Sinosmylites fumosus, Makarkin, Vladimir N., Yang, Qiang & Ren, Dong, 2011

Makarkin, Vladimir N., Yang, Qiang & Ren, Dong, 2011, Two new species of Sinosmylites Hong (Neuroptera, Berothidae) from the Middle Jurassic of China, with notes on Mesoberothidae, ZooKeys 130, pp. 199-215: 202-203

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scientific name

Sinosmylites fumosus

sp. n.

Sinosmylites fumosus  ZBK  sp. n. Figs 4A, B


Differs from Sinosmylites pectinatus  by CuP once forked (twice forked in Sinosmylites pectinatus  ), by presence of one ‘inner’ gradate series of crossveins (two in Sinosmylites pectinatus  ) (see differences from Sinosmylites rasnitsyni  sp. n. under that species.).


Forewing with broad-rounded apex, about 6.0 mm long (as preserved, estimated complete length about 6.5 mm), 2.6 mm wide. Costal space mo derately broad, most dilated at proximal 1/5 of wing length. Subcostal veinlets simple, regularly arranged, less closely spaced than in previous species. Sc distally fused with R1 far from wing apex; Sc+R1 with nine veinlets (eight simple, one forked). Subcostal space broad, with two basal crossveins. R1 space nearly as wide as subcostal space; six crossveins before fusion of Sc and R1, one after. Rs with nine pectinate, regularly spaced branches; four proximal-most branches with 2-4 terminal forks, other branches once forked. Rs1 originating at some distance from origin of Rs. M not fused basally; forked much distal to origin of Rs1. MA, MP almost parallel, distally with one (simple) , two (one simple) branches respectively. Cu divided into CuA, CuP proximal to origin of Rs. CuA pectinate, with 7 branches; proximal-most branch once forked. CuP once deeply forked. Anal veins incompletely preserved; 1A with single marginal fork; 2A with two marginal short branches; 3A very incomplete, with single fork preserved. Four gradate series of crossveins posterior to stem of Rs, all incomplete. First series consists of three crossveins: 1r-m (located at origin of Rs), 1m-cu, 1a1-a2 (longer than previous); Second series includes two crossveins: 2m-cu (connecting MP, CuA), 2icu (connecting CuA, anterior branch of CuP). Third ( ‘inner’) series with six crossveins (3rs-rs7, 3rs5-rs4 to 3rs2-rs1; two between Rs3, Rs2). Fourth ( ‘outer’) series with five crossveins (from 4rs2-rs1 to 4m-cu; two between Rs1, MA). Wing one color, fuscous. Veins mainly dark brown as preserved.

Type material.

Holotype CNU-NEU-NN2011003, deposited in CNUB. A nearly complete forewing.

Type locality and horizon.

Daohugou Village, Shantou township, Ningcheng county, Inner Mongolia, China. Jiulongshan Formation, Middle Jurassic.


From the Latin fumosus, smoked, in reference to the coloration of wings.