Tolpia michaeli, Han & Kononenko, 2023

Han, Hui Lin & Kononenko, Vladimir S., 2023, A review of the genus Tolpia Walker, 1863 in East Asia with descriptions of five new species (Lepidoptera, Erebidae, Hypenodinae, Micronoctuini). Taxonomic study of Micronoctuini. Contribution VII, Zootaxa 5227 (5), pp. 549-567 : 557-558

Tolpia michaeli sp. n.

( Figs. 20 , 39 )

Type materials. Holotype: male, Malaysia, Borneo , Jungle Girl camp, 24.iv–2.v.2016, leg. H.L. Han, genit. prep. HHL-3610-1.

Diagnosis. This species superficially is similar to T. orientis Fibiger, 2007 ( Fig. 19 ), but can be distinguished by the following characters: The head and thorax are yellowish grey (reddish brown in T. orientis ); the abdomen is brown to ochreous grey, in the male with long yellowish-grey anal tufts (the abdomen is brown, with long black to brown anal tufts in male T. orientis ); the forewing is broad, greyish brown, wingspan 12.5 mm (in T. orientis the forewing is rather narrower, brown, wingspan 13 mm); the basal line bears a black dot on the costal margin and costal vein (in T. orientis a black dot appears only on the costal margin); the antemedial and postmedial lines are brownish red, distinct (in T. orientis they are indistinct, only appearing as black dots on the costal margin); the terminal line appears as a row of dots (in T. orientis the terminal line is indistinct). In the male genitalia, the saccus is long (in T. orientis ( Fig. 38 ) it is short); the tegumen is broad (in T. orientis it is narrow); the transtilla is bifurcate, with two processes of equal length on the left side, and trifurcate on the right side (in T. orientis the right process is bifurcate, the inner process a prominent thorn, 3X longer than the outer left process); the saccular process is short, smoothly rounded apically (in T. orientis it is long and apically sharp); the juxta is sharply angled at both sides, with a circular hole at the centre (in T. orientis it is smoothly rounded, with a circular hole on the right side).

Description. Adult ( Fig. 20 ). Wingspan 12.5 mm. Head and thorax yellowish grey; abdomen brown to ochre grey, male with long yellowish grey anal tuft. Forewing broad, greyish brown, costal margin smoothly arched basally; basal line black, only appearing as two dots on costal margin and costal vein; antemedial line brownish red, distinct, slender, with black dot on costal margin, indistinct from costal dot, curved to inner vein of cell, then slightly waved and incurved to inner margin; median line absent; postmedial line brownish red, distinct, with black segment in costal area, then slightly waved and incurved to inner margin; subterminal line distinct, broad, waved; terminal line a row of small, black dots; fringe lighter than ground colour; orbicular spot small and black; reniform spot brownish red to ochre; terminal line diffuse, black. Hindwing dark brown, darker than forewing, smoky brown on inner margin; outer margin emarginate between vein M 3 and anal angle; discal spot darker brown, blurred; fringe lighter than ground colour.

Male genitalia ( Fig. 38 ). Tegumen long and broad. Vinculum thick, 7/10 as long as tegumen. Saccus long, Vshaped. Fultura superior fused with tegumen, irregularly fan-shaped on left side. Juxta sclerotized, tongue-shaped, sharply angled on both sides, with circular hole at centre. Valva broad, asymmetrical, right valva broader than left one; sacculus asymmetrical, left sacculus broad and conical, about 1/2 length of valva, right sacculus same length, narrower than left one, slightly shorter than valva; saccular process short, smoothly rounded apically; transtilla bifurcate, sclerotized, very thick basally, with two processes of equal length on left side, right process trifurcate, sclerotized, very thick basally, with three prominent short thorn process of the same length; costal margin thick, sclerotized, of equal length on left and right valvae; clavus small, swollen, with setose crest; ampulla asymmetrical, golf club-like, right ampulla larger than left one, smoothly rounded apically; digitus curved, hook-shaped, pointed apically, left digitus 5/7 as long as right one. Phallus sclerotized, extremely long, extended by ductus ejaculatorium, coiled counter-clockwise; coecum broad, cone-shaped, about 1/3 length of entire phallus.

Etymology. The species name “ michaeli ” is dedicated to the late Dr. Michael Fibiger, a famous and enthusiastic lepidopterist, who first described and recognized Micronoctuini ( Micronoctuidae sensu Fibiger ) as a taxon belonging to the Noctuoidea and developed its classification.

Distribution. Malaysia (Borneo). The species was collected in the tropical rainforest zone.

Figures

FIGURES 17–24. Tolpia spp., adults. 17. T. multiprocessa, Cambodia, HHL-3531-1; 18. T. kohkonga sp. n., Cambodia, holotype, 3528-1; 19. T. orientis, Thailand, holotype (after Fibiger 2007); 20. T. michaeli sp. n., Malaysia, holotype, HHL- 3610-1; 21. T. alexmadseni, Thailand, holotype (after Fibiger 2007); 22. T. talauti, Malaysia, holotype (after Fibiger 2007); 23. T. palawani, Philippines, Palawan, holotype (after Fibiger 2007); 24. T. conscitulana, Malaysia, HHL-3597-1.

FIGURES33–40. Tolpiaspp.,male genitalia.33. T.conscitulana, Malaysia, genit.prep.HHL-3597-1;34.T.peniculus,Indonesia, holotype (after Fibiger 2007); 35. T. kampungi, Thailand, holotype (after Fibiger 2007); 36. T. multiprocessa, Cambodia, gen. prep. HHL-3531-1; 37. T. kohkonga sp. n., Cambodia, holotype, genit. prep. HHL-3528-1; 38. T. orientis, Thailand, holotype, (after Fibiger 2007); 39. T. michaeli sp. n., Malaysia, holotype genit. prep. HHL-3610-1; 40. T. alexmadseni, Thailand, holotype (after Fibiger 2007).