Issikiomartyria Hashimoto, 2006

Imada, Yume & Kato, Makoto, 2018, Descriptions of new species of Issikiomartyria (Lepidoptera, Micropterigidae) and a new genus Melinopteryx gen. n. with two new species from Japan, Zoosystematics and Evolution 94 (2), pp. 1-25 : 6-7

Genus Issikiomartyria Hashimoto, 2006

Type species.

Neomicropteryx nudata Issiki, 1953, fixed by original designation.

Diagnosis.

Aedeagus with two pairs of dorsal fins.

Description.

The generic description is based on I. nudata (Issiki, 1953), I. akemiae , I. plicata , I. distincta Hashimoto, 2006, I. bisegmentata , I. hyperborea sp. n., I. leptobelos sp. n., I. catapasta sp. n., I. trochos sp. n., and on the previous studies on this group ( Issiki 1953, Hashimoto 2006).

Head capsule densely covered by microtrichia, apart from genal area where glossy and naked; most of clypeus, frons, and vertex covered with brownish yellow piliform scales. Ocelli present. Antenna moniliform, approximately as long as forewing in male, longer than in female; scape the largest segment, twice longer than most basal flagellum; pedicel bulbous, larger than most basal flagellum; basal one or two flagellomeres cylindrical. SOI about 0.4. MIOI about 0.5. Interocellar sulcus almost complete. Postinterocellar sulcus distinct. Epicranial sulcus distinct between occipital foramen and postinterocellar sulcus, being as a short distance anterior to interocellar sulcus. Temporal sulcus as a darker line. Occipital sulcus almost complete, but slightly indistinct on dorso-lateral corner. Occipus fan-shaped. Mandibular teeth greatly reduced. Labial palp 1- or 2-segmented. Maxillary palp 5-segmented. Proximal prelabium obscure. Foretibial epiphysis absent. Antero-lateral processes of pronotum present, strongly sclerotized. Fore- and hindwings obtuse at apex, forewing with brown to purple luster, without any distinct maculation. Forewing with R1 unforked; R3 stalked with R4+5. Hindwing with a main stem of R absent; most anterior vein of hindwing forked near terminal end (Sc1 and Sc2 + R1). Sternum V gland present; orifice of gland a narrow slit.

Male abdomen and genitalia. Sternum VIII membranous. Segment IX a complete ring, well sclerotized, with a posterior expansion dorsally; posterior margin gradually expanded from dorsum to venter. Valva triangular, broadly membranous at inner surface, with a proximo-ventral ridge whose anterior portion fused with median plate; median plate large, roughly fan-shaped. Phallobase strongly curved, with or without longitudinal ventral ridge(s) on midline. Aedeagus with acute apex, ventrally forked slightly at caudal end; with two pairs of basal fins dorso-medially and a pair of lateral triangular fins; gonopore opening horizontally; vesica with serrate minute projections. Tergum X, broader than long, with a pair of long ventral plates (venter X plates) extending antero-ventrally at base of terminal processes.

Female abdomen and genitalia. Segment IX forming a complete ring, strongly sclerotized; anterior margin gradually expanded anteriorly from dorsum to venter; mid-dorsal length generally shorter than 2/5 of mid-ventral length; laterally protruded in some species. Segment X consisting of a pair of lateral sclerites and a dorsal sclerotized plate; lateral sclerites simple, broader than long, with digitate projections having an apical seta at terminal inner margin. Corpus bursae large, globular, membraneous, with signa composed of four sclerites. Ductus spermathecae arising from a hexagonal or round concavity. Genital chamber with small sclerite(s).

Comparative Remarks.

The following characters are regarded as synapomorphies of Issikiomartyia : aedeagus with two pairs of hornlike dorsal projections and without any protrusion vertically in male: sclerite in female genital chamber greatly reduced.