Euphoria candezei Species-Group

Orozco, Jesús, 2012, Monographic Revision of the American Genus Euphoria Burmeister, 1842 (Coleoptera: Scarabaeidae: Cetoniinae), The Coleopterists Bulletin (mo 11) 66, pp. 1-182 : 52-53

Euphoria candezei Species-Group

(Appendix 3: Plate 2j–r )

The species in this group are distinguished by the strongly raised clypeal sides, depressed frons, impunctate scutellum, strongly to moderately emarginate base of the pronotum, mesometasternal process extending anteriorly well beyond the mesocoxae, metatarsi sexually dimorphic in length (shorter than metatibiae and compressed in females, slightly shorter or longer in males), abdomen of males without a deep, longitudinal depression, and distinct form of the parameres. The lateral margins of the pronotum are sexually dimorphic in this group, being more evident in some species (i. e., Euphoria westermanni (Gory and Percheron)) and vague or absent in some specimens at times. The lateral pronotal margins in the females are frequently angulate, appearing subparallel at the base. Males have the lateral pronotal margins more evenly rounded, appearing convergent at the base.

Composition. Three species are included: E. candezei Janson , E. lesueuri (Gory and Percheron) , and E. westermanni (Gory and Percheron) .

KEY TO THE SPECIES OF THE CANDEZEI SPECIES- GROUP

1. Vertex with central protuberance. Antennal club in males frequently as long as head in lateral view. Parameres as in Fig. 24c ........................ E. westermanni (Gory and Percheron) (p. 56)

1′. Vertex flat at middle, without central protuberance. Antennal club in males shorter than head in lateral view......................................2

2. Pronotum tomentous in males ( Fig. 23a, d ), shiny in females ( Fig. 23e ). Elytra tomentous, frequently with whitish, cretaceous markings. Parameres as in Fig. 23c .................................... .... E. lesueuri (Gory and Percheron) (p. 54)

2′. Pronotum and elytra never tomentous. Elytra without cretaceous markings ( Fig. 22a, d ). Parameres as in Fig. 22c ................................ ........................... E. candezei Janson (p. 53)

CLAVE PARA LAS ESPECIES DEL GRUPO CANDEZEI

1. Vertex con protuberancia central. Maza antenal en machos frecuentemente tan larga como la cabeza en vista lateral. Parámeros como en la Fig. 24c .............. E. westermanni (Gory y Percheron) (p. 56)

1′. Vertex plano en el medio, sin protuberancia central. Maza antenal en machos más corta que la cabeza en vista lateral.......................2

2. Pronoto tomentoso en machos ( Fig. 23a, d ), brillante en hembras ( Fig. 23e ). Élitros tomentosos, frecuentemente con máculas cretáceas blanquecinas. Parámeros como en la Fig. 23c ........................................................ ..... E. lesueuri (Gory y Percheron) (p. 54)

2′. Pronoto y élitros nunca tomentosos. Élitros sin máculas cretáceas ( Fig. 22a, d ). Parámeros como en la Fig. 22c ........................................ ........................... E. candezei Janson (p. 53)

Figures

Plate 2. Euphoria species. herbacea species-group: a, b) E. herbacea, c, d) E. leprosa, e, f, g) E. steinheili, h, i) E. subguttata. candezei species-group: j, k) E. candezei, l, m, n) E. lesueuri, o, p, q, r) E. westermanni. hera speciesgroup: s, t, u, v) E. hera.

Fig. 24. Euphoria westermanni. a) Habitus, b) Head, c) Parameres, d) Color variation, e-f) Females, g) Temporal distribution, h) Geographic distribution in Mexico.

Fig. 23. Euphoria lesueuri. a) Habitus, b) Head, c) Parameres, d) Color variation, e) Female, f) Temporal distribution, g) Geographic distribution in Belize, El Salvador, Guatemala, Honduras, Mexico, and Nicaragua; open square indicates country record only.

Fig. 22. Euphoria candezei. a) Habitus, b) Head, c) Parameres, d) Color variation, e) Temporal distribution, f) Geographic distribution in Costa Rica, Guatemala, Honduras, Mexico, Nicaragua, and Panama; open square indicates state record only.