Oryctini, Mulsant & Rey, 1842

Gasca-Álvarez, Héctor & Amat-García, Germán, 2010, Synopsis and key to the genera of Dynastinae (Coleoptera, Scarabaeoidea, Scarabaeidae) of Colombia, ZooKeys 34 (34), pp. 153-192 : 164

Key to the genera of adult Oryctini of Colombia

1. Protibia tridentate...................................................................................... 2

– Protibia quadridentate ............................................................................... 3

2. Elytra smooth, black, shiny. Clypeus emarginate or truncate (bidentate in M. philoctetes ). Mandibles bidentate (Fig. 17).......................... Megaceras Hope

– Elytra with 5 distinct rows of punctures on relatively smooth surface. Clypeus with apex sharply bidentate. Mandibles tridentate (Fig. 18) .... Coelosis Hope

3. Elytra with deeply furrowed rows of punctures. Mandibles broad, with 2 lobes, strongly projecting from beneath clypeus. Clypeus with conical tubercle on dorsal surface. Pronotal fovea of males broadly triangular, extending almost to posterior margin of pronotum ..................... Gibboryctes Endrődi

– Elytra smooth, rugose, or with punctures, never with deeply furrowed rows of punctures. Mandibles variably toothed, never with 2 large lobes. Head with 1–2 conical tubercles in frontoclypeal region, never with 1 on top of clypeus. Males with pronotal fovea variable in form ................................................ 4

4. Body form elongate, subparallel. Protibia with teeth projecting almost at right angles. Apex of metatibia with 2 strong teeth. Males with anterior half of pronotum nearly smooth and with single, median horn or tubercle (Fig. 19). Females without fovea on pronotum ....................... Podischnus Burmeister

– Body form broader, sides rounded (not subparallel). Protibia with teeth projecting obliquely. Apex of metatibia crenulate or with 1, 3, or 4 teeth. Males with anterior half of pronotum densely punctate or rugose or, if nearly smooth, then with lateral horns or tubercles. Females with or without fovea on pronotum ............................................................................................. 5

5. Both males and females with head horn. Prosternal process short, not produced beyond middle of protrochanters. Pronotum with anterior margin distinctly emarginate at center ( Fig. 20)....................................... Enema Hope

– Males with or without head horn; females never with head horn, instead tuberculate at most. Prosternal process long, produced beyond middle of protrochanters. Anterior margin of pronotum lacking emargination at center. 6

6. Frons in males and females unarmed or with 2 tubercles, never with horn on the head. Mandibles tridentate. Pronotum in males with subapical horn and usually with lateral horn or elevated, triangular ridge on each side (Fig 21). Females with fovea in anterior third of pronotum ............... Strategus Kirby

– Males usually with distinct head horn; females with single tubercle. Mandibles without teeth, or with 1–2 rounded teeth. Pronotum in males with horn or prominence arising from posterior half and with or without lateral horns; females usually lacking pronotal fovea.............. Heterogomphus Burmeister