Panamamyia, Woodley, 2008

Woodley, Norman E., 2008, Two new Stratiomyinae, including Panamamyia gen. nov., from the Neotropical Region (Diptera: Stratiomyidae), Zootaxa 1701 (1), pp. 29-39 : 32

Panamamyia gen. nov.

Type species, Panamamyia silbergliedi sp. nov., by present designation.

Diagnosis. I regard the structure of the antennal flagellum ( Figs. 10 , 13 ), with the sixth flagellomere as wide or wider than the fifth, and nearly as wide as flagellomeres 1–4, as autapomorphic for this genus. Also, the moderately elongate abdomen with nearly straight, very gradually posteriorly tapering lateral margins is probably autapomorphic as well ( Figs. 8 , 11 ).

In the key of James & McFadden (1979) this genus keys to Odontomyia Meigen. Odontomyia differs from Panamamyia by having a less strongly produced face, a much more slender sixth antennal flagellomere, and the abdomen less slender with more rounded margins. I regard the reduction in size of the fifth and sixth flagellomeres in Odontomyia as apomorphic for that genus, a character state not found in Panamamyia . Panamamyia also bears some resemblence to Psellidotus Rondani (formerly known as Labostigmina Enderlein ), but the latter has the apical antennal flagellomeres more closely associated and indistinctly separated from each other, and an abdomen similar to that noted for Odontomyia .

Description. Male. Head with eyes strongly holoptic ( Fig. 8 ), upper frons reduced to a very narrow triangle, lower frons and face conically produced with rounded apex, and antenna inserted on dorsal part of this prominence. Ocellar tubercle slightly prominent. Gena only slightly visible in lateral view, occiput not visible in lateral view. Eye bare, ommatidia on upper two-thirds much larger than lower ones ( Fig. 10 ). Antenna ( Figs. 10 , 13 ) slender, pedicel slightly longer than scape; flagellum slender, parallel-sided with six distinct flagellomeres, fifth and sixth not strongly differentiated from preceding ones, sixth subequal to fourth, fifth about one-third length of sixth. Palpus small, cylindrical, about three times longer than its diameter, apparently one-segmented.

Thorax with subscutellum developed, thinly tomentose, and with a few upwardly-directed hairs at lower lateral corners. Scutellum ( Figs. 8 , 11 ) trapezoidal with two short conical spines about one-fourth length of scutellum and separated by about twice the length of one spine. Legs unremarkable, without significant modifications. Wing with microtrichia virtually absent from entire surface; R 2+3 originating well beyond distal margin of discal cell, R 4 absent, crossvein r-m present, short; discal cell small; M 1 and M 3 faint beyond bases at discal cell, no veins reaching posterior margin of wing; about distal third of A 1 faint.

Abdomen ( Fig. 8 ) approximately two times longer than wide, very gradually tapering posteriorly from second segment, and flattened dorsally.

Female. Differs from male as follows: Head ( Figs. 11, 13 ) with eyes smaller than male, strongly dichoptic, upper frons parallel-sided, 0.44 width of head, with medial elevated ridge in lower part that is very narrowly grooved medially ( Fig. 13 ); small depressions present on either side of this ridge, with small protuberances on each side of depressions; lower frons and face with conical projection more developed than in male. Gena and occiput visible in lateral view, forming a broad border on lower and posterior margins of eye that is sharply margined above and more rounded ventrally. Eye bare, ommatidia uniform in size, small.

Abdomen ( Fig. 11 ) slightly more strongly tapering posteriorly. Cercus two-segmented.

Etymology. This genus is named after Panama, the only country from which specimens are known; feminine in gender.

Figures

FIGURES 8–10. Panamamyia silbergliedi, gen. nov., sp. nov., male holotype. 8, dorsal view. 9, lateral view. 10, head and antennae, lateral view.

FIGURES 11–13. Panamamyia silbergliedi, gen. nov., sp. nov., female paratype. 11, dorsal view. 12, lateral view. 13, dorsolateral view of head and antennae.