberian records: South Urals, Mt. Iremel’ ( Malozemov, 1989: 37); Novosibirskaya Oblast’ ( Sorokina, 2006a: 228).

General distribution: Holarctic Region; in the Palaearctic, from Europe, European Russia, and the Russian Far East.

Helina mandschurica Hennig, 1957

Helina mandschurica Hennig, 1957: 150 , 156 (key) (formal description in 1958: 196). Type-locality: China, Heilongjiang, Ha-erh-pin. Holotype ♂, MAKB.

Siberian records:?East Siberia ( Pont, 1986a: 145).

General distribution: Russia (?East Siberia and Far East), Mongolia and North China.

Helina sp. near mandschurica Hennig, 1957

Central Respublika Sakha [Yakutiya]: Yakutsk, Khomustakh nasleg ( Sychevskaya, 1972: 147) .

Helina moedlingensis (Schnabl, 1911)

Mydaea (Spilogaster) moedlingensis Schnabl in Schnabl and Dziedzicki, 1911: 290. Type-localities : Poland, Ruda Guzowska = Żyrardow , Warsaw district , and Toruń , Ciechocinek; Austria, Mödling; and Germany, Erfurt. Syntypes 6♂, formerly in MZPW, Schnabl collection (destroyed 1945).

Siberian records: Novosibirskaya Oblast’, Karasukskiy Rayon, forest-steppe ( Sorokina, 2008: 163).

General distribution: Palaearctic Region, from Europe east to China, and Pakistan.

Helina obscurata ( Meigen, 1826)

Anthomyia obscurata Meigen, 1826: 89 . Type-locality: not stated ( Germany, probably Stolberg). Holotype ♂, MNHN ( Pont, 1986b: 234).

Siberian records: Urals ( Hennig, 1958: 201; Zimin & El’berg, 1970: 535); south part of Tyumenskaya Oblast’, Moskovskiy and Salairka, subtaiga ( Veselkin, 1966: 784,788); Tomsk ( Wnukowsky, 1936: 124); Novosibirskaya Oblast’, Karasukskiy Rayon, forest-steppe ( Sorokina, 2008: 163); Altay Mts ( Sychevskaya, 1978: 82); Irkutskaya Oblast’, Listvyanka (Lavčiev, 1971: 217); Central Respublika Sakha [Yakutiya] ( Sychevskaya (1972: 147); valley of River Yenisey, northern taiga: Turukhansk, Verkhneimbatskoe, Yarzevo, Vorogovo, Kureyka, Igarka ( Sychevskaya, 1974: 129); West and East Siberia ( Pont, 1986a: 145).

General distribution: Holarctic Region; in the Palaearctic from Europe east to Japan.

Helina obscuratoides ( Schnabl, 1887)

Aricia obscuratoides Schnabl, 1887: 347 (also in Polish in 1888a: 209). Type-localities: Poland, Toruń,

Ciechocinek; and Belarus, Mogilev, Vitebsk district, Zachernya near Liozno. Syntypes 3♂, formerly in

MZPW, Schnabl collection (destroyed 1945).

Siberian records: Novosibirskaya Oblast’ ( Sorokina, 2006a: 228: as aff. obscuratoides ); West Siberia ( Pont,

1986a: 146).

General distribution: Holarctic Region; in the Palaearctic, from Europe east to the Russian Far East.

Helina parcepilosa ( Stein, 1907)

Spilogaster parcepilosa Stein, 1907: 325 . (Full formal description as sp.n. by Stein in Becker, 1908: 97.) Type-locality: China, Gansu, “Oase Satschou” (Dunhuang = Minghoshan). Lectotype ♂, ZIN (designated by Pont, 2004: 87).

Siberian records: Novosibirskaya Oblast’ ( Sorokina, 2006a: 228, 2008: 163); West Siberia ( Pont, 1986a: 138).

General distribution: Palaearctic Region, from Europe east to China.

Helina pyrrhopyga Pont, 1975

Helina pyrrhopyga Pont, 1975: 191 . Type-locality: Russia, East Siberia, Respublika Sakha [Yakutiya], Troitsk , Olekminsk. Holotype ♂, ZMUM.

Siberian records: Respublika Sakha [Yakutiya], Troitsk, Olekminsk ( Pont, 1975: 191, type-locality); East Siberia ( Pont, 1986a: 147).

General distribution: Known only from the type-locality.

Helina quadrum ( Fabricius, 1805)

Musca quadrum Fabricius, 1805: 297 . Type-locality: Austria. Holotype,?sex, ZMUC, Fabricius collection ( Michelsen, 1979: 188).

Siberian records: Omsk ( Kertész, 1901: 186); Altay Mts ( Sychevskaya, 1978: 82); West Siberia ( Pont, 1986a: 148).

General distribution: Palaearctic Region, from Europe east to China.

Helina reversio ( Harris, 1780)

Musca reversio Harris, 1780: 146 , plate 43 fig. 62. Type-locality: not stated (South-east England). Type-series lost ( Pont & Michelsen, 1982: 40).

Anthomyia duplicata Meigen, 1826: 92 . Type-locality: not stated ( Germany, probably Stolberg). Lectotype ♂, MNHN (designated by Pont, 1986b: 221).

Siberian records: Urals ( Hennig, 1957: 180); south part of Tyumenskaya Oblast’, subtaiga ( Veselkin, 1966: 788); Novosibirskaya Oblast’, Karasukskiy Rayon, forest-steppe ( Sorokina, 2006a: 228, 2008: 163); Altay Mts ( Sychevskaya, 1978: 82); West and East Siberia ( Pont, 1986a: 148).

General distribution: Holarctic Region, and also Taiwan; in the Palaearctic, abundant from Europe to Japan.

Helina sp. near reversio ( Harris, 1780)

Central Respublika Sakha [Yakutiya]: Yakutsk, Khomustakh nasleg ( Sychevskaya, 1972: 147, identification queried).

Helina setiventris Ringdahl, 1924

Helina setiventris Ringdahl, 1924b: 64 . (As “ setiventris n.nom. pro duplaris Stein nec Zett.”) Type-localities: Germany (Genthin, Kissingen), Denmark (Bornholm Island), Austria (Vienna), Sweden, France (Dauphiné), in Stein (1916: 61); Sweden, at several localities in Skåne, Öland Island, Stockholm. Syntypes ♂ ♀, MZLU, and 4♂, ZMHU ( Pont & Werner, 2006: 95).

Siberian records: Novosibirskaya Oblast’ ( Sorokina, 2006a: 228, 2008: 163); West Siberia ( Pont, 1986a: 150).

General distribution: Palaearctic Region, from Europe east to Russia (West Siberia) and Turkmenistan.

Helina sexmaculata ( Preyssler, 1791)

Musca sexmaculata Preyssler, 1791: 88 , plate 2 fig. 10. Type-locality: Czech Republic, Prague, Botanic Garden. Holotype / syntypes ♀, destroyed ( Rozkošný, Chvála & Pont, 1982: 352).

Musca uliginosa Fallén, 1825b: 81 . Type-locality: not stated ( Sweden, probably Skåne province). Lectotype ♂, NHRS (designated by Pont, 1984b: 295).

Siberian records: south part of Tyumenskaya Oblast’, subtaiga ( Veselkin, 1966: 788); West Siberia ( Pont, 1986a: 150).

General distribution: Holarctic Region, and also parts of the Oriental Region ( Myanmar, China, India, Nepal, Pakistan, Taiwan) and introduced into New Zealand.

Helina subvittata ( Séguy, 1923)

Phaonia subvittata Séguy, 1923: 336 . Type-locality: France, Hautes-Alpes, Lautaret. Holotype ♂, MNHN ( Hennig, 1963c: 899).

Helina rothi Ringdahl, 1939: 150 . (“N. nomen pro H. marmorata Stein (nec Zett.).”) Type-locality: Finland, Kittilä. Lectotype ♂, ZMHU (designated by Hennig, 1958: 217).

Syn: marmorata of Stein (misidentifications), not Zetterstedt, 1860

Siberian records: Urals, Sverdlovskaya Oblast’ (Nevyansk), Chelyabinskaya Oblast’ (Zlatoust) ( Hennig, 1958: 218); South Urals ( Zimin & El’berg, 1970: 529); “Siberia” ( Matsumura, 1911: 3, 82); west shores of Lake Baykal, near Pokoyniki (Lavčiev, 1971: 217); West and East Siberia ( Pont, 1986a: 151).

General distribution: Holarctic Region; in the Palaearctic, from Europe east to Japan.

Helina trivittata ( Zetterstedt, 1860)

Anthomyza trivittata Zetterstedt, 1860: 6267 . Type-locality: Sweden, Skåne, Fågelsång near Lund. Holotype ♀, MZLU (Pont, in press).

Spilogaster atripes Meade, 1889: 425 ). Type-locality: United Kingdom, England, Yorkshire , Hornsea near Hull. Lectotype ♂, BMNH (designated by Pont, 1984a: 10).

Siberian records: Novosibirskaya Oblast’ ( Sorokina, 2006a: 228, 2008: 163); Altay Mts ( Sychevskaya, 1978: 82); West Siberia ( Pont, 1986a: 151).

General distribution: Palaearctic Region, from Europe east to China.

Helina veterana ( Zetterstedt, 1838)

Anthomyza veterana Zetterstedt, 1838: 662 . Type-locality: Sweden, Torne Lappmark , Vittangi. Lectotype ♂, NHRS (designated by Pont, in press).

Siberian records: West Siberia ( Pont, 1986a: 152).

General distribution: Palaearctic Region, in montane and northern Europe, and Russia (West Siberia).

Helina sp. 1 .

Novosibirskaya Oblast’, Karasukskiy Rayon, forest-steppe ( Sorokina, 2008: 163).