Bracon (Foveobracon) megapterus Wesmael, 1838

Papp, Jenő & Xviii, Budapest, 2012, A revision of the Bracon Fabricius species in Wesmael’s collection deposited in Brussels (Hymenoptera: Braconidae: Braconinae), European Journal of Taxonomy 21, pp. 1-154: 50-53

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Bracon (Foveobracon) megapterus Wesmael, 1838


Bracon (Foveobracon) megapterus Wesmael, 1838  

Figs 22 View Fig A-L, 23A

Braco megapterus Wesmael, 1838: 22   ♀ (type material: 5 ♀♀, seen 4 ♀♀), type locality: “...environs de Charleroi et de Bruxelles ” ( Belgium), ♀ lectotype (and three ♀ paralectotypes, present designations) in the Royal Belgian Institute of Natural Sciences, Brussels; examined.

Bracon (Orthobracon) biimpressus Telenga, 1936: 176   (in key), 295 (description) (in Russian) and 378 (in key), 402 (description) (in German) ♀ (type material:? ♀), type locality: “Europäischer Teil der UdSSR, ehem. Gouv. Ufa,”.

Bracon megapterus   – Szépligeti 1901: 263 (in key, in Hungarian); 1904 (1901): 175 (in key, in German), ♀. Bracon (Lucobracon) megapterus   – Fahringer 1927: 248 (in key) and 362 (redescription). — Telenga 1936: as Ipobracon megapterus   94 (in key), 98 (redes-cription) (in Russian) 330 (in key, in German). — Shenefelt 1978: 1621 (literature up to 1957).

Bracon (Foveobracon) biimpressus   – Tobias 1961 (in subgenus Foveobracon   created by him), 1986: 122 (in key, designation of ♀ lectotype and locality: “Bashkirskaya ASSR, Belebey” (European Russia); examined, syn. nov. — Shenefelt 1978: 1554 (as valid species, literature up to 1961).

Designation of the ♀ lectotype of Bracon megapterus  

(first label, printed) “Coll. Wesmael”, (second label, printed) “2047”, (third label, partly handwritten) “ Braco   ♀ / megapterus mihi / dét. C. Wesmael”, (fourth label, printed red) “Type”, fifth label is the lectotype card. - Lectotype is in good condition: (1) micropinned, (2) left flagellum missing, right flagellum apically deficient.

Designation of the three ♀ paralectotypes of Bracon megapterus   (1 ♀: “var. 1.”)

Labels identical to those of the lectotype except paralectotype cards. Paralectotype are in good condition: (1) micropinned, (2) one ♀ with deficient flagellum, left hind wing glued separately, (3) one ♀ with missing both flagelli and left fore leg (except coxa + trochanter), (4) one ♀ with missing head and left fore leg (except coxa).

Material examined

11 ♀♀ + 3 ♁♁ from seven countries: GERMANY: 2 ♀♀ + 1 ♁ from three localities. BOHEMIA: 3 ♀♀ + 1 ♁ from two localities. POLAND: 1 ♀. SLOVAKIA: 1 ♀. CROATIA: 1 ♀: EUROPEAN RUSSIA: 1 ♀ + 1 ♁ from two localties. MONGOLIA: 2 ♀♀ from one locality.

Redescription of the ♀ lectotype of Bracon megapterus   ( Fig. 22 View Fig A-I)

LENGTH. Body 5 mm long.

ANTENNAE. Right antenna apically deficient and with 39 antennomeres (according to the original description “de 39 on 40 articles environ”). First flagellomere 1.4 times as long as broad, further flagellomeres gradually shortening so that 37th flagellomere subcubic, i.e. just longer than broad.

HEAD. In dorsal view less transverse ( Fig. 22A View Fig ), 1.7 times as broad as long, eye one-fourth (or 1.3 times) longer than temple, temple rounded, occiput excavated. Oral opening usual in size, its horizontal diameter somewhat longer than shortest distance between opening and compound eye ( Fig. 22B View Fig ). Head polished, face laterally and clypeus weakly uneven, frons finely granulate.

MESOSOMA. In lateral view 1.4 times as long as high, polished. Propodeum polished, lunule unusually protruding and around it with short rugae ( Fig. 22C View Fig ).

LEGS. Hind femur 3.8 times as long as broad medially ( Fig. 22D View Fig ). Claw strongly curved and with fingerlike basal lobe ( Fig. 22E View Fig ).

WINGS. Forewing as long as body. Pterostigma ( Fig. 22F View Fig ) 2.6 times as long as wide and issuing r somewhat   proximally from its middle, r short   , 0.45 times as long as width of pterostigma; second submarginal cell long, 3-SR 1.75 times as long as 2-SR, SR1 straight, reaching tip of wing and 1.4 times as long as 3-SR; 1-R1 distinctly 1.6 times as long as pterostigma. First discal cell subquadrate, 1-M 1.5 times as long as m-cu, 1-SR-M 1.2 times longer than 1-M ( Fig. 22G View Fig ).

TERGITES. First tergite ( Fig. 22H View Fig ) 1.3 times longer than broad behind, pair of spiracles slightly protruding, beyond spiracles tergite parallel-sided, margin of scutum crenulate, scutum smooth. Second tergite a bit longer than third tergite, antero-medially with a somewhat prominent three-sided areola, lateral margin of areola crenulo-rugulose, tergite laterally with a pair of longitudinal impressions, otherwise together with further tergites polished ( Fig. 22H View Fig ). Suture between tergites 2-3 bisinuate, smooth, fairly deep. Ovipositor sheath as long as hind tibia, posterior end of ovipositor apparatus as in Fig. 22I. View Fig

COLOUR. Antenna, head, mesosoma and first tergite black. Eyes with interrupted reddish yellow pattern. Mandible reddish yellow. Palpi proximo-distally black to brown. Coxae + trochanters black, rest of legs blackish brown, fore femur with yellowish pattern. Metasoma (except black first tergite) yellow, areola of second tergite blackish brown. Ovipositor sheath black. Wings dark brownish fumous, pterostigma and veins brown.

Redescription of the three ♀ paralectotypes of Bracon megapterus  

Similar to the lectotype. Eye in dorsal view 1.5 times as long as temple (1 ♀). Tergites 3-6 medially with faint brownish maculae.

Variable features of the ♀ (5 ♀♀) ( Fig. 22 View Fig J-L)

Areola of second tergite laterally more impressed and here with strong crenulo-rugosity, lateral pair of impressions well distinct ( Fig. 22J View Fig ). Hind femur 3.6-3.8 times as long as broad medially ( Fig. 22K View Fig ). Forewing: 3-SR 1.7-1.8 times as long as 2-SR; first discal cell less subquadrate, 1-M twice as long as m-cu ( Fig. 22L View Fig ).


COL. Cerambycidae   : Molorchus umbellatarum Schreber.   — LEP. Thyatiridae   : Achlya flavicornis Linnaeus   , Polyploca ridens Fabricius.  


Belgium, France, Germany, Bohemia, Slovakia, Slovenia, Croatia, Bulgaria, Finland, European Russia, Mongolia. A less frequent to sporadic species.

Taxonomic position

Within the subgenus Foveobracon   the species B. (Fo.) megapterus   is nearest to B. (Fo.) plugarui Tobias   considering their antero-median areola on second tergite and colour of the body; the two species are separated by a few features:

1 (2) Temple in dorsal view rounded ( Fig. 22A View Fig ). Second submarginal cell long, 3-SR 1.7-1.8 times as long as 2-SR, r shorter   than widht of pterostigma ( Fig. 22F View Fig ). Second tergite along areola crenulate ( Fig. 22H, J View Fig ). Antenna with 35-37 antennomeres. Femora 2-3 black. Body strongly built. ♀ ♁: 5 mm ....................................................................................................... B. (Fo.) megapterus Wesmael, 1838  

2 (1) Temple in dorsal view receded ( Fig. 23B View Fig ). Second submarginal cell less long, 3-SR 1.2-1.3 times as long as 2-SR, r longer   than width of pterostigma ( Fig. 23C View Fig ). Second tergite around areola rugorugulose ( Fig. 23D View Fig ). Antenna with 26-27 antennomeres. Femora 2-3 yellow. Body usually built. ♀ ♁: 2 mm ..................................................................................................... B. (Fo.) plugarui Tobias, 1986  














Bracon (Foveobracon) megapterus Wesmael, 1838

Papp, Jenő & Xviii, Budapest 2012

Bracon (Orthobracon) biimpressus

Telenga N. A. 1936: 176

Braco megapterus

Wesmael C. 1838: 22

Bracon megapterus

Szépligeti 1901: 263

Bracon (Lucobracon) megapterus

Telenga 1936
Shenefelt 1978: 1621

Bracon (Foveobracon) biimpressus

Shenefelt 1978: 1554