Tolpia kohkonga, Han & Kononenko, 2023

Han, Hui Lin & Kononenko, Vladimir S., 2023, A review of the genus Tolpia Walker, 1863 in East Asia with descriptions of five new species (Lepidoptera, Erebidae, Hypenodinae, Micronoctuini). Taxonomic study of Micronoctuini. Contribution VII, Zootaxa 5227 (5), pp. 549-567 : 556

Tolpia kohkonga sp. n.

( Figs. 18 , 37 )

Type materials. Holotype: male, Cambodia, Prov. Kohkong, 7.ii.2014, Y.S. Bae, genit. prep. HHL-3528-1, coll. NEFU . Paratypes: 1 male, Cambodia, Pursat, 2.v.2010, Y.S. Bae et al., coll. NEFU ; 1 male, Cambodia, Cardamom , 29.vi.2011, Y.S. Bae et al., coll. NEFU ; 1 male, Cambodia, Seima , 1-3.vii.2011, Y.S. Bae et al. coll. NEFU ; 1 male, same data as holotype, genit. prep. HHL-3624 (in tube), coll. NEFU .

Diagnosis. This species is superficially similar to T. multiprocessa ( Fig. 17 ), but differs in the following characters: The head is pale yellowish grey, the thorax light brown and the abdomen brown to ochre brown (in T. multiprocessa the head is fuliginous, the thorax is dark brown, and the abdomen blackish brown); the wingspan is 13.5- 15 mm, the forewing is broad, brownish grey (in T. multiprocessa the wingspan is 13 mm, the forewing is narrower, dark brown); the basal line bears a black dot on the costal vein (in T. multiprocessa no black dot is absent); the antemedial region is diffused with smokey coloration at the inner margin (in T. multiprocessa the diffusion is black); the reniform spot is indistinct (in T. multiprocessa it is black, distinct); the terminal line is separated into segments, appearing as a broken line (in T. multiprocessa it is as a row of dots); the hindwing is light brown to brown (in T. multiprocessa it is dark brown, mixed with blackish, with brown and black grain at the inner area). In the male genitalia the saccus is rather long and narrow, V-shaped (in T. multiprocessa ( Fig. 36 ) it is short, and broad, also V-shaped); the tegumen is narrow (in T. multiprocessa it is more expansive); the ampulla bears two processes on the left side, the inner one large, horn-like, curved, and the outer process small and thorn-like (in T. multiprocessa the inner process is slightly hastate with the inner margin entire, the outer process straight and dentate); the juxta is sclerotized, tongue-like, with a narrow finger-like process apically on the left side, and with a circular hole on the right (in T. multiprocessa it is disc-shaped, apically with a small and short process at left side, with a circular hole); the phallus is moderately coiled counter-clockwise (from dorsal view) (in T. multiprocessa it is strongly coiled clockwise).

Discription. Adult ( Fig. 18 ). Wingspan 13.5-15.0 mm. Head yellowish grey; thorax light yellowish grey; abdomen dark reddish brown, male with long reddish-brown anal tuft of scales; hind leg with prominent, long scent brush. Forewing broad, light brownish grey, costal margin smoothly arched; basal line appearing as two black dots on costal margin and costal vein; antemedial line slender, reddish brown, appearing as black dot at costal margin, strongly curved in costal margin, then oblique; median line absent; postmedial line slender, reddish-brown, black and broader at costal margin, slightly waved, distinct; subterminal line indistinct, slightly waved, somewhat darker than ground colour; terminal line appearing as a broken line; fringe same as ground colour; orbicular and reniform spots indistinct. Hindwing darker than forewing, with brownish-black inner margin; discal spot indistinct; outer margin emarginate between vein M 3 and anal angle; fringe same as ground colour.

Male genitalia ( Fig. 37 ). Tegumen long and thick. Vinculum thick, V-shaped, tapering at tip, with broad saccus, Fultura superior fused with tegumen, weakly sclerotized, with triangular processes, distinct on left side, very small on right side. Juxta sclerotized, tongue-like, with narrow finger-like process apically on left side, and with a circular hole on right side. Valva rather narrow, asymmetrical, right valva somewhat broader than left one; sacculus asymmetrical, left sacculus broad, cone-shaped, about 4/7 length of valva, right sacculus narrower and shorter than left one, triangular cone-shaped, about 2/7 length of valva; process of sacculus finger-shaped with setae on left side, right process shorter than left; costa thick, sclerotized, longer on left valva than on right one; clavus without setose crest, elongate spine on left side, stouter spine on right side; ampulla asymmetrical, with two processes on left side, inner process curved, horn-like, other process small and curved, apically asymmetrical, rounded, shovelshaped, right process larger than left, apically smoothly rounded; digitus curved, pointed apically, left digitus with two processes (inner one stout, outer process slender, incurved, hook-shaped), right digitus with two process (inner one a protuberance, outer process long, incurved, hook-shaped), right process longer than left. Phallus sclerotized, coiled counter-clockwise (from dorsal view), with broad ductus ejaculatorium; coecum broad, cone-shaped, about 1/3 as long as entire phallus.

Etymology. The species name is derived from the type locality of the new species “Kohkong” in Cambodia.

Distribution. Cambodia (Kohkong, Pursat, Seima, Cardamom). The species was collected in the tropical rainforest zone.

Figures

FIGURES 17–24. Tolpia spp., adults. 17. T. multiprocessa, Cambodia, HHL-3531-1; 18. T. kohkonga sp. n., Cambodia, holotype, 3528-1; 19. T. orientis, Thailand, holotype (after Fibiger 2007); 20. T. michaeli sp. n., Malaysia, holotype, HHL- 3610-1; 21. T. alexmadseni, Thailand, holotype (after Fibiger 2007); 22. T. talauti, Malaysia, holotype (after Fibiger 2007); 23. T. palawani, Philippines, Palawan, holotype (after Fibiger 2007); 24. T. conscitulana, Malaysia, HHL-3597-1.

FIGURES33–40. Tolpiaspp.,male genitalia.33. T.conscitulana, Malaysia, genit.prep.HHL-3597-1;34.T.peniculus,Indonesia, holotype (after Fibiger 2007); 35. T. kampungi, Thailand, holotype (after Fibiger 2007); 36. T. multiprocessa, Cambodia, gen. prep. HHL-3531-1; 37. T. kohkonga sp. n., Cambodia, holotype, genit. prep. HHL-3528-1; 38. T. orientis, Thailand, holotype, (after Fibiger 2007); 39. T. michaeli sp. n., Malaysia, holotype genit. prep. HHL-3610-1; 40. T. alexmadseni, Thailand, holotype (after Fibiger 2007).