Bracon (Lucobracon) erraticus Wesmael, 1838

Papp, Jenő & Xviii, Budapest, 2012, A revision of the Bracon Fabricius species in Wesmael’s collection deposited in Brussels (Hymenoptera: Braconidae: Braconinae), European Journal of Taxonomy 21, pp. 1-154: 20-27

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http://dx.doi.org/10.5852/ejt.2012.21

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scientific name

Bracon (Lucobracon) erraticus Wesmael, 1838
status

 

Bracon (Lucobracon) erraticus Wesmael, 1838  

Figs 8 View Fig A-L, 9A-G

Braco erraticus Wesmael, 1838: 35   ♀♁ (type material: three ♀♀ + two ♁♁), type locality: “environs de Bruxelles ” ( Belgium), ♀ lectotype (and two ♀ + two ♁ paralectotypes, present designations) in the Royal Belgian Institute of Natural Sciences, Brussels; examined.

Bracon (Lucobracon) bellicosus Papp, 1971a: 339   (description) ♀, type locality: “ Uvs aimak: on eastern slope of Ulaan davaa pass, between Lake Örög nuur and town Ulaangom” ( Mongolia), ♀ holotype and two ♀ paratypes in Magyar Természettudományi Múzeum, Budapest; examined, syn. nov.

Bracon (Bracon) confinis Szépligeti, 1901: 261   (in key), 276 (description) (in Hungarian) and 1904 (1901): 161 (in key), 169 (description) (in German) ♀ (type material: 1 ♀), type locality: “Pilismarót” ( Hungary), ♀ lectotype in Magyar Természettudományi Múzeum, Budapest; examined.

Bracon congruus Szépligeti, 1901: 261   (♀), 263 (♁) (in key), 276 (description) (in Hungarian); 1904 (1901): 161 (♀), 164 (♁) (in key), 165 (description) (in German) ♀ ♁ (syntype series 2 ♀♀ + 2 ♁♁), type locality: “ Budapest ” ( Hungary), ♀ lectotype (and one ♀ and two ♁ paralectotypes, present designation) in Magyar Természettudományi Múzeum, Budapest; examined.

Bracon erythrostictus Marshall, 1885: 14   (in key) and 17 (description) ♀ ♁ (type material: several ♀♀ and ♁♁), type locality: “Milford heaven” (Pembrokeshire, England), syntypes in The Natural History Museum, London; not examined.

Bracon exarator Marshall, 1885: 14   (in key) and 26 (description) ♀ (type material 1 ♀), type locality: “at Brundall, Norfolk ” ( England), ♀ holotype (present designation) in Castle Museum, Norwich; examined, syn. nov.

Bracon foveola Thomson, 1894: 1819   ♀, type locality: “Ringjön... Skåne ” ( Sweden), ♀ lectotype (and one ♀ paralectotype, designated by Papp in 1969) in Zoological Museum, Lund; synonymized by Papp 1969: 197.

Bracon hades Papp, 1965: 409   (description) ♀, type locality: “Somlóvásárhely, Com. Veszprém, Hungary ”, ♀ holotype (and one ♀ paratype) in Magyar Természettudományi Múzeum, Budapest; examined.

Bracon praetermissus Marshall, 1885: 15   (in key) and 37 (description) ♀ ♁ (type material 5 ♀♀ + 1 ♁, seen 3 ♀♀), type locality: “Nunton Wilts.”[hire] ( England), ♀ lectotype (and two ♀ paralectotypes, present designations) in The Natural History Museum, London; examined, syn. nov.

Braco superciliosus Wesmael, 1838: 38   ♀ (type material: one ♀), type locality: “environs de Bruxelles ” ( Belgium), ♀ holotype (“J’ai pris un seul individu de cette espèce”, present designation) in the Royal Belgian Institute of Natural Sciences, Brussels; examined.

Bracon vectensis Marshall, 1885: 15   (in key) and 31 (description) ♁ (type material 1 ♁), type locality: “Niton, Isle of Wight ” ( England), ♁ holotype (present designation) in The Natural History Museum, London; examined, syn. nov.

Further synonyms of B. erraticus Wesmael   (see also Papp 2005a: 199 and Tobias 1986: 149): aestivalis Szépligeti, 1901, confinis Szépligeti, 1901   , congruus Szépligeti, 1901   ,? idrianus Fahringer, 1934, lagodechianus Telenga, 1936, planiceps Telenga, 1936, secernendus Schulz, 1906   , segregatus Telenga, 1936, similis Szépligeti, 1901, talitzkii Telenga, 1936, transcaspicus Telenga, 1936, ventricosus Szépligeti, 1901).

Bracon erraticus   – Szépligeti 1901: 261, 263, 266 (in key, in Hungarian); 1904 (1901): 180 (in key). Bracon (Orthobracon) erraticus   – Fahringer 1927: 270 (♀), 278 (♁) (in key) and 381 (redescription), assigned to “Section Orthobracon   ”. — Telenga 1936: 164 (♀), 169 (♁) (in key), 257 (redescription) (in Russian) and 367 (♀), 371 (♁) (in key, in German). — Shenefelt 1978: 1630 (literature up to 1975). Bracon (Lucobracon) erraticus   – Tobias 1986: 145 and 149 (in key, in Russian). — Tobias & Belokobylskij 2000: 162 and 164 (in key, in Russian). Bracon (Lucobracon) bellicosus   – Shenefelt 1978: 1616 (as valid species, literature up to 1974). — Tobias & Belokobylskij 2000: 164 (synonymized with question mark with B. erraticus   ). Bracon (Bracon) confinis   – Fahringer 1927: as valid species 242 (♀ in key) and 313 (redescription), assigned to “Section Striobracon)”. — Telenga 1936: 165 (♀, in key), 259 (as valid species, redescription, in Russian) and 367 (♀, in key, in German). — Shenefelt 1978: 1628 (as valid species, literature up to 1968). — Tobias 1986: 149 (as new synonym of B. erraticus   ). — Papp 2004: 173 (as synonym of B. erraticus   , type depository). Bracon congruus   – Fahringer 1927: 242 (♀ in key) and 338 (redescription), as synonym of B. secernendus Schulz, 1906   and assigned to “Section Striobracon”. — Telenga 1936: 258 (as synonym of B. secernendus Schulz   ). — Shenefelt 1978: 1629 (as synonym of B. confinis Szépligeti   after Papp 1968: 197, literature up to 1968). — Papp 2004: 173 (type depository). Bracon erythrostictus   – Fahringer 1927: 237 (♀ in key), 316 (redescription) and 425: ♁ identical with B. tenuicornis Wesmael   , ♀ as valid species and assigned to “Section Striobracon”. — Telenga 1936: 162 (♀), 167 (♁) (in key), 249 (redescription) (in Russian) and 364 (♀), 369 (♁) (in key, in German). — Shenefelt 1978: 1481 (as valid species, literature up to 1974) and 1649 (as synonym of B. tenuicornis Wesmael   after Dalla Torre 1898: 291). — Tobias 1986: 125 (in key, as synonym of B. intercessor Nees   ). Bracon exarator   – Fahringer 1927: as valid species 253 (♀), 257 (♁) (in key) and 350 (redescription), assigned to “Section Lucobracon   ”. — Telenga 1936: as valid species 164 (♀), 169 (♁) (in key), 257 (redescription) and 367 (♀), 371 (♁) (in key, in German), assigned to subgenus Bracon   s. str.. Shenefelt 1978: 1618 (as valid species, literature up to 1945). Bracon foveola   – Fahringer 1927: as valid species 273 (♀ in key) and 388 (redescription), assigned to “Section Orthobracon   ”. — Telenga 1936: as valid species 170 (♀, in key), 270 (redescription) (in Russian) and 373 (♀ in key, in German), assigned to subgenus Orthobracon   . — Shenefelt 1978: 1631 (as B. erraticus var. foveola   after Papp l.c., literature up to 1969). Bracon hades   – Papp 1969: 197 (synonymyzation). — Shenefelt 1978: 1631 (as synonym of B. erraticus var. superciliosus Wesmael   after Papp l.c). Bracon praetermissus   – Fahringer 1927: (♀, identical with “? erraticus Wesm.   var.”), 278 (♁ in key) and 412 (redescription), ♁ as valid species and assigned to “Section Orthobracon   ”. — Telenga 1936: as valid species 177 (♀), 178 (♁) (in key), 300 (redescription) and 379 (♀), 381 (♁) (in key, in German). — Shenefelt 1978: 1645 (as valid species, literature up to 1974). Tobias 1986: as valid species 137 (in key, in Russian). — Tobias & Belokobylskij 2000: as valid species 150 (in key, in Russian). Braco superciliosus   – Schmiedeknecht 1897: 559 (15) (synonymization). — Fahringer 1927: 381 (as synonym of B. erraticus   ). — Telenga 1936: 257 (as synonym of B. erraticus   ). — Shenefelt 1978: 1631 (as B. erraticus var. superciliosus Wesmael   after Fischer 1965: 130, literature up to 1974). Bracon vectensis   – Fahringer 1927: as valid species 278 (♀ in key) and 430 (redescription), assigned to “Section Orthobracon   )”. — Telenga 1936: as valid species 178 (♁, in key), 301 (redescription) (in Russian) and 381 (♁, in key, in German). — Shenefelt 1978: 1650 (as valid species, literature up to 1936).

Designation of the types

Designation of the ♀ lectotype of Bracon erraticus   (first label, printed) “ Coll. Wesmael ”, (second label, printed) “2040”, (third label) “ Braco erraticus   mihi ♀ ” (handwritten) “dét. C. Wesmael ” (printed), (fourth label, printed) “ Type ”, (fifth label with my handwriting) “ Belgique / Bruxelles / leg. Wesmael ” (above) “teste J. Papp 1987” (reverse), sixth label is the lectotype card. - Lectotype is in good condition: (1) micropinned by mesosoma, (2) right antenna deficient: with 17 antennomeres, (3) left fore wing close beyond pterostigma slightly torn.

Designations of the three paralectotypes (1 ♀ + 2 ♁♁) of B. erraticus   : with similar labels to that of the lectotype. One ♀ paralectotype is in good condition: flagelli deficient, tarsomeres 3-5 of right fore leg missing. One ♁ paralectotype is in very poor condition: head, right fore and left hind wings missing, mesosoma cracked at prosoma. One ♁ paralectotype is in poor condition: left antenna, left fore wing and metasoma entirely as well as legs partly missing. All three paralectotypes micropinned by mesosoma.

Designation of the ♀ paralectotype of B. erraticus   var. 1. Wesmael with two labels similar to the first and fifth labels of the lectotype. Paralectotype is in fairly poor condition: (1) micropinned by mesosoma, (2) right flagellum missing, left flagellum apically deficient, (3) legs more or less damaged.

Designation of the ♀ holotype of B. exarator Marshall   : (first label, handwritten) “Brundall 15.9.81” (reverse on first label, above on label the wasp itself is glued), (second label, handwritten) “exarator”, third label is the holotype card, fourth label is with the actual name B. erraticus var. superciliosus (Wesmael)   given by me. Holotype is in good condition.

Designation of the ♀ lectotype of B. praetermissus Marshall   : (first label, handwritten) “N [=Nunton, England] 1.IX.84” (reverse on label, above on label the wasp itself is glued), (second label, handwritten) “27 jts”, (third round label with red frame) “ ♀ Type CM.” (♀ and CM with handwriting, Type printed), (third label) “ B.M. Type Hym. ” (printed) “ Bracon praetermissus Marshall, 1885   ” (handscdript), (fourth label,?Marshall’s handwriting) “ praetermissus Marsh.   ”, (fifth label, printed) “ Marshall coll. 1904- 120.”, sixth label is with the full locality ( England, Wilts., Nunton, 1.IX.1884, leg. Marshall) added by me, seventh label is the lectotype card and the eighth label is with the actual name ( B. erraticus   ) given by me. Lectotype is in good condition: specimen glued ventrally and nicely mounted on a card.

Designation of the two ♀ paralectotypes of B. praetermissus   : with similar labels to that of the lectotype; the specimens are representing a variety: B. erraticus var. superciliosus (Wesmael)   . Paralectotypes are in good condition similar to the lectotype.

Remark

One ♀ paralectotype of B. praetermissus   is housed in the Hungarian Natural History Museum, the specimen is representing the species B. romani Fahringer, 1927   .

Designation of the ♀ holotype of B. superciliosus   : (first label, printed) “ Coll. Wesmael ”, (second label, printed) “2052”, (third label) “ Braco superciliosus   mihi ♀ ” (handwritten) “dét. C. Wesmael ”, (fourth label, printed red) “ Type ”, (fifth label with my handwriting) “ Belgique / Bruxelles / VI, leg. Wesmael ” (after Wesmael l.c.), sixth label is the holotype card, seventh label is with the actual name B. erraticus var. superciliosus (Wesmael)   given bu me. Holotype is in fairly good condition: (1) micropinned by mesosoma; (2) left forewing missing; (3) left flagellum apically deficient.

Designation of the ♁ holotype of B. vectensis   : (first label, handwritten) “Niton” (reverse on label, above it the wasp itself is glued), (second label with red frame, printed) “Type”, (third label) “B.M. Type Hym.” (printed) “3.C.2.” (handwritten, (fourh label) “B.M. Hym. Type” (printed) “ Bracon vectensis Marshall 1885   ” (handwritten), (fifth label,?Marshall’s handwriting) “ vectensis Marsh.   ”, (sixth label,

printed) “Marshall coll. 1904-120.” seventh label is the holotype card and eighth label is with the actual name, B. erraticus   , given by me. Holotype is glued by its ventrtal side on a card.

Material examined

201 ♀♀ + 211 ♁♁ from 38 countries. The localities are grouped according to the nominate form and the three varieties of the species.

1) Bracon erraticus Wesmael   , nominate form (62 ♀♀ + 65 ♁♁ from thirty countries): ENGLAND: 6 ♀♀ + 3 ♁♁ from seven localities. SWEDEN: 4 ♀♀ + 2 ♁♁ from six localities. DENMARK: 1 ♀ + 2 ♁ from three localities. FRANCE: 2 ♀♀ + 4 ♁♁ from four localities. THE NETHERLANDS: 2 ♀♀ from two localities. GERMANY: 5 ♀♀ +3 ♁♁ from eight localities. SWITZERLAND: 1 ♁. AUSTRIA: 3 ♀♀ + 2 ♁♁ from five localities. BOHEMIA: 2 ♀♀ + 1 ♁ from three localities. SLOVAKIA: 8 ♀♀ + 7 ♁♁ from 13 localities. HUNGARY: 55 ♀♀ + 81 ♁♁ from 124 localities. ROMANIA: 3 ♀♀ + 6 ♁♁ from seven localities. PORTUGAL: 1 ♁. SPAIN: 5 ♀♀ + 3 ♁♁ from one locality. ITALY: 3 ♀♀ + 2 ♁♁ from five localities. SERBIA: 6 ♀♀ from five localities. ALBANIA: 2 ♁♁ from two localities. MACEDONIA: 3 ♀♀ + 4 ♁♁ from seven localities. CROATIA: 2 ♀♀ + 6 ♁♁ from eight localities. BULGARIA: 3 ♀♀ + 8 ♁♁ from ten localities. GREECE: 1 ♀ + 2 ♁♁ from three localities. TURKEY: 4 ♀♀ + 4 ♁♁ from eight localities. ALGERIA: 1 ♁. SYRIA: 1 ♁. CYPRUS: 1 ♁. ARMENIA: 1 ♁. GEORGIA: 1 ♁. ASIATIC RUSSIA: 2 ♀♀ from two localities. MONGOLIA: 4 ♀♀ + 2 ♁♁ from six localities. NORTH CHINA: 1 ♀ + 2 ♁♁ from one locality.

2) Bracon erraticus var. aestivalis (Szépligeti)   (1 ♀ + 8 ♁♁ from three countries): SLOVAKIA: 1 ♀ + 1 ♁ from two localities. HUNGARY: 6 ♁♁ from five localities. ROMANIA (Transsylvania): 1 ♁.

3) Bracon erraticus var. confinis (Szépligeti)   (70 ♀♀ + 45 ♁♁ from 26 countries): ENGLAND: 1 ♀. SWEDEN: 1 ♀. FRANCE: 2 ♀ + 5 ♁♁ from five localities. GERMANY: 1 ♀. BOHEMIA: 1 ♀ + 1 ♁ from two localities. SLOVAKIA: 2 ♀♀ from two localities. HUNGARY: 30 ♀♀ + 16 ♁♁ from 43 localities. SLOVENIA: 1 ♀. ROMANIA: 4 ♀♀ + 2 ♁♁ from six localities. SPAIN: 1 ♁. ITALY: 2 ♀♀ + 1 ♁ from three localities. CROATIA: 2 ♀♀ + 5 ♁♁ from four localities. BULGARIA: 4 ♀♀ + 3 ♁♁ from seven localities. ALBANIA: 1 ♀. MACEDONIA: 2 ♀♀ + 1 ♁ from three localities. KOSOVO: 2 ♁♁ from one locality. MONTENEGRO: 1 ♁. BOSNIA-HERZEGOVINA: 1 ♀. GREECE: 3 ♀♀ + 3 ♁♁ from four localities. CYPRUS: 2 ♀♀ from one locality. TURKEY: 2 ♁♁ fro, two localities. UKRAINE: 2 ♀♀ from one locality. ARMENIA: 1 ♀ + 1 ♁ from two localities. GEORGIA: 5 ♀♀ from two localities. IRAN: 2 ♀♀ from two localities. NORTH CHINA: 1 ♁.

4) Bracon erraticus var. superciliosus (Wesmael)   (68 ♀♀ + 93 ♁♁ from 18 countries): ENGLAND: 8 ♀♀ + 6 ♁♁ from nine localities. SWEDEN: 1 ♁. DENMARK: 2 ♁♁ from two localties. GERMANY: 3 ♀♀ + 3 ♁♁ from six localities. AUSTRIA: 2 ♀♀ + 1 ♁ from three localities. SWITZERLAND: 1 ♁. SLOVAKIA: 3 ♀♀ + 2 ♁♁ from four localities. HUNGARY: 30 ♀♀ + 49 ♁♁ from 59 localities. ROMANIA: 6 ♀♀ + 11 ♁♁ from sixteen localities. SERBIA: 2 ♁♁ from two localities. KOSOVO: 2 ♀♀ + 2 ♁♁ from one locality. MACEDONIA: 4 ♀♀ + 2 ♁♁ from six localities. MONTENEGRO: 1 ♀. BULGARIA: 4 ♀♀ + 2 ♁♁ from six localities. TURKEY: 2 ♀♀ from two localities. ARMENIA: 1 ♀. IRAN: 1 ♁. MONGOLIA: 1 ♀ + 3 ♁♁ from four localities.

Redescription of the ♀ lectotype of B. erraticus   ( Fig. 8 View Fig A-J)

LENGTH. Body length 3.5 mm.

ANTENNAE. (right flagellum apically deficient) Somewhat shorter than body and with 28 antennomeres (left antenna). First flagellomere 1.8 times, second flagellomere 1.25 times, flagellomeres 3-19 subcubic

(i.e. slightly longer than broad), flagellomeres 20-24 attenuating so that penultimate (or 25th) flagellomere 1.2 times as long as broad ( Fig. 8A View Fig ).

HEAD. In dorsal view transverse ( Fig. 8B View Fig ), 1.7 times as broad as long, eye one-fourth longer than temple, temple rounded, occiput excavated. Eye in lateral view nearly 1.6 times as high as wide, temple beyond eye somewhat less wide than eye ( Fig. 8C View Fig , see arrows). OOL twice as long as POL. Oral opening fairly large: its horizontal diameter 1.4 times as long as shortest distance between opening and compound eye ( Fig. 8D View Fig ). Head polished, face finely granulose, medially polished.

MESOSOMA. In lateral view 1.5 times as long as high, polished. Notaulix distinct, evenly deep, smooth. Propodeum polished, above lunule with oblique striae extending up to middle of propodeum ( Fig. 8E View Fig ).

LEGS. Hind femur thick, 2.5 times as long as broad medially ( Fig. 8F View Fig ). Claw clearly downcurved and its basal lobe fairly large ( Fig. 8G View Fig ).

WINGS. Forewing about as long as body. Pterostigma ( Fig. 8H View Fig ) 2.8 times as long as wide and issuing r from   its middle; r 0.7 times as long as width of pterostigma, second submarginal cell less long, 3-SR 1.25 times as long as 2-SR, SR1 straight, clearly 1.7 times as long as 3-SR and ending near to tip of wing. First discal cell less high, 1-M 1.75 times as long as m-cu, 1-SR-M 1.7 times as long as 1-M ( Fig. 8I View Fig ).

TERGITES. First tergite ( Fig. 8J View Fig ) quadrate in form, somewhat broader behind than long, beyond pair of spiracles weakly broadening, scutum apically and tergite laterally from scutum strongly sculptured. Second tergite 3.4 times as broad behind as long, medially rugose, laterally rugulose. Third tergite somewhat longer than second tergite, rugulose-subrugulose, medially almost smooth. Suture between tergites 2-3 slightly bisinuate, deep, smooth. Further tergites polished. Ovipositor sheath long, as long as hind tibia + basitarsus combined.

COLOUR. Ground colour of body black with light colour pattern. Scape black, flagellum blackish. Margin of eye, cheek and mandible yellow, palpi brown. Tegula brownish. Tergites 2-4 laterally reddish, sternites brownish. Legs dark brown to brown, fore femur apically and fore tibia entirely yellow, tibiae 2-3 basally yellowish. Wings faintly brownish fumous, pterostigma and veins opaque brown.

Redescription of the ♀ paralectotype of B. erraticus  

Similar to the ♀ lectotype. Body 3.4 mm long. Both antennae deficient, right antenna with 22 and left antenna with 20 antennomeres. Head in dorsal view 1.6 times as broad as long ( Fig. 8K View Fig ). Hind femur 2.8 times as long as broad medially ( Fig. 9A View Fig ). Second tergite nearly entirely yellow, i.e. only medially blackish.

Redescription of the ♀ paralectotype of B. erraticus   var. 1

Similar to the ♀ lectotype. Body 3.9 mm long. Left antenna with 28 antennomeres, i.e. flagellum apically deficient. Head in dorsal view ( Fig. 9B View Fig ) 1.7 times as broad as long, temple somewhat receded. Hind femur less thick, 3.1 times as long as broad distally ( Fig. 9C View Fig ). Pterostigma ( Fig. 9D View Fig ) 2.6 times as long as wide, r 0.7 times as long as width of pterostigma, second submarginal cell wide, 3-SR 1.3 times length of 2-SR, SR1 1.4 times as long as 3-SR. First tergite as long as broad behind ( Fig. 9E View Fig ). Tergites 2-6 laterally and tergites 7-8 entirely yellow.

Redescription of the two ♁ paralectotypes is disregarded owing to their very poor condition.

Redescription of the ♀ holotype of B. superciliosus  

Similar to the ♀ holotype of B. erraticus   . Body 4 mm long. Right antenna just shorter than body and with 30 antennomeres. First flagellomere 1.6 times as long as broad, middle flagelloneres subcubic to cubic, penultimate flagellomere 1.3 times as long as broad. Head in dorsal view ( Fig. 9F View Fig ) 1.8 times as broad as long, eye 1.35 times as long as temple, temple rounded. Hind femur 2.6 times as long as broad medially ( Fig. 8L View Fig ). Second submarginal cell short (as usually), 3-SR just longer than 2-SR, SR1 twice length of 3-SR ( Fig. 9G View Fig ). Body almost entirely black, legs more dark coloured. This form is a melanic variety of the nominate form named B. erraticus var. superciliosus (Wesmael)   .

Hosts

COL. Chrysomelidae   : Gastroidea viridula DeGeer.   Curculionidae   : Larinus turbinatus Gyllenhal   ,! Rhinocyllus conicus Frölich.   Scolytidae   : Hylesinus toranio Danthoine et Bernard.   — LEP. Aegeriidae   : Pennisetia hylaeiformis Laspeyres.   Gelechiidae   :! Metzneria lapella Linnaeus.   Tortricidae   : Acleris rhombana Denis et Schiffermüller   ,! Lobesia littoralis Humphrey et Westwood   , Rhyacionia duplana Hübner.   Cochylidae   :! Aethes williana Brahm   ,! Falseuncaria ruficiliana Haworth   , Phalonia posterana Zeller   , Stenodes straminea Haworth.   — DIPT. Agromyzidae   : Melanagromyza lappae Loew.   Tephritidae   : Chaetorella jaceae Robineau-Desvoidy   , Orellia ruficauda Fabricius   , Sitarea lurida (Loew)   , Sitarea scorzonera Robinaeu-Desvoidy   , Tephritis bardanae Schrank   , Terellia serratulae Linnaeus   , Urophora eriolepidis Loew   ,! U. stylata Fabricius   , Xyphosia miliaria Schrank.   — HYM. Tenthredinidae   : Pontania joergenseni Enslin.  

Distribution

Palaearctic Region, in Europe frequent to common.

Colour variabilities

(1) The ground colour of the nominate form ( erraticus   ) is black to blackish with rusty to testaceous pattern on head and tergites 2-3(-4), eventually the line of notaulix is also rusty.

(2) The ground colour of var. confinis (Szépligeti)   is yellow to testaceous with a few dark (light brown to blackish) pattern on vertex, mesoscutum, propodeum and tergites 1-4.

(3) The ground colour of var. superciliosus (Wesmael)   is almost entirely to entirely black with (very) few rusty pattern on fore tergites.

Sculpture variabilities

(1) Nominate form ( erraticus   ): second tergite rugose to rugulose, third tergite rugulose to (almost) smooth.

(2) Var. aestivalis (Szépligeti)   : tergites 2-4(-5) rugose, rugosity slightly weakening posteriorly.

Morphological variabilities

(1) Body 2.5-5.5 mm, usually 3.5-5 mm, long.

(2) Antenna somewhat shorter to somewhat longer than body, the ♀ with (25-)28-33(-40) and the ♁ with (21-)29-35(-40) antennomeres. Flagellomeres cubic to 1.5 times (♀♀) or 1.2-1.3 times to twice longer than broad. Head in dorsal view 1.6-1.85 times as broad as long, eye usually one-fourth, rarely one-fifth to one-third, longer than temple.

(3) Oblique striation of propodeum ( Fig. 8E View Fig ) variable in extent restricting to fine rugulosity close above lunule. Hind femur 2.4-2.8(-3.2) times as long as broad medially ( Figs 8L View Fig ; 9C View Fig ).

(4) SR1 of fore wing clearly approaching, exceptionally rather reaching, tip of wing; second submarginal cell always relatively wide, 3-SR just to 1.3 times longer than 2-SR.

(5) Tergites 1-3 sculptured as in Fig. 8J View Fig , sometimes sculpture (very) weak or relatively rough. Ovipositor sheath short to long, i.e. as long as tarsomeres 1-3(-4) to hind tibia + basitarsus to hind tibia + tarsus combined respectively.

(6) Second tergite rarely as long as, exceptionally just longer than, third tergite (cf. Fig. 28G View Fig ).

(7) First discal cell rarely less high, 1-M 1.5-1.6 times as long as m-cu (cf. Fig. 10G View Fig ; 16G View Fig ).

(8) Male: first tergite sometimes 1.3 times longer than broad behind, beyond pair of spiracles either parallel-sided (cf. Fig. 20I View Fig ) or somewhat broadening posteriorly (cf. Fig. 20J View Fig ).

(9) Male: second submarginal cell of forewing rarely long, 3-SR 1.3 times longer than 2-SR (cf. Fig. 17B View Fig ).

(10) Female: first tergite rarely somewhat broader behind than long (cf. Fig. 20K View Fig ).

The colour pattern and the morphological (eidonomic) features may vary in a highly wide spectrum.

Taxonomic position

The following three species, B. (Gl.) nigriventris (Wesmael)   (Europe), B. (Gl.) peroculatus (Wesmael)   (in Europe sporadic) and B. (Lu.) roberti (Wesmael)   (in Europe sporadic), are near to B. erraticus   , the four species are distinguished by the following key features:

1 (4) Temple in dorsal view receded, eye somewhat protruding ( Figs 36A View Fig ; 44A View Fig ).

2 (3) SR1 reaching tip of forewing, second submarginal cell less wide ( Fig. 36E View Fig ). Second tergites smooth except (sub-)rugulosity close around small areola (subgeneric features of Glabrobracon   ), tergite one-sixth longer than third tergite ( Fig. 36G View Fig ). First discal cell high, 1-M a bit more than twice as long as m-cu ( Fig. 36F View Fig ). ♀: 4.5-5 mm .................................... B. (Gl.) peroculatus Wesmael, 1838  

3 (2) SR1 approaching tip of forewing, second sbmarginal cell wide ( Fig. 44F View Fig ). Second tergite rugorugulose (subgeneric features of Lucobracon   ), just longer than third tergite. ( Fig. 44H View Fig ). First discal cell less high, 1-M 1.8 times as long as m-cu ( Fig. 44G View Fig ). ♀: 4.5-5.1 mm ............................................. .............................................................................................................. B. (Lu.) roberti Wesmael, 1838  

4 (1) Temple n dorsal view rounded, eye not protruding ( Figs 8B, K View Fig ,; 9B, F View Fig ; 28A View Fig ).

5 (6) SR1 approaching tip of forewing, second submarginal cell wide ( Fig. 8H View Fig ; 9D View Fig ). Second tergite rugo-rugulose, third tergite either rugulose-subrugulose ( Fig. 8J View Fig ) or (almost) smooth (subgeneric features of Lucobracon   ). First discal cell less high, 1-M less than twice as long as m-cu ( Fig. 8I View Fig ). Hind femur usually thick, 2.5-2.8 times as long as broad medially ( Fig. 8F, L View Fig ). ♀ ♁: 2.5-4.5 mm ......... ............................................................................................................ B. (Lu.) erraticus Wesmael, 1838  

6 (5) SR1 reaching tip of forewing, second submarginal cell less wide ( Figs 28F View Fig ; 29 View Fig B-C). Second tergite antero-medially striolate, otherwise together with third tergite smooth and shiny (subgeneric features of Glabrobracon   , Figs 28H View Fig ; 29F View Fig ). First discal cell high 1-M clearly twice as long as m-cu ( Fig. 28C View Fig ) or less high ( Fig. 29 View Fig D-E). Hind femur usually 2.8-3 times as long as broad medially ( Fig. 28D, I View Fig ). ♀: 3-3.4 mm ....................................................................................... B. (Gl.) nigriventris Wesmael, 1838  

Bracon erraticus   is also near to Bracon fortipes Wesmael   ; their distinction is given at the latter species.

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Hymenoptera

Family

Braconidae

Genus

Bracon

Loc

Bracon (Lucobracon) erraticus Wesmael, 1838

Papp, Jenő & Xviii, Budapest 2012
2012
Loc

Bracon (Lucobracon) bellicosus

Papp J. 1971: 339
1971
Loc

Bracon hades

Papp J. 1965: 409
1965
Loc

Bracon (Bracon) confinis Szépligeti, 1901: 261

Szepligeti Gy. 1901: 261
1901
Loc

Bracon congruus Szépligeti, 1901: 261

Szepligeti Gy. 1901: 261
1901
Loc

Bracon foveola

Thomson C. G. 1894: 1819
1894
Loc

Bracon erythrostictus

Marshall T. A. 1885: 14
1885
Loc

Bracon exarator

Marshall T. A. 1885: 14
1885
Loc

Bracon praetermissus

Marshall T. A. 1885: 15
1885
Loc

Bracon vectensis

Marshall T. A. 1885: 15
1885
Loc

Braco erraticus

Wesmael C. 1838: 35
1838
Loc

Braco superciliosus

Wesmael C. 1838: 38
1838
Loc

B. erraticus

Papp 2005a: 199
Tobias 1986: 149
Loc

Bracon erraticus

Szépligeti 1901: 261
Fahringer 1927: 270
Telenga 1936: 164
Shenefelt 1978: 1630
Tobias 1986: 145
Tobias & Belokobylskij 2000: 162
Loc

Bracon (Lucobracon) bellicosus

Shenefelt 1978: 1616
Tobias & Belokobylskij 2000: 164
Telenga 1936: 165
Shenefelt 1978: 1628
Tobias 1986: 149
Papp 2004: 173
Loc

Bracon congruus

Fahringer 1927: 242
Telenga 1936: 258
Shenefelt 1978: 1629
Papp 1968: 197
Papp 2004: 173
Loc

Bracon erythrostictus

Fahringer 1927: 237
Telenga 1936: 162
Shenefelt 1978: 1481
Tobias 1986: 125
Telenga 1936 : as valid species 164
Shenefelt 1978: 1618
Loc

Bracon foveola

Telenga 1936 : as valid species 170
Shenefelt 1978: 1631
Shenefelt 1978: 1631
Loc

Bracon praetermissus

Telenga 1936 : as valid species 177
Shenefelt 1978: 1645
Tobias 1986 : as valid species 137
Tobias & Belokobylskij 2000
Loc

Braco superciliosus

Schmiedeknecht 1897: 559
Fahringer 1927: 381
Telenga 1936: 257
Shenefelt 1978: 1631
Fischer 1965: 130
Loc

Bracon vectensis

Telenga 1936 : as valid species 178
Shenefelt 1978: 1650