Tachysphex jokischianus (Panzer, 1809), restored from synonymy

Straka, Jakub, 2016, Tachysphexaustriacus Kohl, 1892 and T. pompiliformis (Panzer, 1804) (Hymenoptera, Crabronidae) are a complex of fourteen species in Europe and Turkey, ZooKeys 577, pp. 63-123 : 80-87

Taxon classification Animalia Hymenoptera Crabronidae

Tachysphex jokischianus (Panzer, 1809), restored from synonymy Figs 5, 17, 23, 31, 39, 45, 51, 57, 61, 65, 72, 84, 88, 97

Larra Jokischiana Panzer, 1809 (Heft 106: pl. 15), ♀.

Type material.

Holotype: ♀, no specific locality at the label but probably from Germany. Holotype by monotypy. "Coll. Sturm [printed] // Ca. // Larra / Jokischi / ana / Panz. [handwriting, card with black frame] // 54 [printed, red card] // Larra / Jokischiana / Panz. Typus / J. de Beaumont / det. 1955 [handwriting on pre-printed card with name of determinator and year] // Tachysphex ♀ / pompiliformis Pz. / J. de Beaumont / det. 1955 [handwriting on pre-printed card with name of determinator and year] // Typus Nr. / Zoologische / Staatsamlung / München. [printed, red card]". Holotype in ZSM, examined.

Additional material examined.

Austria: Seefeld NW, Niederösterreich, 1 ♂, 19. v. 2015, M. Halada lgt., OLML; Czech Republic: Boh. bor., Počerady, ash coal deposit from power station, 1 ♂, 23. v. 2010, 1 ♂, 7. vi. 2010, 1 ♂, 1 ♀, 27. vi. 2010, 1 ♂, 1 ♀, 1. viii. 2010, R. Tropek lgt., JSPC; Boh. centr., Cihelna v Bažantnici, Praha-Hloubětín, 14 ♂♂, 1 ♀, 28. v. 1998, 6♂♂, 1 ♀, 22. vi. 1998, J. Straka lgt., JSPC; Černošice, 1 ♂, 30. v. 1968, Z. Pádr lgt., OLML; Divoká Šárka, Praha-Liboc, 1 ♂, 29. v. 1998, J. Straka lgt., JSPC; Heřmanův Městec, sandpit, 2 ♂♂, 1 ♀, 1. vii. 2012, 1 ♀, 1. viii. 2012, R. Tropek lgt., JSPC; Hrachovka, Praha-Troja, 1 ♀, 28. vi. 1998, J. Straka lgt., JSPC; Chvaletice, ash coal deposit, 1 ♂, 1. viii. 2012, R. Tropek lgt., JSPC; Chvalská navážka, Praha-H. Počernice, 1 ♂, 9. vii. 1997, 1 ♀, 22. viii. 1997, 2 ♂♂, 1 ♀, 6. vii. 1998, 1 ♀, 20. viii. 1998, J. Straka lgt., JSPC; Chvalský lom, Praha-H. Počernice, 1 ♀, 6. vii. 1998, J. Straka lgt., JSPC; Jenštejn, 4 ♂♂, 22. vi. 2001, Jos. Straka lgt., JSPC; Klánovický les, Praha-Klánovice, 1 ♂, 20. vi. 1998, J. Straka lgt., JSPC; Opatovice, ash coal deposit, 1 ♀, 1. vi. 2014, R. Tropek lgt., JSPC; Písečný vršek, Praha-Běchovice, 1 ♀, 2. vii. 1997, 5 ♂♂, 1 ♀, 6. vii. 1998, M. Kolísko & J. Straka lgt., JSPC; Písty, Nymburk env., sand dune, 1 ♀, 1. vii. 2012, 1 ♀, 1. viii. 2012, R. Tropek lgt., JSPC; Travčický les, Terezín env., 1 ♂, 6. viii. 1981, Z. Pádr lgt., OLML; Valy, 1 ♂, 7. viii. 1988, Odehnal lgt., OLML; Boh. mer., Blatná, sand quarry, 1 ♂, 1998, P. Bogusch lgt., JSPC; Cep 1, sand quarry, 1 ♂, 4.-9. vi. 2013, 1 ♀, 1.-5. vii. 2013, R. Tropek, I. Černá, O. Čížek lgt., JSPC; Františkov, Chlum u Třeboně, 1 ♂, 29. vii. 2002, J. Halada lgt., JSPC; Hodějovice, 5 ♂♂, 1 ♀, 3. vi. 2015, 1 ♀, 2. viii. 2013, R. Tropek, I. Černá, O. Čížek lgt., JSPC; Plavsko, 1 ♂, 5. vii. 2013, R. Tropek, I. Černá, O. Čížek lgt., JSPC; Těšínov, 1 ♀, 2. viii. 2013, R. Tropek, I. Černá, O. Čížek lgt., JSPC; Slepičí vršek, Třeboňsko PLA, 1 ♂, 2 ♀♀, 6. viii. 1997, J. Straka lgt., JSPC; Vlkov, Třeboňsko PLA, 1 ♂, 6. viii. 1997, J. Straka lgt., 1 ♂, 1 ♀, 9. vi. 2013, R. Tropek, I. Černá, O. Čížek lgt., JSPC; Boh.occ., Starý lom, ash coal deposit from Prunéřov power station, 1 ♀, 26. vi. 2010, 1 ♀, 31. vii. 2010, R. Tropek lgt., JSPC; Tušimice, ash coal deposit from power station, 1 ♂, 6. vi. 2010, 2 ♂♂, 2 ♀♀, 26. vi. 2010, 1 ♀, 22. viii. 2010, R. Tropek lgt., JSPC; Boh. or., Plachta NR, Hradec Králové, 2 ♂♂, 1. vii. 1998, J. Straka lgt., JSPC; Veská, Pardubice env., sand dune, 1 ♂, 20. v. 2007, 1 ♀, 27. v. 2007, P. Bogusch lgt., 1 ♀, 1. viii. 2012, 3 ♂♂, 1 ♀, 1. vii. 2012, R. Tropek lgt., JSPC; Mor.mer., Bzenec env., 1 ♂, 18. vi. 1942, A. Hoffer lgt., 1 ♀, viii. 1963, M. Kocourek lgt., OLML; Bzenec env., Vojenské cvičiště NR, 1 ♀, 8. vi. 1997, M. Kolísko lgt., 1 ♀, 4. viii. 2015, J. Straka lgt., JSPC; Bzenec-přívoz, Bzenec env., 4 ♂♂, 31. v. 2002; J. Straka lgt., JSPC; Čučice 1 km S, 2 ♂♂, 1.-15.vii.2001, M. Říha lgt., MRBC; Dolní Bojanovice, 1 ♂, viii. 1941, A. Hoffer, OLML; Mistřín, 1 ♂, viii. 1942, A. Hoffer lgt., OLML; Pouzdřany, 1 ♂, 19. vi. 1970, J. Strejček lgt., OLML; Rohatec lgt., 1 ♂, vii. 1941, 2 ♀♀, vii. 1942; A. Hoffer, OLML; France: Falaise, 1 ♂,date and collector unknown, ZSM; Plateau du Coiron, 16 km NW Montélimar, 2 ♂♂, 24. vi. 2001, 2 ♂♂, 1 ♀, 11.-14 vii. 2002, 2 ♂♂, 1 ♀, 20. vi. 2004, J. v. d. Smissen lgt., USMB; Lachamp - Raphael W, D 122, 23 km NW Aubenas, 1 ♂, 2 ♀♀, 18. vi. 2004, 1 ♀, 29. vii. 2003, J. v. d. Smissen lgt., USMB; Cereste, Apt., Provence prov., 1 ♀, 9. ix. 1987, Ch. Schmid-Egger lgt., CSE; Germany: Allersberg, 1 ♂, 17. viii. 1956, Ettinger lgt., ZSM; Blankenförde, Mecklenburg-Vorpommern, mn-hex, 2 ♀, 7. vii. 2013, Ch. Schmid-Egger lgt., ZSM HYM 19879, ZSM HYM 19880; Blankensee, Lübeck, Schleswig-Holstein, 1 ♀, 18. vi. 1989, J. v. d. Smissen lgt., USMB; Brammerau, Nortorf, Schleswig-Holstein, 1 ♂, 1 ♀, 22. vii. 1995, J. v. d. Smissen lgt., USMB; Brünkendorf /Dannenberg, Niedersachsen, 3 ♂♂, 2. viii. 1995, J. v. d. Smissen lgt., USMB; Büchen, Schleswig-Holstein, 10 ♂♂, 10. vii. 1994, J. v. d. Smissen lgt., USMB; Damsdorf, Bornhöved 6km E, Schleswig-Holstein, 1 ♂, 18. vii. 1999, J. v. d. Smissen lgt., USMB; Eichholz, Lübeck, Schleswig-Holstein, 1 ♀, 25. vii. 1989, J. v. d. Smissen lgt., USMB; Friedrichsfeld, Mannheim, Baden-Württenberg, 1 ♂, 22. v. 1989, Ch. Schmid-Egger lgt., CSE; Friedrichsort, Kiel, Lehmgrube, 1 ♀, 1. vii. 1961, collector unknown, ZSM; Gabow, Bad Freienwalde 4km NE, gab, 1 ♀, 25. vi. 2001, Ch. Schmid-Egger lgt., CSE; Gemünda, Sesslach, Bayern, 2 ♂♂, 9. vii. 1994, J. v. d. Smissen lgt., USMB; Grissheim, Baden-Württemberg, gr, 1 ♂, 18. vii. 2012, Ch. Schmid-Egger, ZSM HYM 17628; Grissheim, Rhein, Freiburg 24 km SW, Baden-Württemberg, 1 ♀, 1. vii. 1999, J. v. d. Smissen lgt., USMB; Gudow, Schleswig-Holstein, 1 ♀, 1. ix. 1991, J. v. d. Smissen lgt., USMB; Ingolstadt, 1 ♀, 21. vii. 1948, H. Freude lgt., ZSM; Istein, Baden-Württenberg, 1 ♂, 26. vii. 1989, Ch. Schmid-Egger lgt., CSE; Klein Kühren /Dannenberg, Niedersachsen, 1 ♂, 15. vi. 1996, 1 ♀, 10. viii. 1996, J. v. d. Smissen lgt., USMB; Laasche/Dannenberg, Niedersachsen, 3 ♂♂, 5 ♀♀, 19. vii. 1994, 12 ♂♂, 23. viii. 1994, J. v. d. Smissen lgt., USMB; Lenggries, Isarauen, 1 ♀, 27. vii. 2003, Ch. Schmid-Egger lgt., ZSM; Malk Göhren, Mecklenburg-Vorpommern, 2 ♂♂, 22. vii. 1997, J. v. d. Smissen lgt., USMB; Mallnow, Framkfurt an der Oder 13km NNW, Brandenburg, br-ma, 1 ♀, 22. vii. 2009, ZSM HYM 14826, 1 ♀, 26. vi. 2010, Ch. Schmid-Egger lgt., ZSM HYM 14825; Malsch, 1 ♀, 25. vi. 1967, R. Degen lgt., ZSM; Oldenburg 5km NW, Schleswig-Holstein, 1 ♀, 26. vii. 1997, 2 ♀♀, 4. viii. 1997, J. v. d. Smissen lgt., USMB; Putlos, Oldenburg, Schleswig-Holstein, 1 ♀, 29. vi. 1997, 1 ♀, 4. viii. 1997, J. v. d. Smissen lgt., USMB; Sandhausen, Heidelberg, Baden-Württenberg, 1 ♂, 20. v. 1989, Ch. Schmid-Egger lgt., CSE; Spreenhagen 4km W, Brandenburg, brsp, 1 ♂, 27. vi. 2011, Ch. Schmid-Egger lgt., ZSM HYM 14827; Steinenstadt, Baden-Württemberg, 1 ♂, 31. vii. 2005, W. Schlaefle lgt., WSKC; Zossen, Brandenburg, zos, 1 ♀, 14. vi. 2011, Ch. Schmid-Egger lgt., ZSM HYM 09867; Hungary: Dunatetétlen, saline lake, Kecskemét 45km SEE, 1 ♂, 17. vii. 2013, J. Habermannová, J. Straka lgt., JSPC; Hegyshalom N, 1 ♂, 1 ♀, 6. vi. 2015, M. Halada lgt., OLML; Kecskemét env., 1 ♂, 16. vii. 2006, J. Halada lgt., JSPC; Kesthely env., lime stone, 1 ♂, 1 ♀, 23. v. 2014, A. Astapenková, J. Habermannová, J. Straka lgt., JSPC; Kunpeszér env., sandy hill, 1 ♀, 7. vi. 2013, D. Benda, P. Bogusch & J. Straka lgt., JSPC; Kunszentmiklós env., salt marsh, 3 ♀♀, 16. vii. 2013, J. Habermannová, J. Straka lgt., JSPC; Örkeny, puszta, 5 ♂♂, 7. vi. 2015, 1 ♀, 27. vi. 2015, M. Halada lgt., OLML; Pákozd env., pasture stepe, 4 ♂♂, 18. vii. 2013, J. Habermannová, J. Straka lgt., JSPC, 6 ♂♂, 5 ♀♀, 10. vi. 2015, 7 ♂♂, 11 ♀♀, 25. vi. 2015M. Halada lgt., OLML, 1 ♂, 9. vi. 2013, D. Benda, P. Bogusch & J. Straka lgt., JSPC; Pusztavacs, sand dune, 1 ♂, 8. vi. 2015, M. Halada lgt., OLML; Súr 1km S, loes steppe, 1 ♀, 18. vii. 2013, J. Habermannová, J. Straka lgt., JSPC; Szenna, Kaposvár env., 1 ♀, 3. ix. 2007, W. Żyła, USMB; Velencei-tó env., steppe, 1 ♀, 9. vi. 2013, D. Benda, P. Bogusch & J. Straka lgt., JSPC; Poland: Białowieża forest, 1 ♂, 19. vii. 1995, R. Dobosz lgt., USMB; Borne Sulinowo, Baltic see coast, 1 ♂, 8. vii. 1997, Kaka lgt., USMB; Grabowiec, Małopolska Upland, 1 ♀, 18. vi. 1995, 2 ♂♂, 4 ♀♀, 12. vii. 1996, W. Żyła, USMB; Jastrzębia Góra, Władysławowo, Baltic see coast, 1 ♂, 16. vi. 1996, 1 ♀, 24. vi. 1996, R. Dobosz lgt., USMB; Kików, Malopolska Upland, 4 ♂♂, 2 ♀♀, 19. vi. 1995, 1 ♀, 14. vii. 1996, W. Żyła lgt., USMB; Kołczewo, Baltic see coast, 1 ♀, 10. vii. 2009, R. Dobosz lgt., USMB; Krzyżanowice, Malopolska Upland, 4 ♂♂, 16. vi. 1995, 1 ♂, 21. vi. 1995, W. Żyła lgt., 1 ♀, 30. vii. 1996, R. Dobosz lgt., USMB; Kuźnica, Baltic see coast, 1 ♂, 1 ♀, 25. vi. 1996, R. Dobosz lgt., USMB; Lubliniec, Upper Silesia, 1 ♂, 12. vi. 1994, R. Dobosz lgt., USMB; Osowo Stare, 1 ♂, 2 ♀♀, 12. viii. 1992, 1 ♂, 28. v. 1993, M. Bunalski lgt., USMB; Pasturka, Pińczów env., Małopolska Upland, 2 ♂♂, 19. vi. 1997, R. Dobosz, W. Żyła lgt., USMB; Ponice, Rabki, Beskid Zachodni, 1 ♀, 17. vii. 1997, W. Żyła lgt., USMB; Przedbórz, Małopolska Upland, 1 ♀, 2. viii. 2007, W. Żyła lgt., USMB; Smołdziński Las, Słowiński NP, 1 ♂, 29. vii. 2004, W. Żyła lgt., USMB; Sycyn Dolny, 2 ♂♂, 2 ♀♀, 22. vi. 1992, 1 ♂, 2 ♀♀, 28. vii. 1992, 1 ♀, 12. viii. 1992, M. Bunalski lgt., USMB; Szombierki, Bytom, 1 ♀, 29. v. 1993, W. Żyła lgt., USMB; Wisła, 1 ♀, 1. viii. 1994, 1 ♂, 8. viii. 1994, W. Żyła lgt., USMB; Władysławowo, 2 ♂♂, 2 ♀♀, 11. vi. 1993, 2 ♂♂, 1 ♀, 13. vi. 1993, 1 ♂,18. vi. 1993, R. Dobosz lgt., USMB; Slovakia: Abrod NR, Velké Leváre env., 1 ♀, 6. vii. 2008, J. Straka, P. Janšta, P. Šípek, D. Král lgt., JSPC; Chotín, 2 ♂♂, 3. viii. 1961, 1 ♂, 9. viii. 1961, 1 ♂, 12. vii. 1981, Z. Pádr lgt., 1 ♂, 15. vi. 1977, M. Kocourek lgt., 1 ♀, 26. vii. 1993, M. Halada lgt., OLML; Královský Chlmec, 3 ♂♂, 1 ♀, 25. vii. 1969, Z. Pádr lgt., OLML; Kúty 1 ♂, 9. vi. 1973, J. Strejček lgt., OLML; Lakšárska Nová Ves env., sandy area, 2 ♂♂, 13. vi. 2008, J. Straka lgt., JSPC; Somotor, 2 ♂♂, vii. 1959, M. Kocourek, OLML; Streda nad Bodrogom, 1 ♂, 26. vi. 1977, Z. Pádr lgt., OLML; Štúrovo, 1 ♀, vii. 1957, M. Kocourek lgt., 1 ♂, 24. vi. 1960, 1 ♂, 8. vi. 1964, Z. Pádr lgt., OLML; Velký Kamenec, 1 ♀, 14. vii. 1975, Z. Pádr lgt., OLML; Sweden: Dammersryet, Ravlunda, 1 ♂, 9. vii. 2011, N. E. Johansson lgt., NJHC; Gårdby, Öland isl., 1 ♀, 27. vii. 1998, collector unknown, USMB; Havängsdösen, Ravlunda, 1 ♂, 27. vi. 2011, N. E. Johansson lgt., NJHC; Lillesjö, Mörlunda env., 1 ♀, 27. vi. 2009, N. E. Johansson lgt., NJHC; Sandhajd, Fårö env., 2 ♂♂, 1 ♀, 10. vii. 2012, 1 ♀, 12. vii. 2012, N. E. Johansson lgt., NJHC; Sandviken, Gannelgarn env., 1 ♀, 9. vii. 2012, N. E. Johansson lgt., NJHC

Diagnosis.

Tachysphex jokischianus (Panzer) is difficult to distinguish from Tachysphex dimidiatus (Panzer), Tachysphex pompiliformis (Panzer) and Tachysphex punctipleuris sp. n. It is exceptionally variable in most characters, and no single character differentiates it from the other morphologically similar species. It can be recognized by the following combination of characters: supraclypeal area with distinctly microsculptured interspaces between punctures, usually appearing dull. Gena dorsally robustly built. Mesopleuron, including hypoepimeral area, finely sculptured, punctures in lower parts of mesopleuron distinct but small. Propodeal dorsum usually lacking longitudinal ridges, if present, then only basally. Dorsolateral cuticular projection of propodeal spiracle arcuate to semicircular, with dark apex. Posteromedial margin of propodeal dorsum elevated but not produced between the marginal ridges, marginal ridges positioned nearly horizontally above groove on posterior side. Wings only slightly infumated to pale. ♀: Apical depression of terga IV and V finely microsculptured and impunctate or nearly so, contrasting with coarsely and densely punctate central part of these terga, punctures of terga IV and V well developed. Pygidium often wide, with convex, shiny surface (but not always). Tergum III all red or red laterally and basally, with black area apicomedially. ♂: Surface of supraclypeal area concave, supraantennal tubercle distinctly elevated ventromedially. Forefemoral notch light red, large, its longitudinal diameter usually longer than the space separating it from forefemoral base. Propodeal side densely and finely ridged basally in most specimens. Terga densely micropunctate.

Description of female (holotype).

Body length: 7.2 mm.

Head. Clypeus distinctly convex, its top at clypeal midlength; basomedian area relatively large, as densely punctate as lateral section; bevel large, distinctly convex, as long as basomedian area laterally, slightly longer medially, obtusely triangular, with sparse large punctures, not reaching base of clypeus, shiny; lip slightly arcuate, with lateral in cision, separated from bevel by punctate groove, WML:LCL = 1.5, WCL:WML = 1.8. Supraclypeal area flat, distinctly punctate, punctures ill-defined, interspaces microsculptured, slightly shiny. Supraantennal tubercle small, slightly elevated on inner side. Both antennae broken off (only three segments present). Frons uniformly punctate, punctures well defined, about half diameter apart or less, most interspaces uniform in size; frontal median line distinct, narrow, finely impressed. Vertex finely punctate, punctures well defined, less than half to one diameter apart, laterally more sparsely punctate than medially, interspaces finely microsculptured, slightly shiny. Vertex setae short, semierect, slightly longer laterally than medially, about 0.5 × MOD; postocellar impression well developed, widely Y-shaped; vertex moderately wider than long; WV:LV = 1.5. Gena dorsally well developed.

Mesosoma. Scutum without distinct anterior impression; scutum and scutellum densely punctate, punctures well defined, less than half to one diameter apart, interspaces well developed, variable in size, microsculptured, slightly shiny, setae about 1.0 × MOD or less. Mesopleuron coarsely microsculptured, without distinct punctures; hypoepimeral area irregularly microsculptured without punctures; ventral part of mesopleuron with punctures ill defined, less than one diameter apart, interspaces small, slightly shiny. Propodeal dorsum relatively finely, regularly rugose, with few indistinct irregular, longitudinal ridges basally, dull, but slightly shiny posteromedially; propodeal side regularly longitudinally ridged, ridges inconspicuous basally, interspaces microsculptured, slightly shiny; posteromedial margin of propodeal dorsum elevated, but not produced between posterior marginal ridges, marginal ridges positioned nearly horizontally above groove on posterior surface. Legs densely punctate, punctures small; forebasitarsal rake light yellow with reddish opalescence at distal half, with three apical spines and two to three additional spines. Wings slightly infumate with brown veins.

Metasoma. Terga I-III with ill defined silvery apical fasciae. Terga I-III densely, finely micropunctate, punctures ill defined, interspaces microsculptured, slightly shiny; apical depressions shallow, with distinct micropunctures; sculpture of terga IV-V distinctly coarser than on previous terga, also slightly more densely punctate; tergum V coarsely punctate, punctures ill-defined, but large, half to one diameter apart, interspaces microsculptured, apical depression of tergum V very sparsely and finely punctate. Pygidium large, sparsely punctate, punctures large, ill defined, interspaces nearly unsculptured, shiny. Central part of sternum II with several larger punctures, interspaces microsculptured, shiny; lateral part slightly shiny, densely micropunctate; remaining sterna with uniform sculpture similar to that on sternum II, but more or less reduced laterally.

Coloration. Central part of mandibles, terga and sterna I-III red. Distal tarsomeres dark red. Tegulae reddish translucent. Apical parts of terga I-III slightly translucent. Remaining body parts black.

Variation of females. Very variable in sculpture, but finely sculptured over all body in most specimens except apical terga (in comparison to other species). Clypeal bevel reaching base of clypeus in some specimens, but well separated from the base in most. Antenna short, LA3:WA3 = 2.2-2.4, LA5:WA5 = 2.6-2.7. Pygidium distinctly microsculptured, of usual width and dull in some specimens, but shiny and wide in most of specimens. Some specimens appear dark in coloration, with only terga I and II dark red and nearly all sternum II black. The most characteristic coloration of metasoma is as follows: segments I and II red, tergum III red basally and laterally, with larger or smaller black area apicomedially and sternum III basally red and apically dark.

Description of male.

Body length: 5.6-8.7 mm.

Head. Mandibular inner margin with one distinct rectangular tooth and distinct furrow next to tooth distally. Clypeus slightly convex, uniformly curved, or more steeply declining ventrally than dorsally, top about in clypeal midlength; basomedian area very large, about two thirds of bevel or more, densely punctate; bevel small, slightly convex, shiny with several larger punctures; lip conspicuously arcuate or slightly sinuate, with small or large lateral corner, lip separated from bevel by distinct groove with large punctures; WML:LCL = 1.2-1.5, WCL:WML = 2.2-2.4. Supraclypeal area groove-like, concave, supraantennal tubercle distinctly elevated ventromedially; surface sparsely punctate, punctures ill defined, interspaces densely microsculptured, slightly shiny to dull. Antenna relatively short, LA3:WA3 = 1.5-1.6, LA5:WA5 = 1.9-2.1. Frons finely punctate, punctures well defined, one to less than half diameter apart, interspaces variable in size; frontal median line distinct, narrow, finely impressed. Vertex punctate, punctures well defined, less than one diameter apart, interspaces microsculptured, slightly shiny. Vertex setae short, semierect, less than 1 × MOD; postocellar impression well developed, open, widely Y-shaped; vertex wider than long; WV:LV = 1.4-1.5. Gena dorsally well developed.

Mesosoma. Scutum without distinct anterior impression; scutum and scutellum densely punctate, punctures well defined, most punctures less than half diameter apart, interspaces distinct, microsculptured, slightly shiny to dull, setae about 1.0 × MOD or less. Mesopleuron coarsely microsculptured with distinct or indistinct punctures in lower areas, most punctures ill defined, interspaces dull; hypoepimeral area coarsely microsculptured to finely rugose, dull; ventral part of mesopleuron with small, ill defined punctures, interspaces slightly shiny. Propodeal dorsum variably sculptured, irregularly ridged, areolate, or coarsely irregularly rugose, without distinct longitudinal ridges, short longitudinal ridges developed basally; propodeal side distinctly ridged, more densely and finely ridged basally than in the center in most specimens; posteromedial margin of propodeal dorsum elevated, but not produced between marginal ridges, marginal ridges positioned nearly horizontally above groove on posterior side. Legs densely punctate, punctures small; forefemoral notch light red and large, its diameter usually longer than space that separates it from forefemoral base, central part of notch slightly elevated, anterior and posterior margins lined by small distinct ledge, notch surface without distinct setae, microsculptured, dull. Wings slightly infumate, veins brown.

Metasoma. Terga I-III with silvery, apical fasciae. Apical depressions of all terga distinct, with micropunctures similar as on central part of terga. Terga I-III densely and distinctly micropunctate, punctures ill defined, interspaces microsculptured, slightly shiny; sculpture of terga IV-VII slightly coarser and slightly more densely punctate than other terga. Sterna uniformly punctate nearly like terga.

Coloration. Apical part of mandibles, terga and sterna I and II and forefemoral notch red. Metasoma very variable in coloration; some specimens with only tergum I dark red, or terga I and II and sternum II basally red, or terga and sterna I-III equally red. Distal tarsomeres dark reddish. Tegulae brown to reddish. Apical parts of all terga and tegulae translucent. Remaining body parts black.

Geographic distribution.

Austria, France, Germany, Hungary, Poland, Slovakia and Sweden.