Euploca polyphylla (Lehm.) J.I.M.Melo & Semir, 2009

Feuillet, Christian, 2016, Two new combinations in Euploca Nutt. (Heliotropiaceae, Boraginales) and a conspectus of the species of the Guiana Shield area, PhytoKeys 61, pp. 101-124 : 108

Euploca polyphylla (Lehm.) J.I.M.Melo & Semir, 2009

Euploca polyphylla (Lehm.) J.I.M.Melo & Semir, Kew Bull. 64(2): 289. 2009.

Heliotropium polyphyllum Lehm., Neue Schriften Naturf. Ges. Halle 3(2): 9. 1817.

Euploca polyphylla Type (fide Förther 1998: 214). Venezuela. near Orinoco, Humboldt s.n. (holotype MEL 233310 n.v.; isotypes B-Willd 3252 n.v., P! [scan!], S 11-21939 n.v. [scan!]).

Heliotropium foliosum Roem. & Schult., Syst. Veg. 4: 737. 1819.

Euploca polyphylla Type. Venezuela. Orinoco, Apr 1800, Humboldt and Bonpland 808 (holotype B-WILLD 3252-01 n.v.).

Preslaea linifolia A.St.-Hil., Voy. Distr. Diam. 2: 433. 1833.

Euploca polyphylla Type. Brazil. Cabo Frio, St.-Hilaire 433 (lectotype, designated by Förther 1998, pg. 234: P!; isotypes B †, M n.v., MPU!, P! [2 sheets]).

Heliotropium bahiense DC., Prodr. 9: 544. 1845.

Euploca polyphylla Syntype: Brazil. Blanchet 92 (G n.v., NY!, P!). Type. Brazil. Bahia, 1836, Salzmann 368 (lectotype, designated by Förther 1998, pg. 181: G-DC n.v.; isotypes B †, F n.v., H n.v., HAL n.v., K!, M n.v., MPU!, P! [4 sheets]).

Heliotropium polyphyllum var. blanchetii DC., Prodr. 9: 544. 1845.

Euploca polyphylla Type. Brazil. Bahia, Serra Jacobina, 1837, Blanchet 2651 (lectotype, designated by Förther 1998, pg. 214: G-DC n.v.; isotypes B† [photo F-17340, GH, NY, US], BM n.v. [fragment GH n.v.], BR n.v., F-520830 n.v. [photo F-57735], FI-W!, HAL n.v., K! [scan!], LE n.v., M n.v., NY!, P! [2 sheets], TUB n.v., US-292391!, W-118162 n.v.).

Schleidenia linifolia (A.St.-Hil.) DC., Prodr. 9: 558. 1845. Type. Based on Preslaea linifolia A.St.-Hil.

Schleidenia pubescens Fresen., Fl. Bras. 8(1): 35. 1857.

Euploca polyphylla Type. Brazil. Rio de Janeiro, Gaudichaud [524] (lectotype, designated by Förther 1998, pg. 236: P!; isotype B†).

Schleidenia polyphylla (Lehm.) Fresen., Fl. Bras. 8(1): 36. 1857. Type. Based on Heliotropium polyphyllum Lehm.

Schleidenia polyphylla var. blanchetii (DC.) Fresen., Fl. Bras. 8(1): 36. 1857. Type. Based on Heliotropium polyphyllum var. blanchetii DC.

Schleidenia bahiensis (DC.) Fresen., Fl. Bras. 8(1): 43. 1857. Type. Based on Heliotropium bahiense DC.

Heliotropium polyphyllum var. laevenworthii A.Gray, Syn. Fl. N. Amer. 2(1): 185. 1878.

Euploca polyphylla Syntypes: USA. Florida: Everglades, Leavenworth s.n.; Palmer s.n. (F). Type. USA. Florida: Everglades, 1877, Garber s.n. (lectotype, designated by Förther 1998, pg. 214: GH n.v.; isotype K n.v. [scan!]).

Heliotropium pubescens (Fresen.) Gürke, in Engler and Prantl, Nat. Pflanzenfam. 4(3a): 97. 1893. Type. Based on Schleidenia pubescens Fresen.

Heliotropium linifolium (A.St.-Hil.) Gürke, in Engler and Prantl, Nat. Pflanzenfam. 4(3a): 97. 1893; illegitimate: not Lehm. 1818. Type. Based on Preslaea linifolia A.St.-Hil.

Heliotropium laevenworthii (A.Gray) Small, Fl. S. E. U.S.: 1006. 1903. Type. Based on Heliotropium polyphyllum var. laevenworthii A.Gray

Heliotropium horizontale Small, Bull. New York Bot. Gard. 3: 435. 1905.

Euploca polyphylla Type. USA. Florida: 9-12 Nov 1903, Small and Carter 742 (holotype NY! [photo GH]; isotypes F n.v., GH n.v. [fragment], US!).

Heliotropium polyphyllum var. horizontale (Small) R.W.Long, Rhodora 72: 33. 1970. Type. Based on Heliotropium horizontale Small

Type.

Based on Heliotropium polyphyllum Lehm.

Description.

Herb or subshrub. Stems prostrate or decumbent, sericeous. Leaves alternate or subopposite: petiole 0.03-0.1 cm long, sericeous; lamina narrow lanceolate or oblanceolate, 0.6-1.3 × 0.1-0.2 cm, apex acute, base cuneate, margin entire, both surfaces sericeous. Inflorescences single or paired, cymose, 1-15 cm long, bracts ovate, 0.3-0.4 cm long, sericeous inside, glabrous outside. Flower sub-sessile: calyx slightly shorter than the corolla tube, lobes, 3-4 mm long at anthesis, glabrous inside, sericeous outside; corolla yellow or white, tubular, outside sericeous, inside sericeous at throat, tube widened in the middle, lobes ovate-deltoid, with narrow sinuses, 2.5-3 mm long; stamens subsessile, anther with a long appendage; ovary subglobose, 4-sulcate, glabrous; style 0.3-0.6 mm long; stigma conical. Fruit subglobose, entirely covered by the calyx; nutlets almost 1.5 mm diam.

Distribution.

Venezuela ( Bolívar), Brazil ( Amapá, Amazonas, Pará); also USA (Florida), Bahamas, Venezuela, eastern Brazil.