Panthea greyi Anweiler, 2009

Anweiler, Gary G., 2009, Revision of the New World Panthea Hübner (Lepidoptera, Noctuidae) with descriptions of 5 new species and 2 new subspecies, ZooKeys 9 (9), pp. 97-134 : 115-117

Panthea greyi Anweiler , sp. n.

urn:lsid:zoobank.org:act:0225DA43-EDA5-458B-AE4D-64188BAB8944

Figs. 25-27 , 57 , 68 , 71

Type material. Holotype male – UNITED STATES “AZ [Arizona] Cochise Co. [ County ] 7700 ft / Chiricahua Mtns [Mountains], Onion / Saddle MV lights / 4 Aug 2000 J.B. Walsh leg.”; “ HOLOTYPE / Panthea greyi / Anweiler” [red label]; [31° 56.006' N 109° 15.804' W]. Deposited in CNC . Paratypes (32 ♁, 4 ♀): UNITED STATES. Arizona: Cochise County: same data as holotype (1 ♁) ; Chiricahua Mtns, Pinery Canyon, Upper Camp , 5.vii.1956, 1 ♁, 1 ♀ ); same locality and coll., 6.vii.1956 (1 ♁) ; same locality, 7.vii.1956, Lloyd Martin, John A. Comstock and Will. A. Rees, coll. (2 ♁) ; Chiricahua Mtns, Pinery Canyon , 6800', 26.vii.1965, RH Leuschner, coll. (2 ♁) ; same locality and coll., 26.vi.1992 (1 ♁) ; Pinery Canyon cmpgd, 31.93° N 109.27° W, 6500' 5.viii.08 C. Schmidt and B. Walsh (3 ♁). Apache Co., White Mountains , Greer , 8300', 1-3.vii.1993, R. Leuschner, coll. (2 ♁) ; same locality and coll., 24-25.vii.1965 (2 ♁) ; same locality and coll., 8200', 6.vi.1990 (1 ♁) ; Greer , 28-30.vii.2005, James Adams and Doug Yanega, coll. (1 ♁) ; Junction Hwy. 260-373, 2.vii.1986, Pat Savage, coll. (1 ♁). Graham Co.: Pinaleno Mtns , Mt. Graham , Pine Crest , 7300', 28.vi.1955, Lloyd Martin, John A. Comstock and Will. A. Rees, coll. (1 ♁). Coconino Co. : Oak Creek Canyon, 5000', 6.viii.1986, RH. Leuschner, coll. (1 ♁) ; same locality and coll., 7.viii.1986 (1 ♁) ; same locality and coll., 5500', 23.vii.1989 (1 ♁). Gila Co.: Mogollon Rim, Tonto Creek State Fish Hatchery, 6400', 21.vi.957, Lloyd M. Martin, Robert J. Ford and William A. Rees, coll. (1 ♁). New Mexico: Colfax Co.: Sangre de Cristo Mts. , Cimarron Canyon , 7900', 7.vii.1962, E. and I. Munroe, coll. (1 ♁, 1 ♀) ; same locality and coll., 10.vii.1962 (1 ♁). Otero Co.: 3 mi sw Cloudcroft , 8830', 23.v.2006, Greg Forbes, coll. (1 ♀) ; Sacramento Mtns, Dry Canyon Rd., 2.1 mi. n. Rte 82, 6 mi. east Rt. 244 at Cloudcroft., G. Forbes, coll. (1 ♁). Lincoln Co.: Ruidosa , Cedar Creek campground, 7000', 28.vii.1962, E. and I. Munroe, coll. (1 ♀). Grant Co. : Gila National Forest at Emory Pass , 32°52.5'N 107°45.0'W, 7160', 5.viii.1991, E. Metzler, coll. (1 ♁). Colorado: Larimer Co. : Big Thompson Canyon , 6500', 9.vii.1955, R.H. Leuschner, coll. (1 ♁). Archuleta Co. : Pagosa Springs , 7000', 7.vii.1988, R.H. Leuschner, coll. (1 ♁). Teller Co. : Florrisant , Big Springs Ranch , 14.viii.1960, TC. Emmel, coll. (1 ♁) ; same locality and coll., 12.viii.1960 (1 ♁). Boul- der Co.: Left Hand Canyon, 6000', 22.vii.958, RH. Leuschner, coll. (2 ♁) .

Etymology. I take pleasure in naming this species in honor of the late L. Paul Grey, who encouraged my budding interest in noctuid moths and who sent me the first specimen of this new Panthea .

Diagnosis. Panthea greyi belongs to the P. furcilla species-group, and can be separated from all Panthea with the exception of P. furcilla by the characters listed in the diagnosis for the group, namely the elongate vesica with one terminal spine in the male and the two-chambered corpus bursae in the female. It can be separated from P. furcilla by range, with P. greyi occurring in southwestern United States west of the Great Plains and P. furcilla east of the Great Plains in eastern United States and north of the Great Plains in Canada. Externally, P. greyi is most similar to Panthea virginarius and Panthea acronytoides nigra . P. virginarius occurs west and north of the range of P. greyi . Panthea acronyctoides nigra may occur with P. greyi in northern Colorado where P. greyi can be recognized by the light, banded hindwing and P. acronyctoides by the dark hindwing, as well as by the genitalic characters. Panthea gigantea is a larger species with two cornuti in the male vesica and a single chambered, thin-walled corpus bursae in the female.

Description. Sexes similar and overlapping in size, female only slightly darker than male; male forewing length 19-22.5 mm, female 21-25 mm. Head – Male antenna bipectinate, with pectinations about 2 × as long as width of antennal shaft; female antenna simple; palps reduced, both palps and lower part of frons clothed in dark redbrown or black hairs; remainder of head a mixture of light-gray, dark-brown and black hair-like scales. Thorax – collar, thorax and tegulae a mixture of light gray, dark brown and black hair-like scales; tegulae crossed by two oblique dark bands midway and along outer edge; legs clothed in dense long light gray and dark gray hair, tarsus banded black and white. Dorsal forewing – ground a mix of white or very light gray and dark brownish-black scales, appearing powdery gray; crossed by five black lines; basal line marked by one or two small patches of black scales; antemedial line straight or nearly so, bending distad in fold before continuing to wing margin; medial line most prominent, straight except for bending distad slightly in fold before reaching lower margin; postmedial line narrower, erratic, bent distad at each vein and angling basad below vein CUA1, contacting or closely approaching medial line below veins CuA2 before bending distad to lower margin of wing; subterminal line incomplete, often reduced to a few dark patches of scales near upper margin, erratic, defined by white scales along distal side; fringe dark gray black, checkered with a few white scales at veins; small black bar or crescent marking the end of the cell. Abdomen – clothed in short, stiff dark gray-brown hair, paler at joints. Dorsal hindwing – white with long gray hair-like scales along inner margin, crossed by poorly defined light gray antemedial, medial and postmedial bands and with a narrow dark gray terminal line; fringe checkered with dark gray and white; veins narrowly lined with dark scales. Male genitalia – ( Fig. 57 ) valve simple, elongate, cucullus rounded, clasper a simple prominent blade near apex, about as long as width of valve; tegumen with large triangular earlike subuncal lobes; uncus laterally compressed with crown approximately as high as wide; ending in a rounded terminus resembling a duck’s beak viewed from above; inflated and everted vesica long, narrow, about 3-4 × as long as wide, angled to left midway and armed with a single massive slightly bent or curved terminal cornutus, oriented toward head; ductus seminalis exiting midway at right angle to right. Female genitalia – ( Fig. 68 ) papillae anales a pair of soft curved bands with sparse hairs; posterior and anterior apophyses about equal in size and of average length; sterigma well-developed, average in size and amount of sclerotization; ductus bursae wide, about 2 × as long as wide, mostly heavily sclerotized with deep creases and folds, expanding gradually into posterior section of corpus bursae; corpus bursae slightly constricted midway, forming a thick-walled and partially sclerotized posterior section united with lower half of ductus, and a thin-walled, translucent globular anterior section.

Distribution and biology. Panthea greyi has been collected in the mountains of Arizona, New Mexico, Colorado and southern Utah, at elevations of 1524-2545 m ( Fig. 71 ). Collection dates range from June 6 through September 23. It has been collected in Arizona in oak-pine woodland.

Figures

Figures 12-27. Panthea adults 12. P. f. furcilla ♀, Berkshire Co., MA, USA; 13. P. f. furcilla ♁, holotype of pallescens, Queen’s Co., NS, CAN; 14. P.f.furcilla ♁ Franklin Co., MA, USA; 15. P.f.furcilla ♁ Redwater, AB, CAN; 16. P. f.furcilla ♁, holotype of atrescens, QC, CAN; 17. P. f. furcilla ♁, holotype of centralis, QC, CAN; 18. P. f. furcilla ♁ (melanic), Windham Co., CT., USA; 19. P. f. furcilla – f. australis intermediate ♁, Owsley Co., KY, USA; 20. P. f. furcilla – f. australis intermediate ♁, Owsley Co., KY, USA; 21. P. f. australis ♁ holotype, Whitfield Co, GA, USA; 22. P. f. australis ♀, Whitfield Co, GA, USA; 23. P. f. australis ♀, Oregon Co., MO, USA; 24. P.f.australis ♁, Jefferson Co.TX, USA; 25. P. greyi ♁, Cochise Co., AZ, USA; 26. P. greyi ♁ holotype, Cochise Co., AZ, USA; 27. P. greyi ♀, Cochise Co., AZ, USA.

Figures 50-57. Panthea male genitalia 50. P. virginarius, Josephine Co. OR., USA; 51. P. virginarius Wyoming variant, Lincoln Co., WY., USA; 52. P. acronyctoides nigra, Albany Co., WY, USA; 53. P. a. acronyctoides, Steuben, ME, USA; 54. P. furcilla furcilla, Norfolk Co., MA., USA; 55. P.furcilla australis, Whitfield Co., GA.USA; 56. Panthea f.furcilla - f. australis intermediate specimens, showing variation in size of basal cornuti; a. P.G. Co., MD; b, c. Owsley Co., KY; d. Elko Co., PA. 57. P. greyi. Coconino Co., AZ, USA.

Figures 58-68. Panthea female genitalia 58. P. apanthea, Coconino Co, AZ. USA; 59. P. judyae, Chihuahua, MEX; 60. P. gigantea, Chihuahua, MEX; 61. P. guatemala, Oaxaca, MEX; 62. P. virginarius, Missoula Co., MT., USA; 63. P.acronyctoides nigra, Teller Co., CO., USA;. 64. P. a. acronyctoides, Ashland Co., WI, USA; 65. P. f.furcilla, Windham Co., CT, USA; 66. P. f.furcilla-f. nigra intermediate, Dauphin Co., PA., USA; 67. P. furcilla australis, Whitfield Co., GA, USA; 68. Panthea greyi, Apache Co., AZ., USA.

Fig. 71. Distribution of examined specimens of P. furcilla and P. greyi.