Heteropimpla pulverulenta, Kopylov & Zhang & Zhang, 2021

Kopylov, Dmitry S., Zhang, Qi & Zhang, Hai-Chun, 2021, The Darwin wasps (Hymenoptera: Ichneumonidae) of Burmese amber, Palaeoentomology 4 (6), pp. 592-603 : 595

Heteropimpla pulverulenta sp. nov.

( Fig. 1 )

LSID: urn:lsid:zoobank.org:act:0BBEC7D3-A4E2-4E9A-9893-DF59BB54DA7C

Holotype. NIGP174880 , sex unknown. Head, fore and hind wings well-preserved; apical metasomal segments lost; amber piece is densely stuffed with dust, almost obscuring tiny body details.

Material. Holotype only.

Etymology. The species is named pulverulenta (Latin) —dusty.

Diagnosis. Fore wing with 1-Rs&1-M equal to r-rs, 2-Rs+M significantly longer than 1m-cu, 2m-cu parallel to 3-Cu; hind wing with 1-Rs equal to r-m, nervellus broken below middle (1-Cu longer than cu-a).

Description. Head with eyes large, globular. Distance between ocelli equal to ocellus diameter, distance between eye and lateral ocellus shorter than ocellus diameter. Antennae filiform, 1.2× as long as wing length; scape fusiform, pedicel not distinctly separated from scape; flagellum 20-segmented, with flagellomeres gradually shortening from base to apex, and apical segment twice as long as subapical one. Mandible slender, strongly curved, with 2 teeth.

Fore wing with pterostigma as long as wide, with rrs issuing from its midlength; 1-Rs&1-M 0.9× as long as r-rs; 2-Cu 1.9× as long as 1-Cu; 2-Rs+M 1.9× as long as 1m-cu, meeting at distinct angle; short ramulus present; 2-Rs 0.8× as long as 2+3-M; 2-Rs 1.6× as long as 3- Rs; 2+3-M 2.4× as long as 4-M; 2m-cu slightly arched, parallel to 3-Cu; 1cu-a arched.

Hind wing with 7 distal hamuli; 1-Rs as long as r-m; 1-Cu 1.6× as long as cu-a.

Measurements. Fore wing length 3.8 mm, width 1.3 mm; hind wing length 2.7 mm, width 0.8 mm; antennal length 4.4 mm; estimated body length 5.2 mm.

Remarks. Differs from H. megista in shorter 1- Rs&1-M, longer 2-Rs+M, 1-Cu longer, 2m-cu parallel to 3-Cu in fore wing; in 1-Rs equal to r-m, nervellus broken below middle in hind wing; and in slightly smaller body size.

The holotype of H. pulverulenta has a short stub of aberrant crossvein close to r-m in fore wing. The presence of an additional r-m crossvein is uncharacteristic of ichneumonids, although Praeichneumonidae possess a second r-m crossvein. However basal ichneumonids (Tanychorinae) never displayed any traces of an extra r-m vein. Thus, we treat this structure as incidental developmental deviation, not atavism.

Heteropimpla pulverulenta is the only known novichneumonine with 1-Rs as long as r-m. In all other genera and species 1-Rs is much shorter.

Figures

FIGURE 1. Heteropimpla pulverulenta sp. nov., holotype NIGP174880. A, Lateral view. B, Wings in transmitting light. C, Habitus drawing. D, Wings reconstruction. Scale bars = 1 mm.