Glyptasida sordida ( LeConte 1853 )

Lockwood, Stephanie A. & Pollock, Darren A., 2009, A review of the genus Glyptasida Casey (Coleoptera: Tenebrionidae: Asidini), Zootaxa 2023, pp. 1-27 : 7-17

Glyptasida sordida ( LeConte 1853)

( Figs. 1, 4, 7 , 10, 12 , 14–23, 26–28 )

Pelecyphorus sordidus LeConte 1853:445 [Type locality: Arkansas River, near the mountains. Type Repository: MCZ]; LeConte 1859:14; LeConte 1866:59; Champion 1892:492. The female lectotype (here designated) has the following label information: [round label dark green above and white below]/ “ TYPE 4526/ [handwritten det. label] “ sordidus Lec / “ Jan. – Jul. 2005 MCZ Image Database. The lectotype has the following identifying features: good condition with all appendages intact, slight debris on posterior end of abdomen, ovipositor slightly extended. According to Phil Perkins (pers. comm.), round labels were used by LeConte for locality designation. Different colors and in some cases cuts on the labels were used for specific localities. The dark green locality label represents a locality in New Mexico (Phil Perkins pers. comm.).

Asida sordida: Horn 1871:283 ; Crotch 1873:103; Champion 1884:53; Champion 1892:492; Fall and Cockerell 1907:202.

Glyptasida sordida: Casey 1912:96 ; Leng 1920:224; Gebien 1937:704; Pallister 1954:18; Broadus 1974:16, 21, Fig. 12 ; Blackwelder and Arnett 1975:R73.22; Lavigne 1980:39,41; Bousquet 1991:254; Poole and Gentili 1996:435; Stapp 1997a:353; Stapp 1997b:304; Fagerlund 2000:36; NMGF 2008; NAU 2006.

Pelecyphorus costipennis LeConte 1858:20 [Type locality: Arizona. Type Repository: MCZ]; LeConte 1866:59; Horn 1871:283; Champion 1892:492.

Asida costipennis: Crotch 1873:103 ; Fall and Cockerell 1907:202.

Glyptasida costipennis Casey 1912:100 ; Leng 1920:224; Gebien 1937:704; Blackwelder and Arnett 1975:R73.22; Poole and Gentili 1996:435; Arnett 2000:467.

Pelecyphorus irregularis LeConte 1858:19 [Type locality: Llano Estacado. Type Repository: MCZ]; LeConte 1866:59; Champion 1892:492.

Pelecyphorus inaequalis: Horn 1871:283 . Lapsus for Pelecyphorus irregularis .

Glyptasida aegra irregularis: Casey 1912:101 ; Leng 1920:224; Blackwelder and Arnett 1975:R73.22; Poole and Gentili 1996:435.

Asida interrupta Champion 1884:53 [Type locality: Mexico: La Cuidad in Durango. Type repository: BMNH]; Champion 1892:492; Blackwelder 1945:516.

Glyptasida interrupta: Casey 1912:104 ; Gebien 1937:704; Blackwelder and Arnett 1975:R73.22; Poole and Gentili 1996:435.

Glyptasida aegra interrupta: Wilke 1922:269 .

Glyptasida aegra imperfecta Casey 1912:102 [Type locality: unknown, probably from Colorado. Type repository: USNM]; Leng 1920:224; Blackwelder and Arnett 1975:R73.22; Poole and Gentili 1996:435. New synonym.

Glyptasida irregularis imperfecta: Gebien 1937:705 ; Lapsus for Glyptasida aegra imperfecta .

Glyptasida aegra pigra Casey 1912:101 –102 [Type locality: unknown. Type repository: USNM]; Leng 1920:224; Blackwelder and Arnett 1975:R73.22; Poole and Gentili 1996:435. New synonym.

Glyptasida irregularis pigra: Gebien 1937:705 ; lapsus for Glyptasida aegra pigra .

Glyptasida aegra plena Casey 1912:102 [Type locality: unknown. Type repository: USNM]; Leng 1920:224; Blackwelder and Arnett 1975:R73.22; Poole and Gentili 1996:435. New synonym.

Glyptasida irregularis plena: Gebien 1937:705 ; Lapsus for Glyptasida aegra plena .

Glyptasida costipennis fulvisetis Casey 1912:100 [Type locality: unknown. Type repository: USNM]; Leng 1920:224; Gebien 1937:704; Blackwelder and Arnett 1975:R73.22; Poole and Gentili 1996:435. New synonym.

Glyptasida crenicollis Casey 1912:103 [Type locality: Wyoming (Wind River). Type repository: USNM]; Leng 1920:224; Gebien 1937:705; Blackwelder and Arnett 1975:R73.22; Poole and Gentili 1996:435. New synonym.

Glyptasida heres Casey 1912:102 –103 [Type locality: Texas. Type repository: USNM]; Leng 1920:224; Gebien 1937:705; Blackwelder and Arnett 1975:R73.22; Poole and Gentili 1996:435. New synonym.

Glyptasida parvicollis Casey 1912:97 [Type locality: Arizona. Type repository: USNM]; Leng 1920:224; Gebien 1937:704; Blackwelder and Arnett 1975:R73.22; Poole and Gentili 1996:435. New synonym.

Glyptasida procrustes Casey 1912:99 [Type locality: Kansas (Wallace). Type repository: USNM]; Leng 1920:224; Gebien 1937:704; Blackwelder and Arnett 1975:R73.22; Poole and Gentili 1996:435. New synonym.

Glyptasida sordida porcatula Casey 1912:96 –97 [Type locality: Colorado. Type repository: USNM]; Leng 1920:224; Gebien 1937:704; Blackwelder and Arnett 1975:R73.22; Poole and Gentili 1996:435. New synonym.

Glyptasida strigipennis Casey 1912:99 –100 [Type locality: unknown, probably from western Kansas. Type repository: USNM]; Leng 1920:224; Gebien 1937:704; Blackwelder and Arnett 1975:R73.22; Poole and Gentili 1996:435. New synonym.

Glyptasida subpubescens Casey 1912:97 –98 [Type locality: New Mexico (Escondido). Type repository: USNM]; Leng 1920:224; Gebien 1937:704; Blackwelder and Arnett 1975:R73.22; Poole and Gentili 1996:435; Fagerlund 2000:36; NMGF 2008. New synonym.

Glyptasida turbulenta Casey 1912:100 –101 [Type locality: New Mexico. Type repository: USNM]; Leng 1920:224; Gebien 1937:704; Blackwelder and Arnett 1975:R73.22; Poole and Gentili 1996:435; Fagerlund 2000:36; NMGF 2008. New synonym.

Glyptasida turgescens Casey 1912:98 [Type locality: Kansas. Type repository: USNM]; Leng 1920:224; Gebien 1937:704; Blackwelder and Arnett 1975:R73.22; Poole and Gentili 1996:435. New synonym.

Glyptasida turgescens furtiva Casey 1912:99 [Type locality: Kansas. Type repository: USNM]; Leng 1920:224; Gebien 1937:704; Blackwelder and Arnett 1975:R73.22; Poole and Gentili 1996:435. New synonym.

Glyptasida turgescens obesa Casey 1912:98 –99 [Type locality: Kansas (Wallace). Type repository: USNM]; Leng 1920:224; Gebien 1937:704; Blackwelder and Arnett 1975:R73.22; Poole and Gentili 1996:435. New synonym.

Diagnosis. Specimens of Glyptasida sordida are diagnosed on the basis of the following unique combination of characteristics: Pronotum with very coarse and close punctures ( Fig. 10 ), discal costae of elytra elevated, with gradual change in elevation between the lateral elytral margin and the lateral costa ( Fig. 4 ), elytral setae with node at base ( Fig. 12 ).

Description. With general features of Glyptasida , and the following specific characteristics: GHW 2.5–4.4 mm; GPW 5.2–9.6 mm; GEW 7.2–13.2 mm; HL 1.9–3.9 mm; PL 3.3–5.5 mm; EL 7.1–18.7 mm; TL 15.3–27.0 mm; Ratio of PW/GEW 0.50–0.95. Body stout, dull black, in some specimens appearing brown due to debris between costae; pronotum with or without pronounced acute hind angles; pronotal punctures coarse ( Fig. 10 ), punctures close, in some specimens overlapping ( Fig. 10 ); discal costae of elytra elevated, with gradual change in elevation between the lateral elytron margin and the lateral costa ( Fig. 4 ); elytral setae with nodes at base ( Fig. 12 ). Male genitalia with median lobe rounded basally ( Fig. 20 ).

Taxonomic Notes. After examining the type specimens, non-type specimens borrowed from collections, and the original species descriptions, we determined that G. sordida is a highly variable species. Many of the distinguishing characteristics (i.e., pronotal width, elytral width, pronotal serrations) used by Casey (1912) are variable in nature. Because of this variability in shape and size, all of the species and subspecies described by Casey (1912) are reduced to synonyms. This variability in several different characteristics allows for many diverse forms to occur through different combinations of the different characteristics; these combinations of characteristics were often observed in series of specimens from the same location and collection date. Unfortunately, almost all of the species and subspecies described by Casey (synonymized above), were based on isolated specimens and therefore the complete range of variation was unknown to him. Details on the types of the synonyms of G. sordida , and information on condition of the type (s) are presented below.

Notes on type specimens. Pelycophorus costipennis LeConte. The male lectotype (here designated) has the following label information: [round label silver above and white below]/ “ TYPE. 4527”/ [handwritten det. label] “ P. costipennis Webb. Lec. ”/ “ Jan.–Jul. 2005 MCZ Image Database.” The lectotype has the following identifying features: good condition with all appendages intact. The silver locality label represents localities in Arizona, “Valley of Gila” (Phil Perkins pers. comm.).

Pelycophorus irregularis LeConte. The female holotype has the following label information: [round label dark green above and white below]/ “ Type 4529”/ [handwritten det. label] “ P. irregularis Lec. ”/ “ Jan.- Jul. 2005 MCZ Image Database.” The holotype has the following identifying features: pin enters right elytra and exits centrally on the ventral side, pin hole through the right elytron through venter, head glued on, left hind tarsus absent, left middle tibia broken off about ¼ of the way, left middle tarsus absent, right hind tarsus absent, ovipositor extended, debris under ventral side. The dark green locality label represents a locality in New Mexico (Phil Perkins pers. comm.).

Asida interrupta Champion. The male holotype has the following label information: [round label white with orange border] “ Type ”/ “Ciudad, Mex., 8100ft. Forrer.”/ [handwritten det. label] “ Asida interrupta Champ. ”/ [handwritten det. label] “= sordida . var. Lec.”/ “ B.C.A.Col. IV.1. Asida sordida rugosissima interrupta ”/ [red handwritten det. label] “ HOLOTYPE ɗ Asida interrupta Champ. Examined by K.W. Brown Sep. 1968.” The holotype has the following identifying features: left apical maxillary palpomere absent, left antenna with only one basal segment, right antenna with only three basal segments, right middle tarsus absent, right hind leg absent.

Glyptasida aegra imperfecta Casey. The male holotype has the following label information: “ɗ”/ “CASEY bequest 1925”/ “ TYPE USNM 46438”/ [handwritten det. label] “imperfecta Csy.” The holotype has the following identifying features: right antenna with only five basal segments, elytra split open, aedeagus exposed, copper filaments on dorsal and ventral sides near pin.

Glyptasida aegra pigra Casey. The holotype, sex undetermined, has the following label information: “Cal.”/ “CASEY bequest 1925”/ “ TYPE USNM 46436”/ [handwritten det. label] “piger Csy”. The holotype has the following identifying features: left and right apical maxillary palpomere absent, right antenna with only three basal segments, right middle tarsus with only one tarsomere, left front tarsus absent, small pin hole through right elytron.

Glyptasida aegra plena Casey. The female holotype has the following label information: “Cal.”/ “CASEY bequest 1925”/ “ TYPE USNM 46437”/ [handwritten det. label] “plena Csy.” The holotype has the following identifying features: right antenna with only four basal segments, left antenna with only two basal segments, left front tarsus with only three tarsomeres, left middle tarsus with only three tarsomeres, left hind leg absent, right front tarsus with only two tarsomeres, right middle leg absent, right hind tarsus with only one tarsomere.

Glyptasida costipennis fulvisetis Casey. The male holotype has the following label information: “ɗ”/ “CASEY bequest 1925”/ “ TYPE USNM 46434”/ [handwritten det. label] “fulvisetis Csy.” The holotype has the following identifying features: right front tarsus with only one tarsomere.

Glyptasida crenicollis Casey. The female holotype has the following label information: “Wind R. Wyo”/ “CASEY bequest 1925”/ “ TYPE USNM 46439”/ [handwritten det. label] “ crenicollis Csy. ” The holotype has the following identifying features: right antenna with only six basal segments, left antenna with only one segment, right front tarsus absent, left middle tarsus with only one tarsomere, left hind leg absent, ovipositor extended.

Glyptasida heres Casey. One male specimen was mentioned in the original description ( Casey 1912:103). We examined one male specimen. The male holotype has the following label information: “Tex.”/ “CASEY bequest 1925”/ “ TYPE USNM 46435”/ [handwritten det. label] “ heres Csy. ” The holotype has the following identifying features: left and right apical maxillary palpomere are absent, left antenna with only one basal segment, right antenna with only six basal segments, left front tarsus with only three tarsomeres, left middle tarsus with only three tarsomeres, right front leg absent, right middle tarsus with only one tarsomere.

Glyptasida parvicollis Casey. The female holotype has the following label information: “ariz Wickham”/ “CASEY bequest 1925”/ “ TYPE USNM 46426”/ [handwritten det. label] “ parvicollis Csy. ” The holotype has the following identifying features: pin hole through right elytron through venter, tergite and sternite exposed, all appendages intact.

Glyptasida procrustes Casey. The female holotype has the following label information: “Wallace Ks 8/ 10/98 ”/ “Ψ”/ “CASEY bequest 1925”/ “ TYPE USNM 46427”/ [handwritten det. label] “ procrustes Csy. ” The holotype has the following identifying features: right antenna with only one basal segment, right front tarsus with only two tarsomeres, left antenna with only six basal segments, left middle tarsus with only one tarsomere, ovipositor extended.

Glyptasida sordida porcatula Casey. The female lectotype (here designated) has the following label information: “Col”/ “Schaupp”/ “ɗ”/ “CASEY bequest 1925”/ “ TYPE USNM 46425”/ “CASEY determ. sordida-7”/ [handwritten det. label] “porcatula Csy.” The lectotype has the following identifying features: pin hole through right elytron, right antenna with only six basal segments.

Glyptasida strigipennis Casey. The male holotype has the following label information: “CASEY bequest 1925”/ “ TYPE USNM 46431”/ [handwritten det. label] “ strigipennis Csy. ” The holotype has the following identifying features: all appendages intact, pin hole through right elytron through venter, aedeagus extended.

Glyptasida subpubescens Casey. One male and one female specimen were examined. The male lectotype (here designated) has the following label information: “Escondido N.M. Wickh Jul 30 ”/ “CASEY bequest 1925”/ “ TYPE USNM 46428”/ [handwritten det. label] “ subpubescens Csy. ” The lectotype has the following identifying features: all appendages intact, pin hole through the right elytron, male genitalia extended.

Glyptasida turbulenta Casey. The male holotype has the following label information: “N.M.”/ “Gissler”/ “CASEY bequest 1925”/ “ TYPE USNM 46433”/ [handwritten det. label] “ turbulenta Csy. ” The holotype has the following identifying features: all appendages intact, pin hole in the first abdominal segment.

Glyptasida turgescens Casey. One male and one female specimen were examined. The male lectotype (here designated) has the following label information: “Ks”/ “CASEY bequest 1925”/ “ TYPE USNM 46429”/ [handwritten det. label] “ turgescens Csy. ” The lectotype has the following identifying features: apical segments of both maxillary palp absent, right middle leg absent, left front tarsus with only two tarsomeres.

Glyptasida turgescens furtiva Casey. One male and one female specimen were examined. The female specimen has a label marked “CASEY determ.?furtiva-2,” this specimen is not considered a primary type. The male holotype has the following label information: “Kansas”/ “ɗ”/ “CASEY bequest 1925”/ “ TYPE USNM 46432”/ [handwritten det. label] “furtiva Csy.” The holotype has the following identifying features: all appendages intact, pin hole through right elytron through venter, aedeagus extended.

Glyptasida turgescens obesa Casey. The female holotype has the following label information: “Wallace, Ks W Knaus”/ “Ψ”/ “CASEY bequest 1925”/ “ TYPE USNM 46430”/ [handwritten det. label] “obesa Csy.” The holotype has the following identifying features: pin hole in the mesosternum, hole in the posterior end of the right elytron, ovipositor extended.

Distribution. The geographic range of this species is northern Mexico to southern Canada and western Utah to mid Kansas ( Fig. 28 ).

Material examined. 1590 non-type specimens were examined, from the following localities:

CANADA, Alberta: 30 mi S Manyberries, 10.ix.1966, N. Wilkinson, (2, CNCI); Deer Creek, 30.viii.1934, (1, CNCI); Empress, 17.viii.1973, K. Shaw, (2, SEMC); Hwy 41 X S. Sask River, 9.viii.1986, C. and A. Nidek, (1, OSUC); Jct 12 mi Coulee and Bow R., 22.viii.1984, J. and B. Carr and D. Pollock, (3, ENMU); Jct 12 mi Coulee and Bow R., W Vauxhall, Pollock and Reichert, 38. ix.1989, (4, ENMU); Lethbridge, 9.ix.1964, T. Owens, (1, CNCI); Lethbridge, 26.viii.1912, J. Wallis, (1, CNCI); Lethbridge, 4.x.1932, R. White, (1, CNCI); Lethbridge, 29.viii.1912, J. Wallis, (1, CNCI), (1, KWBC); Lethbridge, 6.ix.1924, H. Seamus, (3, CNCI); Lethbridge, 2.ix.1930, N. Criddle, (1, CNCI); Lethbridge, 5.ix.1965, D. Larson, (1, USNM); Lethbridge, 28.viii.1912, J. Wallis, (1, CNCI); Medicine Hat, 1.viii.1963, (2, USNM); Medicine Hat, 22.viii.1923, F. Carr, (3, UASM); Medicine Hat, 18.x.1925, F. Carr, (1, UASM); Medicine Hat, 15.viii.1926, F. Carr, (1, UASM), (1, KWBC); Medicine Hat, 27.viii.1924, F. Carr, (1, UASM), (1, CNCI); Medicine Hat, 17.ix.1934, F. Carr, (1, UASM); Medicine Hat, 21.viii.1930, J. Pepper, (1, CNCI); Medicine Hat, 23.ix.1927, (5, CNCI); Medicine Hat, B. and J. Carr, 10.viii.1963, (5, CNCI); Medicine Hat, 22.viii.1924, N. Criddle, (4, CNCI); Medicine Hat, 22.viii.1924, F. Carr, (1, UASM), (1, CNCI); Medicine Hat, 24.viii.1926, F. Carr, (1, BYUC); Medicine Hat, 7.ix.1925, F. Carr, (2, CUIC); Medicine Hat, August, F. Carr, (1, CUIC); Medicine Hat; 23.viii.1924, F. Carr, (1, UASM); Medicine Hat, 16.viii.1924, F. Carr, (1, UASM); Medicine Hat, 31.viii.1924, F. Carr, (1, UASM); Medicine Hat, (1, UASM), (1, BYUC); Medicine Hat, 5.x.1924, H. Wenzel, (1, OSUC); Medicine Hat, 10.viii.1923, F. Carr, (1, KWBC); Medicine Hat; 29.viii.1928, F. Carr, (1, KWBC); Medicine Hat, F. Carr, (3, BYUC); Milk R. Township 4 Range 1, 1.vii.1966, D. Taylor, (2, USNM); Rolling Hills, 4.ix.1985, C. and A. Nidek, (1, OSUC); Tp 1 Rge 13 W4 Mer, 13.x.1971, B. and J. Carr, (1, CNCI); Tp 13 Rge 12 W4 Mer, 2.ix.1968, B. and J. Carr, (1, CNCI).

MEXICO, Chihuahua: 1.1 mi S Col. Alvaro Obregón, 19.vii.1969, R. and M. Murray, (1, OSUC), (1, TAMU); 1.1 mi S Col. Alvaro Obregón, 20.vii.1969, R. and M. Murray, (1, OSUC), (1, TAMU); 2 mi W Samalayuca, 2.ix.1969, J. Doyen, (7, EMEC); 65 mi NNW Casas Grandes, 27.viii.1957, J. Schaffner, (1, KWBC); 8 mi S Gallego, 27.vii.1953, (1, SEMC); Bachimba Cyn., 25 mi SE Chihuahua, 19.viii.1972, Powell and Veirs, (1, EMEC); Chihuahua City, Höge, (1, USNM), (2, KWBC); Colonia Dublan, D. Beck, (11, BYUC); Colonia Dublan, Beck and Call, ~1931, (2, BYUC); Col. Juarez, 22.vii.1966, Wood and Karren, (1, BYUC); Galeana, 2.viii.1915, W. Foshag, (1, KWBC); Rte 16, 7.6 mi SE Madera, 1.ix.1974, Frania and Ball, (2, UASM). Coahuila: 20.9 km S Cuatro Ciénagas, Gypsum Dunes, 30.ix.1972, M. Mispagel, (1, RLAC); 9 mi SW Cuatro Ciénagas, 28.v.1981, J. Lebherr, (1, EMEC); Mesa de Hilcoat, Sierra del Carmen, 24.vii.1934, R. Baker, (1, KWBC). Sonora: state record only, C. Lumholtz, (6, AMNH).

UNITED STATES OF AMERICA, Arizona: Apache Co. : 12.2 mi S Rock Point, 28.viii.1967, J. Hall, (1, UCRC); 3 mi SE Chinle, 14.ix.1986, Doyen, (15, EMEC); Adamana, Petrified Forest, 27.vii.1976, W. Steiner, (1, USNM); Canyon de Chelly CG, nr. Chinle, 26.viii.1992, E. and C. Fisher, (3, CDFA); Chinle, 13.viii.1994, A. and N. Menke, (1, USNM); Springerville, 17.viii.1935, I. Cantrell, (1, CNCI); Springerville, 8.viii.1933, E. Ball, (1, EMEC); St. Johns, 8.viii.1937, R. Allen, (1, EMEC); Window Rock, 3.viii.1975, R. Aalbu, (1, RLAC). Cochise Co. : 1 mi N Portal, 5.viii.1969, L. Anderson, (1, UCRC); 1 mi S Apache Pass, Dos Cabenzas Mts. , 3.ix.1986, A. Menke, (1, USNM); 1.5 mi E Chiricahua Nat. Mon., 8.viii.1971, (12, AMNH); 1.5 mi E Chiricahua Nat. Mon., 11.viii.1971, (1, AMNH); 10 mi NW Douglas, 13.ix.1950, W. Gertsch and M. Cazier, (2, AMNH); 2 mi W Chiricahua Nat. Mon., 3.viii.1972, D. Sumlin, (1, UCRC); 2 mi N Paradise, 1.viii.1973, Frommer, (1, UCRC); 2 mi E Apache , 2.ix.1966, F. Andrews, (2, KWBC); 3 mi E Apache , 3.ix.1966, F. Andrews, (3, CDFA); 4 mi N Kansas Settlement, 17.viii.1972, Wilcox, (1, WFBM); 5 mi S Tombstone, 26.viii.1959, H. Evans, (1, CUIC); 50 mi E Douglas, 22.viii.1966, L. Anderson, (1, UCRC); 6 mi W Willcox, Ariz 86, 9.viii.1948, G. Ball, (1, UASM); 8 mi E Tombstone, 7.ix.1965, L. and C. O’Brien, (1, CWOB); 8 mi NW Chiricahua Mtns, 21.viii.1974, (1, TTCC); Apache , 31.viii.1941, G. Mackenzie, (2, UCRC); Ca 2.2 mi NE Paradise, 18.viii.1972, S. Frommer, (5, UCRC); Carr Cn., 15 mi S Sierra Vista, 20.viii.1964, R. Sternitzky, (1, CNCI); Chiricahua Mts, W side, 7.viii.1971, D. Richman, (1, USNM); Chiricahua Mtn, 11.ix.1962, D. and J. Knull, (1, OSUC); Chiricahua Mts Nat. Monum., 24.viii.1973, (1, USNM); Chiricahua Mts, W side, 8.viii.1971, D. Richman, (1, USNM); Douglas, 1.x.1931, F. Parker, (1, USNM); Douglas, August, F. Snow, (1, SEMC); Douglas, 26.ix.1947, (1, EMEC); Garden Canyon, Huachuca Mts., 9.vii.1950, G. Brandy, (1, AMNH); Huachuca Mts., Ash Canyon Road, 29.viii.1988, N. McFarland, (1, WFBM); Jct Hereford Rd and Hwy 92, S Sierra Vista, 30.vii.1976, A. Evans, (2, RLAC); Jct 186 and 181, 19.viii.1972, F. and V. Beer, (1, WFBM); Jhus Canyon, Chiricahua Mts, 12.viii.1967, E. Schliuger, (1, UCRC); Lower French Joe Can., app. 1mi W Hwy 50, 9.viii.1992, R. Duff, (1, KWBC); McNeal, 13.ix.1950, W. Gertsch and M. Cazier, (1, AMNH); Pinery Cyn., 1. mi E State 131 on 42, 2.ix.1976, F. Andrews, (3, CDFA); Portal, 26.viii.1957, W. Barr, (1, WFBM); Portal, 1.ix.1955, J. Hood, (2, OSUC); Portal, 1.vii.1966, W. Rosenberg, (1, USNM); Portal, 21.viii.1960, Ball Family, (1, UASM); Portal, 18.vii.1968, L. Anderson, (1, UCRC), (1, KWBC); Portal, 10.viii.1970, L. and M. Anderson, (3, UCRC); Portal, 1.ix.1968, L. Anderson, (1, KWBC); Price Cn Chiricahua Mt, 1.viii.1981, Lenczy, (1, USNM); Ramsey Cn, 15 mi S Sierra Vista, 7.xi.1964, R. Sternitzky, (1, CNCI); S slope Chiricahua Mts., 48.8 km N Douglas, 28.viii.1977, G. and K. Ball, (1, UASM); Scotty Anderson’s Ranch, 3 mi N Paradise Ranch, Chiri. Mts., 2.ix.1966, K. Brown, (54, KWBC), (1, USNM); Sierra Vista, 9.viii.1961, R. Sternitzky, (1, CNCI); Sierra Vista, 13.ix.1966, R. Sternitzky, (1, CNCI), (1, KWBC); Skeleton Canyon, 24.viii.1962, C. Triplehorn, (1, OSUC); Skeleton Cyn at Coronado Nat For boundary gate, 29.viii.1965, K. Brown, (1, KWBC); Skeleton Cyn., 6 mi SE Apache , 4.ix.1958, E. Linsley, (1, EMEC); Southwest Research Station, Chiricahua Mts, 10.viii.1966, E. Schliuger, (1, UCRC); Southwestern Research Station nr Portal, 17.vii.1973, (1, USNM); Stronghold, 6.ix.1960, L. Stange, (1, KWBC); SW Research St’n, 21.ix.1970, F. Hovore, (1, EMEC); Turkey Creek Rd, 2 mi N Gayleville, 8.viii.1971, D. Carlson, (1, CDFA); Willcox, 9.viii.1975, (1, AMNH); Willcox Dry Lake, 26.vii.1969, A. Hardy, F. Andrews, and J. Smith, (1, UCRC). Coconino Co.: 10 mi W Gray Mt., 2.x.1953, M. Cazier, (1, KWBC), (31, AMNH); 11 mi E Jacobs Lake, 10.viii.1950, T. Cohn, P. Boome, and M. Cazier, (1, AMNH); 15 mi E Jacobs Lake, 17.vii.1940, Gertsch and Hook, (1, AMNH); 12 mi S Page, (3, BYUC); 16 mi W Tuba City, 24.ix.1964, C. Peth, (1, UCRC); 17 mi W Leupp, 28.vii.1965, L. Jones, (3, NAUF); (1, KWBC); 2 mi NW Winslow, 1.viii.1963, C. Hysong, (1, NAUF); 2 mi SE Moenkopi, 30.iv.1978, Andrews and Hardy, (2, CDFA); 20 mi N Flagstaff, 7.viii.1969, D. Tiemann, (3, EMEC); 20 mi N Flagstaff, 6.viii.1969, D. Tiemann, (2, EMEC); 20 mi W Leupp, 9.viii.1931, H. Bell, (8, AMNH); 25 mi N Flagstaff, Wupatki Nat. Mon., Rte 377, 19.vii.1979, S. and J. Peck, (2, CNCI); 30 mi E Flagstaff, 2.viii.1936, D. Tiemann, (9, EMEC); 30 mi E Flagstaff, D. Tiemann, 1.viii.1936, (20. EMEC); 30 mi N Flagstaff, 3.xi.1965, G. Denipuk, (1, KWBC); 30 mi N Flagstaff, 3.xi.1963, G. Denipuk, (2, NAUF); 48 km N Cameron, Navajoland, 19.viii.1977, L. Masner, (5, CNCI); 5 mi W Moenkopi, sand dunes, 13.ix.1983, R. Aalbu, (4, RLAC); 7 mi NE Winslow, 5.viii.1963, C. Hysong, (1, NAUF); 7.6 mi SE Moenkopi, 17.vii.1975, Andrews and Hardy, (6, CDFA); Cameron, 19.viii.1950, D. and J. Knull, (1, OSUC); Cameron, 6.ix.1968, L. Anderson, (1, KWBC); Cameron, 11.iv.1954, A. Lewis, (1, KWBC); Cameron, 23.x.1944, R. Beal, (1, EMEC); Coal Mine Canyon, 25.x.1965, R. Beal, (1, USNM); Coconino Nat’l Forest, Flagstaff, 5.ix.1935, T. and G. Hubbell, (3, CNCI); Flagstaff, 7.viii.1948, R. Sanders, (1, WSUC); Flagstaff, 31.viii.1928, (2, FMNH); Flagstaff, 12.viii.1945, (4, UCRC); Fredonia, B. and J. Carr, (1, CNCI); Fredonia, 19.viii.1966, B. and J. Carr, (1, CNCI); Fredonia, 7.ix.1934, Ball, (1, KWBC); Grand Canyon, 11.viii.1959, (2, EMEC); Gray Mountain, 1.viii.1977, Lenczy, (10, USNM); Gray Mountain, 25.ix.1980, B. and J. Carr, (1, CNCI); Jct I-40 and Meteor Crater Road, 22.viii.1999, Riley and Yoder, (1, TAMU); Leupp, 2.viii.1931, H. Bell, (11, AMNH); Leupp, 9.viii.1931, H. Bell, (1, AMNH); Leupp, 4.vii.1931, H. Bell, (1, AMNH); Meteor Crater, 30.vii.1983, C. Barr, (1, CBBC); Meteor Crater Nat’l Landmark, 20.viii.1999, Riley and Yoder, (5, TAMU); Meteor Crater Nat’l Landmark, 21.viii.1999, Riley and Yoder, (1, TAMU); Meteor Crater Nat’l Landmark, 22.viii.1999, Riley and Yoder, (1, TAMU); Meteor Crater near Winslow, 28.vii.1930, E. Tinkham, (1, CASC); Meteor Crater, 26.viii.1963, E. Rogers, (1, EMEC); Navajo Pwr Plant, Page, 14.vii.1972, (2, BYUC); Page, 30.vii.1976, W. Steiner, (1, USNM); Page, 14.vii.1972, (12, BYUC); Page, (12, BYUC); Page, 1.x.1960, Dunk, (1, NAUF); Painted Desert, 6.vii.1947, Cameron, (1, KWBC); Tonalea, 26.vii.1951, W. Boyle, (1, USNM); Tuba City, 12.viii.1948, C. and P. Vaurie, (2, AMNH); Tuba City, 22.ix.1964, S. Welles, (1, EMEC); Two guns, 14.viii.1948, C. and P. Vaurie, (1, AMNH); US 89, between Flagstaff and Cameron, 7.viii.1955, (3, CNCI), (1, USNM); Wupatki, 13.viii.1935, (2, FMNH); Wupacki, 10.x.1935, (1, KWBC); Wupatki Nat. Mon., 4.ix.1977, S. Peck, (4, CNCI); Wupatki Nat. Mon., 2.xii.1971, J. McKoski, (1, FMNH); county record only, 18.viii.1927, R. Beamer, (1, SEMC). Graham Co.: Galiuro Mts., near Sunset, 19.vii.1956, M. Westfall, (2, OSUC); Deer Creek Rnch., Galiuro Mt 65 mi N Wilcox, 18.vi.1956, E. Ordway, (2, AMNH). Greenlee Co.: US 666, ca 30 mi N Cliffton, 14.viii.1969, G. and K. Ball, (3, UASM). Maricopa Co. : Mesa, 1.vi.1931.D. Beck, (1, BYUC). Mohave Co.: 25 mi W Fredonia, 31.viii.1967, J. Hall, (1, UCRC); Base of Vermillion Cliffs, 11 mi W Pipe Springs, 4.ix.1926, (1, KWBC). Navajo Co.: 1.8 mi W Joseph City, 13.ix.1995, K. Brown, (1, KWBC); 22.5 km NE Heber, Rte 377, 21.viii.1967, G. and K. Ball, (1, UASM); 4 mi W Holbrook, 25.vii.1966, S. Welles, (1, EMEC); 5–8 mi NE Winslow, 5.ix.1969, J. Doyen, (7, EMEC); 9 mi N Holbrook, 27.ix.1964, C. Peth, (1, UCRC); Holbrook, 24.vii.1938, (1, OSAC), Holbrook, Wickham, (1, CNCI), (1, EMEC); Holbrook, 5.ix.1969, J. Doyen, (1, EMEC); Joseph City, 4.viii.1938, F.B., (1, OSAC); Joseph City, 24.viii.1952, Leech and Green, (1, KWBC); Joseph City, 7.viii.1937, R. Allen, (2, EMEC); Kayenta, D. and J. Knull, (2, OSUC); Kayenta, 27.viii.1963, Ball Family, (1, OSUC), (10, UASM); Show Low, 12.vii.1954, Cazier and Gertsch, (1, AMNH); Winslow, Wickham, (2, OSAC), (4, CUIC); (1, CNCI), (1, NMSU), (1, EMEC); Winslow, ii.1896, Wickham (10, AMNH); Winslow, ix.1903, Wickham, (1, EMEC); Winslow, (2, CNCI); Winslow, 28.viii.1968, K. Brown, (1, KWBC); Winslow, 10.viii.1935, (1, FMNH); county record only, 15.viii.1927, L. Anderson, (1, SEMC). Pima Co. : Arivaca, 26.viii.1941, B. Hodgen, (1, SEMC). Santa Cruz Co.: Mustang Mts, 30.vii.1941, L. Banker, (2, SEMC); Nogales, 24.x.1939, R. Crandall, (1, KWBC); Santa Rita Mts, E. Smyth, (1, KSUC); Santa Rita Mts, E. Smyth, July, (2, KSUC); Santa Rita Mts, E. Smyth, August, (1, KSUC); Santa Rita Mts, F. Snow, June, (2, KWBC), (1, FMNH); (19, SEMC); Santa Rita Mts, F. Snow, (5, SEMC); Santa Rita Mts., 10.vii.1950, W. Arnold, (1, SEMC). Yavapai Co.: 2 mi N Prescott, 17.vii.1978, A. Gilbert, (1, CDFA); Chino Valley, 29.vii.1961, Bowlan, (1, NAUF); Prescott, 1.ix.1982, Lenczy, (1, USNM); Prescott, 29.vii.1954, L. Anderson, (1, UCRC); US 89, 6.5 mi S Chino Valley, 5.ix.1966, Brown and Quezada, (7, KWBC). Unknown county: Palmerlee, H. Kaeber, (1, USNM); Palmerlee, (1, KWBC); Huachuca Mts, C. Palm, (1, AMNH); south, ii.1896, White Mts, D. Duncan, (1, EMEC); Wickham, (1, AMNH); Horn, (1, AMNH); C. Palm, (5, AMNH); Wickham, (2, AMNH); E. Dickerson, (1, AMNH); state only record, (1, AMNH), (1, WSUC), (1, CUIC), (1, CNCI), (1, FMNH). Colorado: Baca Co. : 18 mi N Two Buttes, 29.x.1982, B. and J. Carr, (1, CNCI), (1, ENMU); Reginer, about 37° 0’N 102° 50’W, 4500ft, vi, (1, AMNH). Bent Co.: Las Animas, 12.ix.1964, A. and R. Raske, (1, RLAC); Las Animas, 12.ix.1964, A. Raske, (2, EMEC), (7, KWBC); Las Animas, 7 mi N, 12.ix.1964, A. Raske, (75, EMEC); Las Animas, 7 mi N, 12.ix.1964, A. and R. Raske, (3, EMEC); county record only, August, Lantz, (8, KSUC); county record only, October, Lantz, (1, KSUC). Boulder Co.: Boulder, 10.vii.1923, D. Vancleave, (1, BYUC); Boulder, 21.ix.1947, R. Gregg, (1, FMNH); Boulder, 21.ix.1947, D. Hamilton, (1, FMNH). Cheyenne Co.: Kit Carson, 1.viii.1971, (1, AMNH). Costilla Co.: 4 mi W Blanca, 11.ix.1964, A. Raske, (1, EMEC); Denver Co.: Denver, 4.v.1971, R. Gregg, (1, FMNH). El Paso Co.: 40 mi SSW Colorado Spgs., 13.viii.1948, M. May, (1, CNCI); Fountain Valley, 25.viii.1938, B. Potts, (1, KWBC). Fremont Co.: Royal Gorge, 25.vii.1978, R. Clark, (1, RLAC). Huerfano Co.: 10 mi NW Walsenburg, 26.vii.1968, E. Becker, (1, CNCI); 3.2 mi E Tioga, 20.ix.1986, J. Stamov, (3, AMNH); 5 mi E Walsenburg, 4.ix.1981, Wappes and Downie, (1, EMEC); Gardner, 10.viii.1918, (1, AMNH); near Cucharas, 3.viii.1942, B. Rotger, (1, KWBC); near Tioga, 19.ix.1942, B.

Rotger, (1, KWBC); Walsenburg, 5.viii.1925, F. Gaige (1, KWBC); Walsenburg, 7.viii.1925, F. Gaige, (1, KWBC); county record only, 4.ix.1981, Wappes and Downie, (1, FMNH). Larimer Co.: Ft. Collins, 1.viii.1956, S. Forsythe, (1, KWBC); Ft. Collins, (2, WFBM); county record only, 13.viii.1952, R. Driesbach, (1, USNM). La Plata Co.: Hermesa, 27.viii.1969, J. Hall, (1, UCRC). Las Animas Co.: Trinidad, August, Lantz, (3, KSUC), (1, KWBC). Lincoln Co.: Limon, 21.ix.1938, R. Osburn, (1, OSUC). Mesa Co.: Mesa, 1.viii.1932, (1, FMNH), (2, KWBC). Montezuma Co. : nr Four Corners Colo. 41, 2 mi NW Jct US 164, 19.ix.1967, F. Andrews, (16, KWBC). Otero Co.: 6 mi SW Timpas, 16.ix.1967, D. Calvert, (2, KWBC). Park Co.: Fairway, (1, AMNH). Rio Blanco Co.: 10 mi W Mecker, 14.viii.1947, (1, FMNH). Saguache Co.: 5 mi S Great Sand Dunes Mon, 9.viii.1976, B. and J. Carr, (2, ENMU); Great Sand Dunes Mon., 10.viii.1976, B. and J. Carr, (1, CNCI); Great Sand Dunes Mon., 8.viii.1976, B. and J. Carr, (1, CNCI). Washington Co.: Akron, 23.x.1971, S. Price, (1, WFBM). Weld Co.: 7 mi E Pierce, 4.viii.1973, R. Gordon, (5, USNM); Central Plains Exp. Range, 8 km N Nunn, 9.viii.1976, J. Scott, (2, EMEC); Pawnee Buttes, 23.viii.1925, Beamer and Lawson, (3, SEMC); Pawnee Nat. Grasslands, 25.x.1976, DLW, (1, EMEC). Yuma Co.: 5 mi NW Joes, 24.viii.1969, R. Biggam, (1, WFBM); 5 mi NW Joes, 26.viii.1969, R. Biggham, (2, EMEC). Unknown County: morr, J. Schuh, (2, AMNH); state record only, (1, KSUC), (1, AMNH). Kansas: Ellis Co.: Hays, 10.ix.1972, (1, FHKS); Yocenento, 6.ix.1984, P. Hubbard, (1, FHKS). Ford Co.: Dodge City, 4.viii.1935, (1, FMNH); Dodge City, Lantz, (2, KSUC). Greenley Co.: county record only, F. Williams, (1, KWBC), (10, SEMC); county record only, (1, FMNH). Hamilton Co.: 6 mi N Syracuse, 16.ix.1964, (1, FMNH). Logan Co.: Smoky Valley Ranch, 8.ix.2000, J. Storm, (1, FHKS). Meade Co.: county record only, (4, KSUC), (1, OSUC). Ness Co.: Arnold, 30.xi.1947, F. Nima, (1, FHKS). Scott Co.: county record only, 12.x.1939, R. Fritz, (1, KSUC). Wallace Co.: Sharon Spgs, Lantz, (1, KSUC); Sharon Spgs, (2, KSUC); county record only, F. Snow, (2, SEMC), (1, CUIC), (1, KWBC), county record only, F. Williams, (1, SEMC), county record only, (1, KSUC); county record only, Lantz, (9, KSUC). Unknown County: West Kansas, Popenoe, (1, KSUC); West Kansas, 1.x.1930, (1, KWBC); state record only, Popenoe, (3, KSUC); state record only, F. Webster, (1, NDSU); state record only, (2, CUIC), (1, TTCC), (1, EMEC). Montana: Cascade Co.: Great Falls, 25.ix.1937, F. Smith, (3, OSAC); Great Falls, 7.ix.1938, F. Smith, (16, OSAC); Great Falls, 20.ix.1938, F. Smith, (13, OSAC). Chouteau Co.: Marias Riv, 5.ix.1924, N. Criddle, (2, CNCI). Dawson Co.: Glendive, 3.viii.1891, (3, UNSM); Maco Sico Park, SW Glendive, 6.ix.51, V. Buck, (2, OSAC). Garfield Co.: Sand Springs, 1 mi S, 27.viii.1972, Lafontaine and Lewis, (1, CNCI). Toole Co.: Shelby, 5.ix.1924, H. Seamus, (7, CNCI), (1, KWBC). Wheatland Co.: Harlowton, 18.viii.1932, (1, KWBC). Yellowstone Co.: Billings, 22.viii.1941, H. Scudmore, (1, FMNH); Billings, 28.viii.1941, (1, FMNH). Nebraska: Banner Co.: Harrisburg, August, R. Wolcott, (2, KWBC), (10, UNSM). Cheyenne Co.: Sidney, (4, UNSM). Dawes Co.: Chadron, 17.ix.1949, M. Bacon, (1, WSUC). Dundy Co.: Benkelman, October, (1, KSUC). Sioux Co.: Hat Creek Basin, Bad Lands, (1, UNSM); Hat Creek Valley, 19.viii.1911, F. Shoemaker, (2, UASM), (1, BYUC); Monroe Canyon, August, R. Wolcott, (17, UNSM), (1, KWBC); Monroe Canyon, 6.viii.1908, (1, UNSM); Monroe Canyon, 18.viii.1911, F. Shoemaker, (3, UASM); Monroe Canyon, (1, OSUC), (2, UASM). Unknown County: state record only, (1, CUIC), (4, USNM). New Mexico: Bernalillo Co.: Albuquerque, 1929–1930, A. Dickens, (1, KSUC); Albuquerque, 11.ix.1977, Guandenover, (1, UNMC); Albuquerque, (1, KWBC); Albuquerque, 10.viii.1959, (1, KWBC); Albuquerque, 21.x.1973, D. Jennings, (1, CUIC); Albuquerque, 30.ix.1973, D. Jennings, (1, CUIC); Albuquerque, 23.x.1973, D. Jennings, (1, CUIC); Albuquerque, 19.viii.1973, D. Jennings, (1, CUIC); Albuquerque, 14.x.1973, D. Jennings, (1, CUIC); Albuquerque, 30.ix.1973, D. Jennings, (2, CUIC); Albuquerque, 16.ix.1973, D. Jennings, (1, CUIC); Albuquerque, 12.viii.1973, D. Jennings, (1, CUIC); Albuquerque, 5.ix.1973, D. Jennings, (1, CUIC); Albuquerque, 30.vi.1946, C. Hoff, (2, AMNH); at S. volcano, W of Albuquerque, C. Hoff, (2, AMNH); Albuquerque, J. Doyen, (1, EMEC); mixed grassland, C. Hoff, (5, AMNH); Petroglyph Park, West Mesa, 5.ix.1980, S. Evans, (1, UNMC); Rio Puerco, 28.viii.1982, C. Fullington, (1, UNMC); rt. 66, 15 mi W Albuquerque, C. Hoff, (2, AMNH); UNM Campus, Albuquerque, C. Hoff, (1, AMNH); county record only, C. Hoff, (1, AMNH). Catron Co.: 10 mi N Apache , 23.vii.1972, W. and C. Triplehorn, (2, OSUC); 10 mi N Apache , 1.viii.1972, B. Triplehorn, (1, OSUC); Hwy 60, 2 km E State Line, 1.viii.1993, (1, CASC); Rte 78, 63.2 mi SW Magdalena, 17.viii.1969, G. and K. Ball, (3, UASM). Chavez Co.: 15 mi E Elk, 23.ix.1950, Gertsch and Cazier, (2, AMNH); 15 mi E Elk, 23.ix.1950, Cazier, Cazier, and Francke, (2, AMNH); Roswell, 1.vii.1968, Lenczy, (1, USNM); county record only, 24.x.1964, J. Green, (1, ENMU). Cibola Co.: 13 mi SW Grants, 8.viii.1990, C. and B. Triplehorn, (1, OSUC); 2 mi E Grants, C. Hoff, (1, AMNH). Colfax Co. : Chico, D. Caffrey, (1, USNM); Koehler, (4, USNM); Koehler, Wickham, (1, USNM); Koehler Junct, 16.viii.1913, E. Kalmbach, (2, USNM); Koehler Junction, 5.viii.1912, (1, USNM); Praire near Koehler, H. Wickham, (1, CASC); Raton, 25.iv.1951, Bryant, (1, BYUC); Raton, 27.x.1964, G. Neilson, (1, NMSU). Curry Co.: county record only, 26.ix.1969, M. Hastain, (1, ENMU); county record only, 10.x.1969, M. Hastain, (2, ENMU); county record only, 10.x.1965, B. Klebold, (1, ENMU). De Baca Co .: 16 mi NW Ft. Sumner, 24.x.1968, R. Longwill, (1, ENMU); 16 mi NW Ft. Sumner, 4.x.1968, D. Whytlaw, (1, ENMU); Alamogordo Dam, 16.x.1965, T. Best, (1, ENMU); county record only, 16.x.1965, Jackson, (1, ENMU). Doña Ana Co.: 2 m E Gemefary, 23.ix.1977, (1, NMSU); Aguirre Spgs Pk, E side Organ Mts, 6.x.1975, W. Logan, (1, NMSU); Jordana Range, 15 mi N Las Cruces, 22.xi.1979, D. Richman, (1, NMSU); Jordana Range, 15 mi N Las Cruces, 20.ix.1979, D. Richman, (1, NMSU); Lac Cruces, 13.viii.1927, (1, KWBC), (3, FMNH); Lac Cruces, 6.vii.1927, (1, KWBC); Las Cruces, 29.vii.1927, (1, FMNH); Las Cruces, 1.xi.1925, (2, FMNH); Las Cruces, 1.viii.1925, (1, FMNH); Las Cruces, 13.vii.1931, (1, FMNH); Las Cruces, (1, NMSU). Eddy Co.: 21–28 mi NE El Paso Gap, 22.ix.1986, J. Doyen, (15, EMEC); 27 mi NE El Paso Gap, 23.ix.1986, J. Doyen, (1, EMEC); Artesia, 8.ix.1951, D. Neill, (1, ENMU); Artesia, 21.vii.1951, W. Neill, (1, ENMU); El Paso Gap, 23.ix.1986, J. Doyen, (2, EMEC); Guadalupe Mts., 9 mi N Queen, 6.x.1993, S. Clark, (2, RLAC). Grant Co.: 1 mi W Silver City Airport, 3.ix.1969, L. and M. Anderson, (1, UCRC); 26 mi W Deming, 6.ix.1970, R. Turnbow, (1, FMNH); 4.5 mi SW Hurley on rd. to Silver City, 7.ix.1971, Brown and Petrulis, (9, KWBC); City of Rock S.P., 26.vii.1976, Cazier, Cazier, and Fracke, (2, WSUC), (8, TTCC); Rest area W of Continental Divide, 25.vii.1991, R. Duff, (1, KWBC); Rte 90, 3 mi W Continental Divide, 12.viii.1971, (2, AMNH); Silver City, J. Wallis, (3, CNCI); Silver City, 11.vii.1974, B. Triplehorn, (2, OSUC); county record only, 8.viii.1935, (1, FMNH). Guadalupe Co.: I-40, 6 mi E Santa Rosa, 22.ix.1979, D. Brzoska, (2, OSUC); Vaughn, 1.vii.1968, Lenczy, (3, USNM); Vaughn, 22.vii.1965, (2, USNM); Vaughn, 5.xi.1949, O’Bryant, (5, BYUC); Vaughn, 12.ix.1957, H. Dogde, (3, WSUC); Vaughn, 22.viii.1965, (1, USNM). Harding Co.: Roy, 23.x.1971, L. Cain, (2, ENMU). Hidalgo Co.: 1 mi N Lordsburg, 31.viii.1965, J. Doyen, (1, EMEC); Cloverdale, 12.viii.1955, R. Dreisbach, (1, USNM); Double Adobe Ranch, 28.vii.1997, MSB Bat Crew, (1, UNMC); Playas Valley, 11.viii.2001, S. Clark, (1, BYUC); Post Office Cyn, Peloncillo Mtn, 29.vii.1976, Cazier, Cazier, and Fracke, (1, TTCC). Lea Co. : 32°24.7'N 103° 40.9W, 16.viii.1979, Delorme and McHugh, (1, TAMU); 32°24.8'N 103°41.5'W, 16.viii.1979, J. Schaffer, (1, TAMU); 5 mi E Maljamar, 2.x.1971, C. O’Brien, (4, KWBC); Buckeye, 1.viii.1969, C. Smith, (1, ENMU); Hwy 380, 15 mi W Tatum, 2.ix.2001, C. Graham, (2, ENMU); Hwy 380, 15 mi W Tatum, 18.x.2001, C. Graham, (1, ENMU); county record only, 28.ix.1964, J. Ingle, (1, ENMU). Lincoln Co.: Carrizozo, 31.viii.1938, R. Allen, (1, EMEC); Carrizozo, 25.vii.1938, (1, OSAC); Corona, 1.ix.1970, Lenczy, (6, USNM); Corona, 23.ix.1969, Lenczy, (1, USNM); Escondido, ~ 10 mi N Tinnie, (1, KWBC); Escondido, (2, USNM); Ruidoso, 14.iv.1954, (1, KWBC); Sierra Blanca Mt, 1.vii.1963, Lenczy, (1, USNM). Luna Co.: 11 mi N Deming, 7.viii.1950, J. MacSwain, (3, EMEC); 20 mi NW Deming, 7.ix.1971, K. Brown, (7, KWBC); 7 mi NW Deming, 22.ix.1950, W. Gertsch and M. Cazier, (5, AMNH). McKinley Co.: 30 mi. N Gallup, 14.viii.1965, Gertsch and Roth, (2, AMNH); 6 mi N Tocatechi, 14.vii.1956, V. Roth, (2, CNCI); near Thoreau, 9.x.1980, J. Goodson; Thoreau, 1.vii.1942, R. Russo, (1, FMNH); county record only, C. Hoff, (3, AMNH). Mora Co. : Mora , 8.x.1969, G. Naler, (1, KWBC). Otero Co.: Hwy 54, 16 mi N Orogrande, 18.viii.1976, B. and J. Carr, (1, CNCI); White Sands, 21.vii.1978, Hardy and Andrews, (35, CDFA), (5, RLAC); White Sands, 11.vii.1956, H. and A. Howden, (2, CNCI), (1, USNM); White Sands, 1.vii.1968, Lenczy, (5, USNM); White Sands, 1.viii.1973, Chandler, (5, USNM); White Sands, 22.viii.1971, Lenczy, (1, USNM); White Sands, 19.viii.1962, W. and C. Triplehorn, (4, OSUC), (1, KWBC); White Sands, 23.vii.1970, J. Gruwell, (1, RLAC); White Sands, 30.viii.1982, R. Aalbu, (1, RLAC); White Sands, 18.viii.1973, Iselin and Chandler, (1, OSUC); White Sands, 10.ix.1948, W. and C. Necker, (1, FMNH); county record only, 20.xi.1963, C. M., (1, ENMU). Quay Co.: Tucumcari, 31.viii.1970, Lenczy, (1, USNM). Rio Arriba Co.: 0.5 mi. S Cebolla, C. Hoff, (1, AMNH). Roosevelt Co.: 1100 Block Leo Dr., 2 mi E of ENMU, 18.x.2001, C. Scott, (1, ENMU); 3 mi S Portales, 10.ix.1966, R. Clark, (1, ENMU); 8 mi NE Portales, 12.ix.1971, (1, ENMU); 8 mi NE Portales, 1.ix.1971, (1, ENMU); 8 mi NE Portales, 23.viii.1971, (1, ENMU); 8 mi NE Portales, 4.x.1971, (1, ENMU); 8 mi W Dora, 4.xii.1968, C. Hays, (1, ENMU); Portales, 16.x.1968, Brown, (1, ENMU); Portales, 30.x.1969, Brown, (1, ENMU); Portales, 20.x.1966, J. Diersing, (1, ENMU); Portales, city limits, 20.x.1965, O’Quinn, (1, ENMU); county record only, 6.ix.1965, Wesson, (2, ENMU); county record only, 16.x.1965, S. Benton, (1, ENMU); county record only, 8.x.1964, A. Berg, (1, ENMU); county record only, 25.ix.1964, E. Dunn, (1, ENMU). Sandoval Co.: Coronado State Park, Bernalillo, 21.viii.1985, (3, BYUC); San Ysidro, 15.vii.1950, T. Cohn, P. Boome, and M. Cazier, (8, AMNH). San Juan Co.: Newcomb, 27.vii.1938, D. and J. Knull, (2, OSUC); Shiprock, 20.viii.1940, B. Rotger, (2, KWNC); county record only, 26.viii.1962, F. Moore, (7, OSUC), (1, KWBC). San Miguel Co.: Las Vegas, 11.vii.1976, B. and J. Carr, (1, ENMU). Santa Fe Co.: Puve Indian Reservation, D. Howell, (1, OSEC); rt. 66, just E Edgewood, C. Hoff, (1, AMNH); Santa Fe, 1.x.1973, D. Lowrie, (1, UNMC); county record only, 18.vii.1928, W. Benedict, (2, SEMC); country record only, C. Hoff, (1, AMNH). Sierra Co.: Hillsboro, 22.vii.1936, J. Beamer, (1, SEMC). Socorro Co.: 12.5 mi E Magdalena, C. Peth, (1, UCRC); 13 km SE Bernado, 3.ix.1992, S. Brantley, (1, UNMC); 13 km SE Bernardo, 26.x.1992, S. Brantley, (1, UNMC); 15 km SE Bernardo, 6.xii.1991, S. Brantley, (1, UNMC); 16 mi W Socorro, 13.viii.1952, A. Cole, (1, CNCI); 17 km W Bernardo, 18.x.1991, S. Brantley, (1, UNMC); 18 km SE Bernardo, 30.viii.1994, S. Brantley, (2, UNMC); 18 km SE Bernardo, 1.ix.1992, S. Brantley, (1, UNMC); 2 km SE/E Bernardo, iiiii.1990, S. Brantley, (1, UNMC); 20 mi W Magdalena, C. Hoff, (1, AMNH); Magdalena, Strickler, (1, KWBC), (3, KSUC); Magdalena, (1, KWBC); Sevilleta Sand Dunes, 9.ix.1983, K. Nislow, (1, UNMC); Socorro, 1.viii.1981, F. Snow, (2, CUIC); Socorro, 4.viii.1969, J. McClellan, (2, BYUC). Torrance Co.: 5 mi W Encino, 1.ix.1971, Brown and Petrulis, (1, KWBC); 6 mi E Moriarty, 18.ix.2003, T. Massey, (1, ENMU); 7 mi E Moriarty, 18.x.2003, T. Massey, (1, ENMU); 8 mi E Moriarty, 18.x.2003, T. Massey, (1, ENMU), Clines Corner, (1, KWBC); Clines Corner, 24.viii.1970, C. O’Brien, (1, KWBC); Willard, 1.vii.1978, Lenczy, (8, CMNC); Willard, 22.viii.1975, (1, EMEC); rt. 41, 1 mi N McIntosh, C. Hoff, (1, AMNH); county record only, 1.vi.1925, C. Martin, (2, SEMC); county record only, 21.viii.1968, G. Noonan, (1, KWBC). Union Co.: Cimarron Valley, 13.viii.1973, H. B., (5, KWBC); De Moines, 27.ix.1979, (1, NMSU); Seneca, 25.ix.1965, T. Best, (3, ENMU). Valencia Co. : county record only, 25.ix.1914, (1, USNM); county record only, C. Hoff, (1, AMNH). Unknown County: state only record, 21.ix.1933, (2, KWBC), state record only, (1, UNSM), (1, KSUC), (2, OSAC), (3, USNM), (1, SEMC), (3, EMEC). North Dakota: Billings Co.: county record only, 22.viii.1962, (2, TTCC), (1, NDSU). Slope Co.: Burning Coalvein Cmpgd., 10 mi NW Amidon, 12.ix.1985, P. Aiken, (1, EMEC); county record only, 18.viii.1961, J. Oberfoell, (1, USNM). Oklahoma: Cimarron Co.: Black Mesa State Park, 5.ix.2004, C. Smith, (1, OSEC). South Dakota: Custer Co.: FR 715, 8 mi E Dewey, 17.viii.1988, B. and J.Carr, (2, CNCI). Fall River Co.: Angostura Res, 17.viii.1988, B. and J. Carr, (1, CNCI); Hot Springs, 27.viii.2003, W. Pierce, (1, UNSM); Hot Springs, 27.ix.2003, (1, UNSM). Haakon Co.: Billsburg, 29.ix.1968, C. Calkins, (1, NDSU). Harding Co.: Buffalo, 29.ix.1965, V. Kirk, (1, USNM), (1, NDSU); Buffalo, 30.ix.1965, V. Kirk, (1, USNM); Buffalo, 13.ix.1965, V. Kirk, (1, USNM); Buffalo, 21.ix.1965, V. Kirk, (1, NDSU); Buffalo, 13.ix.1968, C. Calkins, (1, NDSU), (3, USNM); Buffalo, 5.ix.1968, C. Calkins, (3, NDSU); Buffalo, 28.viii.1968, C. Calkins, (1, NDSU); Buffalo, 21.ix.1966, V. Kirk, (1, NDSU); Buffalo, 21.ix.1968, V. Kirk, (1, NDSU); Buffalo, 23.viii.1968, V. Kirk, (1, USNM); Buffalo, 29.ix.1968, V. Kirk, (1, USNM). Unknown County: West River, 10.x.1967, C. Calkins, (2, USNM); West River, 13.vii.1968, C. Calkins, (1, USNM); state record only, (2, CUIC). Texas: Andrews Co.: Shafter Lake, 16.ix.1986, J. Stamov, (1, AMNH). Armstrong Co.: N Rim Palo Duro Cyn, 14 mi S Claude, 29.viii.1971, Brown and Petrulis, (1, KWBC). Brewster Co.: Green Valley, H. Wenzel, (1, OSUC); Marathon, 14.vi.1948, Cazier, (1, AMNH). Crockett Co: 8 mi W Ozona, 30.ix.1950, T. Cohn, P. Boome, and M. Cazier, (1, AMNH). El Paso Co.: El Paso, (1, OSUC); El Paso, 1.ix.1929, (1, OSUC); El Paso, 29.vii.1938, (1, OSUC); Fabens, 10.x.1943, J. Robinson, (1, KWBC); Fabens, 18.viii.1978, W. Suter, (1, FMNH); Fabens, 1.x.1926, (1, OSUC); county record only, 15.x.1943, A. Everline, (1, TTCC). Gaines Co.: Seminole, 22.viii.1948, (1, USNM). Hansford Co.: Spearman, 10.x.1981, S. Jones, (1, TAMU). Jeff Davis Co.: 4 mi W Fort Davis, 4.viii.1968, (1, FMNH); 6.5 mi W Fort Davis, 10.viii.1947, B. Valentine, (1, OSUC); Valentine, 25.vi.1947, A. Michener, (1, SEMC). Motley Co.: Matador, 1.ix.1966, F. Nonini, (2, WFBM). Reeves Co.: 1.4 mi NW Pecos, 5.ix.1997, R. Turnbow, (2, OSUC); 2 mi N Pecos, 5.ix.1997, Wappes and Turnbow, (3, EMEC). Yoa ku m Co.: 8 mi E Plains, 27.viii.1939, Rehn and Rehn, (1, KWBC); Plains, 2.xi.1981, W. Steiner, (5, USNM). Unknown County: Hueca, 28.vii.1938, (1, OSAC); Cedar Lake, 2.vii.1962, (3, USNM). Utah: Duchesne Co.: Lake Powell, 3.ix.1999, J. and R. Walters, (2, OSUC); Lake Powell, 4.ix.1999, J. and R. Walters, (1, OSUC); Monument Butte, S Myton, 6.viii.1998, Ellison and Mendel, (1, BYUC); Monument Butte, S Myton, 5.viii.1998, Ellison and Mendel, (3, BYUC); Monument Butte, S Myton, 30.vii.1998, Ellison and Mendel, (3, BYUC); Monument Butte, S Myton, 12.viii.1998, Ellison and Mendel, (2, BYUC); Monument Butte, S Myton, 22.viii.1998, Ellison and Mendel, (1, BYUC); Monument Butte, S Myton, 28.vii.1998, Ellison and Mendel, (1, BYUC); county record only, 1.viii.1947, R. Olmstead, (1, SEMC). Emery Co.: 40 mi SW Green R., 10.ix.1972, (1, OSUC); NE side nr. Black Dragon Canyon, 14.ix.1943, H. Vokes, (7, AMNH); San Rafael Desert, near Little Gilson Butte, 19.viii.1980, (2, USNM); county record only, 23.ix.1972, A. Shen, (2, OSUC). Garfield Co.: Escalante, 20.x.1999, J. Dunlap, (3, BYUC); Harris Wash Trailhead off Hole in the Rock Rd, 26.x.1999, C. and R. Nelson, (1, BYUC). Grand Co.: 3 mi S Green River, 6.ix.1943, H. Vokes, (1, AMNH); Canyon Lands N.P., 13.vii.1970, J. Petty, (1, BYUC); Crescent Jct, 8 mi W, 7.ix.1964, A. Raske, (2, KWBC), (3, EMEC); Dead Horse Point State Park; 1.v.1963, H. McCoard, (1, BYUC); Dead Horse Point, 13.v.1969, H. McCoard, (1, BYUC); Green River, 19.ix.1986, V.S., (2, BYUC); Green River, 6 mi S Green River, 19.x.1986, Baumann and Nelson, (16, BYUC); Green River, NE of Green River, 19.x.1986, Baumann and Nelson, (1, BYUC); probably near Moab, 30.viii.1965, R. Hardy, (3, KWBC); Thompson’s Spgs, 9.vi.1999, (1, KWBC). Kane Co.: 5 mi N 40 mi E Kanab, 1.iv.1985, D. Giuliani, (2, CDFA); Cedar Mtn, Glen Cyn Cty, 18.viii.1971, (1, BYUC); Cedar Mtn, 15.viii.1971, (2, BYUC); Nipple Bnch, Glen Cnyn Cty, 17.viii.1972, (1, BYUC); Nipple Bnch, Glen Cnyn Cty, 11.viii.1971, (1, BYUC); Nipple Bnch, Glen Cnyn Cty, 16.viii.1972, (1, BYUC); Nipple Bnch, Glen Cnyn Cty, 12.viii.1971, (1, BYUC); Nipple Bnch, Glen Cnyn Cty, 13.viii.1971, (1, BYUC); Red Well off Hole in the Rock Rd, 28.x.1999, C. and R. Nelson, (1, BYUC). Salt Lake Co.: county record only, 1.vii.1920, (1, KWBC). San Juan Co.: 15 mi NE Mexican Hat, 19.ix.1967, F. Andrews, (5, KWBC); 2 mi S Bluff Sand Island Campground, 22.viii.1984, Nelson and Baumann, (11, BYUC); Bluff, 12.x.1967, V. Tanner, (1, BYUC); Cottonwood Wash, 10 mi SW Blanding, (2, BYUC); Monument V., 1.vi.1968, Lenczy, (1, USNM); San Juan River, 3 mi SSW Bluff, 1.iv.1984, D. Guiliani, (1, CDFA); San Juan River, San Juan Cmpg., 22.viii.1984, (1, BYUC); county record only, 1.viii.1953, M. Coffey, (1, BYUC). Uintah Co.: 10 mi SSW Vernal, 28.vii.1983, R. Aalbu, (1, RLAC); 8 mi W Vernal, 22.viii.1949, R. Fredrickson, (1, SEMC); Quarry, Dinosaur Nat. Mon., 10.viii.1970, M. Goldware, (1, RLAC); SW of Vernal, 18.ix.1999, T. Maxwell, (2, BYUC); Whiterocks R., 2.vi.1971, J. Petty, (1, BYUC). Washington Co.: Dixie Nat. For., 1.vi.1968, Lenczy, (1, USNM). Wayne Co.: 13 mi S Hanksville, 2.viii.1968, A. Howden, (4, CNCI), (1, KWBC); 13 mi S Hanksville, 4.viii.1968, A. Howden, (1, CNCI); 13 mi S Hanksville, 9.viii.1968, A. Howden, (1, CNCI); 14 mi S Hanksville, 25.vi.1978, Hardy and Andrews, (1,CDFA); Fairview Ranch, 13 mi S Hanksville, 9.viii.1968, A. Howden, (2, CNCI);); Fairview Ranch, 13 mi S Hanksville, 4.viii.1968, H. Howden, (1, CNCI); Fairview Ranch, 13 mi S Hanksville, 4.viii.1968, J. Martin; (1, CNCI); Fairview Ranch, 13 mi S Hanksville, 2.viii.1968, H. Howden, (2, KWBC), (1,CNCI). Wyoming: Albany Co.: 13 mi SW Garrett, 26.vii.1986, W. Suter, (1, FMNH); 5 mi SW Garrett, 29.vi.1985, D. Field, (1, FMNH). Big Horn Co.: 10 mi E Lovell, 17.viii.1979, S. and J. Peck, (1, CNCI); Big Horn Basin, 13.viii.1910, (1, UNSM). Carbon Co.: county record only, (1, FMNH), (4, AMNH). Converse Co.: 10 mi S Douglas, 25.viii.1962, (1, UNSM); 13 mi W Bill, 25.ix.1982, M. Rice, (7, WFBM). Crook Co.: 15 mi S Sundance, 2.ix.1962, Isakson, Rolofson, and Valder, (1, UNSM). Fremont Co.: Lander, 10.ix.1933, L. Shultz, (1, OSAC); Riverton, 14.viii.1957, G. Knowlton, (3, OSUC); Shoshone, 14.viii.1957, G. Knowlton, (2, OSUC). Laramie Co.: Cheyenne, (1, UNSM); Wind River, ~1984, (1, KSUC); county record only, 2.v.1927, L. Ward, (1, OSAC). Platte Co.: Chugwater, 19.v.1955, (1, KWBC); Guernsey, 10.viii.1961, (1, KWBC). Washakie Co.: Worland, 1.viii.1911, (1, KWBC), (1, UNSM); Worland, (1, UNSM). Weston Co.: Newcastle, (1, UNSM). Unknown County: Natural bridge, 16.viii.1968, N. Baker, (1, KWBC), state record only, (1, USNM), (1, CNCI). Unknown Localities: Ca., (1, WSUC), (1, CUIC); Dak., (1, KWBC); Or., (2, KWBC); No locality Data, (1, TAMU), (2, CUIC); (1, BYUC), (1, NMSU), (4, OSAC), (1, NDSU), (1, USNM), (1, KWBC), (1, UNSM), (3, AMNH).

Figures

FIGURE 28. Geographic distribution of Glyptasida species. Circles indicate localities of G. sordida, triangles indicate G. aegra, and squares indicate G. rugosissima.

FIGURES 1 – 9. Habitus. Dorsal view: 1, 2, 3) Posterior view: 4, 5, 6) Lateral view: 7, 8, 9) Images of G. sordida (1, 4, 7); total length = 18.0 mm. Images of G. aegra (2, 5, 8); total length = 20.6 mm. Images of G. rugosissima (3, 6, 9); total length = 19.3 mm.

FIGURES 10 – 13. Discal pronotal punctures: 10) G. sordida. 11) G. aegra. Scale bar = 100 µm (10, 11). Elytral microsculpturing: 12) G. sordida (showing nodes at base of setae). 13) G. aegra (without nodes at base of setae). Scale bar = 30 µm (12, 13).

FIGURES 14 – 17. Head Structures of G. sordida: 14) Anterior view of head. Scale bars = 300 µm (14, 16). 15) Antennomeres 10 and 11; scale bar = 60 µm. 16) Dorsolateral view of eye. 17) Right mandible, dorsal view; scale bar = 200 µm.

FIGURES 18 – 27. Sexual Dimorphism of G. sordida: 18) Male maxillary palp (outline). 19) Female maxillary palp (outline). Scale bar = 1 mm (18, 19). Male Genitalia of Glyptasida spp.: 21, 25) Median lobe, ventral view; scale bar = 1 mm. 20, 24) Median lobe, lateral view; scale bar = 1 mm. 22) Tegmen, ventral view; scale bar = 1 mm. 23) Tegmen, lateral view; scale bar = 1 mm. Images of G. sordida (20, 21, 22, 23), G. rugosissima (24, 25). Ap = Apicale; Ba = Basale. Female Genitalia of G. sordida: 26) Genital tract; scale bar = 1 mm. 27) Ovipositor, ventral view; scale bar = 1 mm. Ba = Baculi; Co = coxite; Dp = depression; Mg = Medial groove; Ov = Oviduct; SAG = Spermathecal accessory gland; Sp = spermatheca; Va = Vagina; Val = Valvifer; Vu = Vulva.