Miliusa, Lesch. ex De Candolle, 1832

Damthongdee, Anissara, Sinbumroong, Aroon, Rueangruea, Sukid, Kaitongsuk, Saksan, Ue-Aree, Phasit, Jongsook, Hathaichanok & Chaowasku, Tanawat, 2022, Miliusa microphylla (Annonaceae), a new species from Thailand as evidenced by morphology and plastid phylogeny, with M. glandulifera, a new record for Thailand, Phytotaxa 532 (3), pp. 259-274 : 269

Revised key to the species of Miliusa in clade C in Thailand

1. Leaf base cuneate, obtuse to rounded, generally slightly unequal, not clasping twigs ......................................................................2

- Leaf base subcordate to cordate, slightly to notably unequal, not or (slightly to distinctly) clasping twigs......................................4

2. Leaf blade 3.8–5.6 cm wide; flowering pedicels (18–) 22–27 mm long; inner petals 6.5–7.5 × 5.5–8 mm ............... M. glandulifera

- Leaf blade 1.1–3.2 cm wide; flowering pedicels ≤ 15 mm long; inner petals ≤ 3.5 mm long and wide ...........................................3

3. Plants semideciduous or deciduous; bud scales present; inflorescences borne on new sprouting shoots; inner petals densely hairy inside (mostly near margin and on apical half), glandular structures at base semicircular; stamens ca. 16 per flower... M. fragrans

- Plants evergreen; bud scales absent; inflorescences not borne on new sprouting shoots; inner petals glabrous inside, glandular structures at base in middle between crescent-shaped and semicircular; stamens (6–)8–13 per flower............................... M. fusca

4. Leaf blade 9.3–26 × 4–11 cm; inner petals 5–7.3 × 5–7.5 mm ............................................................................... M. amplexicaulis

- Leaf blade 1.8–11 × 0.9–3.5 cm; inner petals ≤ 4 mm long and wide ...............................................................................................5

5. Lower leaf surface densely hairy........................................................................................................................................................6

- Lower leaf surface glabrous to almost glabrous.................................................................................................................................7

6. Flowering pedicels 12–15 mm long; glandular structures inside inner petals distinct (brownish black in living plants), ± crescentshaped, located about 1 mm above base; with short intermixed with long dense hairs on margin (and area near margin on adaxial side) of inner petals (best observable on apical half) ................................................................................................. M. microphylla

- Flowering pedicels 4–10 mm long; glandular structures inside inner petals less distinct (± light brown in living plants), ± semicircular, located at base; with only short dense hairs on margin (and apical area near margin on adaxial side) of inner petals.. .............................................................................................................................................................................................. M. mollis

7. Carpels ≥ 10 per flower; stipe 5–7 mm long ............................................................................................................. M. nakhonsiana

- Carpels 3–4 per flower; stipe ≤ 1.5 mm long .....................................................................................................................................8

8. Leaf blade usually narrowly elliptic-oblong, base always distinctly clasping twigs; flower buds pink-orange in living plants; monocarps globose, nearly sessile....................................................................................................................................... M. sessilis

- Leaf blade elliptic to elliptic-ovate, base often only slightly clasping twigs; flower buds pale green in living plants; monocarps subglobose, stipe 1–1.5 mm long ................................................................................................................................. M. intermedia