PLANORBIDAE Rafinesque, 1815

Coan, Eugene V., Pacaud, Jean-Michel & Kabat, Alan R., 2024, Gérard-Paul Deshayes (1796 - 1875) and his taxa, Geodiversitas 46 (16), pp. 553-930 : 680-681

Family PLANORBIDAE Rafinesque, 1815

ambiguus , Planorbis – Deshayes, 1863-DescrCoqFoss: 744-745, 47, pl. 47, figs 10-13. — Type localities: Le Fayel, Ducy, Mareuil-en-Dole, La Ferté-sous-Jouarre, Nantheuil-sur Marne, Avize, Saint-Aubin & Saint-Saturin (Sarthe). — Type age: Eocene (Bartonian). — Type material: MNHN.F.J04167, syntype. — Current status: synonym of Hippeutis (Hippeutis) obtusus (J. Sowerby, 1818) (Wenz 1923: 1643; Kadolsky 1975: 131) .

boissyi , Planorbis – Deshayes, 1863-DescrCoqFoss: 741, 45, pl. 45, figs 20-21, non Potiez & Michaud, 1838. — Type locality: Rilly-la-Montagne. — Type age: Paleocene (Thanetian). — Type material: UCBL-EM 32584, 2 syntypes. Planorbis rillyensis Bayan, 1870 ( Bayan 1870: 4), replacement name. — Current status: Headonia rillyensis ( Bayan, 1870) (Harzhauser & Neubauer, in Harzhauser et al. 2020: 24).

bourgeoisi , Ancylus – Deshayes, 1863-DescrCoqFoss: 700-701, 42, pl. 42, figs 22-25. — Type locality: Montabuzard. — Type age: Miocene (Burdigalian). — Current status: Ancylus bourgeoisi Deshayes, 1863 (Wenz 1923: 1862).

campaniensis , Planorbis – Deshayes, 1863-DescrCoqFoss: 747-748, 45, pl. 45, fig. 22. — Type locality: Cuis. — Type age: Eocene (Ypresian). Biomphalaria sparnacensis [Deshayes, 1824] campaniensis (Deshayes, 1863) ( Le Renard & Pacaud 1995: 129) . — Current status: Headonia campaniensis (Deshayes, 1863) (Harzhauser & Neubauer, in Harzhauser et al. 2020: 24).

chertieri , Planorbis – Deshayes, 1863-DescrCoqFoss: 753-754, 47, pl. 46, figs 5-8. — Type localities: Saint-Parres-lès-Vaudes & Morancez. — Type age: Eocene (Lutetian). — Type material: MNHN.F.J04179, 11 syntypes (Hébert coll.) from Morancez; UCBL-EM 32586, syntypes (Deshayes coll.) from Saint-Parres-lès-Vaudes. — Current status: Hippeutis (Hippeutis) chertieri (Deshayes, 1863) (Wenz 1923: 1636; Le Renard & Pacaud 1995: 130; Pacaud 2008: 76).

concavus , Planorbis – Deshayes, 1863-DescrCoqFoss: 745-746, 46, pl. 46, figs 9-12. — Type locality: Passy (Paris). — Type age: Eocene (Lutetian). — Type material: MNHN.F.J04180, 2 syntypes (Hébert coll.); UCBL-EM 32587, syntype (Deshayes coll.). — Current status: Menetus concavus (Deshayes, 1863) ( Le Renard & Pacaud 1995: 130; Pacaud 2008: 76).

depressus , Ancylus – Deshayes, 1824-DescrCoqFoss: 5, pl. 10, fig. 13; 1825-DescrCoqFoss: 101; 1830a-EncyMeth: 48-49; 1836-HistNatAnim2: 657; 1843-HistNatAnim3: 242; 1863-DescrCoqFoss: 699. — Type locality: Jouy. — Type age: Oligocene (Chattian). Preoccupies Ancyclus depressus Haldeman, 1844 , which was renamed Ancylus haldemani Boettger, 1853 . — Current status: Ancylus depressus Deshayes, 1824 (Wenz 1923: 1686).

duveyrieri , Planorbis –Deshayes, 1864: 44-45, l. 3, fig. 1 [Deshayes, in Duveyrier]; Bourguignat 1864: 25-26, 30, pl. 27, fig. 6 [see under: Deshayes, in Duveyrier]. — Type locality: White soil near Ghoûrd- Ma’ammer, on the road from El-Quâd to Ghadâmè, Libya. — Type age: Recent. — Current status: synonym of Biomphalaria pfeifferi (Krauss, 1848) [ Planorbis ] ( Chevallier 1969: 278-280), an extinct species closely related to Biomphalaria glabrata (Say, 1818) , an intermediate snail host for the trematode parasite Schistosoma mansoni .

exustus , Planorbis – Deshayes, 1832b: 417-418, 439, 522, pl. 1, figs 11-13; 1838-HistNatAnim2: 392; 1843-HistNatAnim3: 409. — Type locality: Malabar, India, freshwater; Bélanger. — Type age: Recent. — Type species (OD) of Indoplanorbis Prashad & Annandale, 1921 . — Current status: Indoplanorbis exustus (Deshayes, 1832) (D. Lamy & Pointier 2018: 623, introduced on Caribbean islands; Koudenoukpo et al. 2020: 48). However, this species, often misdated as 1833 or 1834, is currently believed to be a complex of up to six closely related biological species in India and Southeast Asia, with populations introduced elsewhere in the world ( Sil et al. 2022). Preece et al. (2022: 112-113) noted that the Deshayes species was described from the Malabar coast in southwestern India, while its supposed junior synonym Planorbis indicus Benson, 1836 , was described from the Gangetic provinces of eastern India, so that Benson’s name could be used for a distinct species if the Deshayes name was to be limited to the western India populations.

inflatus , Planorbis – Deshayes, 1824-DescrCoqFoss: 5, pl. 10, 3-5; 1825-DescrCoqFoss: 86-87; 1863-DescrCoqFoss: 745. — Type localities: Septeuil & La Villette (Paris). — Type age: Eocene (Bartonian). Hippeutis (Hippeutis) inflatus (Deshayes, 1825) (Wenz 1923: 1637; Le Renard & Pacaud 1995: 130). — Current status: synonym of Hippeutis (Hippeutis) lens (Brongniart, 1810) (Pacaud herein).

inversus, Planorbis – Deshayes, 1824-DescrCoqFoss: 5, pl. 9, figs 16- 18; 1825-DescrCoqFoss: 88; 1863-DescrCoqFoss: 755. — Type locality: La Villette (Paris). — Type age: Eocene (Bartonian). — Type material: UCBL-EM 32381, syntype from Septeuil. — Current status: synonym of Hippeutis (Hippeutis) inflatus (Deshayes, 1824) (Wenz 1923: 1637) .

laevigatus , Planorbis – Deshayes, 1824-DescrCoqFoss: 5, pl. 9, figs 19-21; 1825-DescrCoqFoss: 85-86; 1832-EncyMeth: 782; 1838-HistNatAnim2: 396; 1843-HistNatAnim3: 410; 1863-DescrCoqFoss: 746. — Type locality: Épernay. — Type age: Eocene (Ypresian). Biomphalaria laevigata (Deshayes, 1824) ( Le Renard & Pacaud 1995: 129) . — Current status: Headonia laevigata (Deshayes, 1824) (Harzhauser & Neubauer, in Harzhauser et al. 2020: 24).

leymerii , Planorbis – Deshayes, 1863-DescrCoqFoss: 739-740, 46, pl. 46, figs 1-4. — Type localities: Côte Saint-Parres (Saint-Nicolasla-Chapelle), Les Éparmailles, Provins & Morances. — Type age: Eocene (Lutetian). — Type material: UCBL-EM 32585, syntype from the Côte Saint-Parres. — Current status: Headonia pseudoammonia [Schlothem, 1820; Helicites ] leymerii (Deshayes, 1863) (Harzhauser & Neubauer, in Harzhauser et al. 2020: 24).

paciacensis , Planorbis – Deshayes, 1863-DescrCoqFoss: 740-741, 46, pl. 46, figs 13-16. — Type locality: Passy (Paris), Chéry-Chartreuve & Boursault. — Type age: Eocene (Lutetian-Bartonian). — Type material: UCBL-EM 32588, syntype from Passy. Biomphalaria paciacensis (Deshayes, 1863) ( Le Renard & Pacaud 1995: 129; Pacaud 2008: 76, as B. “ paciaciensis ”). — Current status: Headonia paciacensis (Deshayes, 1863) (Harzhauser & Neubauer, in Harzhauser et al. 2020: 24).

planulatus, Planorbis – Deshayes, 1824-DescrCoqFoss: 5, pl. 10, figs 8-10; 1825-DescrCoqFoss: 88-89; 1838-HistNatAnim2: 397; 1843-HistNatAnim3: 410-411; 1863-DescrCoqFoss: 753. — Type locality: Pantin & La Villette (Paris). — Type age: Eocene (Bartonian). Hippeutis (Hippeutis) inflatus [Deshayes, 1825] planulatus (Deshayes, 1824) ( Le Renard & Pacaud 1995: 130). — Current status: synonym of Hippeutis (Hippeutis) lens (Brongniart, 1810) (Pacaud herein).

pygmaeus , Planorbis – Deshayes, 1863-DescrCoqFoss: 750, 46, pl. 46, figs 36-39. — Type locality: Mouchy-le-Châtel. — Type age: Eocene (Lutetian). — Type material: UCBL-EM 32591, syntype. — Current status: Menetus pygmaeus (Deshayes, 1863) (Le Renard & Pacaud 195: 129; Pacaud 2008: 76).

sparnacensis , Planorbis – Deshayes, 1824-DescrCoqFoss: 5, pl. 10, figs 6-7; 1825-DescrCoqFoss: 86; 1832-EncyMeth: 782-783; 1838-HistNatAnim2: 396; 1843-HistNatAnim3: 410; 1863-DescrCoqFoss: 746-747, 45, pl. 45, figs 27-30 [in text as figs 27-29]; 1865-DescrCoqFoss: 666 [correction noted]. — Type locality: Mont-Bernon (Épernay). — Type age: Eocene (Ypresian). — Type material: UCBL-EM 32138, syntype. — Current status: Biomphalaria sparnacensis (Deshayes, 1824) ( Le Renard & Pacaud 1995: 129) , now Headonia sparnacensis (Deshayes, 1824) (Harzhauser & Neubauer in Harzhauser et al. 2020: 24).

spiruloides , Planorbis – Deshayes, 1863-DescrCoqFoss: 747, 46, pl. 47, figs 6-9. — Type locality: Ducy. — Type age: Eocene (Bartonian). — Type material: UCBL-EM 32594, syntypes. — Current status: Menetus spiruloides (Deshayes, 1863) ( Le Renard & Pacaud 1995: 129) .

subangulatus , Planorbis – Deshayes, 1824-DescrCoqFoss: 5, pl. 9, figs 14-15; 1825-DescrCoqFoss: 87, non Lamarck, 1804; 1832-EncyMeth: 783. — Type localities: Pantin & La Villette (Paris). — Type age: Oligocene (Rupelian). — Current status: synonym of Gyraulus depressus (Nyst, 1845) (Wenz 1923: 1549) [ Planorbis ].

subovatus , Planorbis –Deshayes, 1824-DescrCoqFoss: 5, pl. 9, figs 19- 21; 1825-DescrCoqFoss: 85; 1832-EncyMeth: 780; 1838-HistNatAnim2: 396; 1843-HistNatAnim3: 410; 1863-DescrCoqFoss: 742. — Type locality: Épernay. — Type age: Eocene (Ypresian). — Current status: Biomphalaria subovata (Deshayes, 1824) ( Le Renard & Pacaud 1995: 130) .

thibetanus, Planorbis –Deshayes, 1870b: 27. — Type locality:Moupin, Tibet. — Type age: Recent. — Current status: Taxon inquirendum.