Horologionini Jeannel, 1949, 1932

Harden, Curt W., Davidson, Robert L., Malabad, Thomas E., Caterino, Michael S. & Maddison, David R., 2024, Phylogenetic systematics of the enigmatic genus Horologion Valentine, 1932 (Coleoptera, Carabidae, Trechinae, Horologionini), with description of a new species from Bath County, Virginia, Subterranean Biology 48, pp. 1-49 : 1

Horologionini Jeannel, 1949

Horologionidae Jeannel, 1949: 91. Type genus: Horologion Valentine, 1932.

Horologionini : Barr (1964: 1).

Horologionina : Bousquet (2012: 711); Ortuño and Arillo (2015: 584), Maddison et al. (2019: 171).

Diagnosis.

The tribe consists of a single genus, Horologion , described in detail below. Members of the tribe possess characteristics of the supertribe Trechitae and several character states that, in combination, distinguish them from other tribes of Carabidae : eyes absent, frontal furrows weakly impressed, penultimate maxillary palpomeres glabrous, apical maxillary palpomeres elongate and narrow, pronotal lateral margins and hind angles absent, elytral humeri with carinate shelf, elytron with a single discal setigerous puncture, apical recurrent stria absent, mesoventrite bell-shaped and extended anteriorly, mesocoxae and metacoxae separated by processes of mesoventrite and metaventrite, all surfaces of external integument except for mouthparts and abdominal tergites generally setose, median lobe of aedeagus open dorsally, spermatheca with attached gland, female genital segment subtriangular, gonocoxites slender and largely glabrous.