Parahabetia, Ingrisch, 2021
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publication ID |
https://doi.org/ 10.11646/zootaxa.5020.2.1 |
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publication LSID |
lsid:zoobank.org:pub:4FF882DF-334F-49C8-A576-4192B5F2654C |
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persistent identifier |
https://treatment.plazi.org/id/FA08E75D-AA12-8071-FDC0-307894F46F82 |
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treatment provided by |
Plazi |
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scientific name |
Parahabetia |
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gen. nov. |
Parahabetia View in CoL gen. nov.
Typus generis: Habetia pictifrons Karny, 1911 , here designated
Diagnosis. In general habitus the new genus resembles Habetia: The pronotum has the posterior area not prolonged but leaves the stridulatory apparatus in male largely uncovered; the lateral lobes are not deep, and the ventral margin is little descending posteriorly; the hind margin of the lateral lobes is rather strongly ascending and substraight or faintly concave, with a humeral sinus only faintly indicated or missing, and the hind margin of the disc is substraight to faintly convex in both genera, Parahabetia gen. nov. and Habetia . Also, the rather narrow fore wings that have the fore margin rather suddenly, obliquely narrowed into the narrow apical area is similar in both genera. However, Parahabetia differs from Habetia by the shapes of the male cerci and titillators and by the female ovipositor and subgenital plate. The male cerci are only little curved or nearly straight and provided in Habetia with an apical or subapical internal process that basically divides at end into two acute tips while in Parahabetia the cerci are markedly curved and provided with two or three internal projections and have the apex widened. The titillators are basically U-shaped sclerites with various modifications in Habetia , while basically straight in Parahabetia . The titillators in Parahabetia have the base widened, the central area narrow and the apical area little widened again and at end they are strongly sclerotized and carrying a curved lamellar crest at tip. The female ovipositor is straight and long or very long and with obtuse or subobtuse tip in Habetia , but rather short, curved, at end slightly recurved, and with acute tip in Parahabetia . The shape of the rather simple male titillators with an apical crest in the new genus resembles that in Philmontis Willemse, 1966 . Parahabetia differs from Philmontis by the absence of the black lateral bands on pronotum and tegmen and by the pronotum with subtruncate instead of rounded hind margin that in the type species Philmontis nigrofasciatus Willemse, 1966 covers the stridulatory area of the tegmen. There are however also species with a shorter pronotum, e.g. Philmontis lobatus Naskrecki & Rentz, 2010 . The rather short female ovipositor in Parahabetia is upcurved as in Philmontis but has the end re-curved instead of running straight into the tip as in the latter genus. Moreover, the female subgenital plate in Parahabetia is provided with a medial furrow and with large, oval, lateral swellings, which is quite different from the shape in P. nigrofasciatus that has a narrow transverse swelling in basal area, and more than apical half divided into a pair of narrow lateral lobes. From both, Habetia and Philmontis , Parahabetia gen. nov. also differs by the high number of stridulatory teeth on the rather short stridulatory file on underside of male tegmen which carries high numbers of densely packed stridulatory teeth.
Description. Rather small to medium sized species with wings that reach about the hind knees and little surpass the abdomen. Head with fastigium verticis narrow conical, moderately projecting anteriorly and separated by a transverse furrow from fastigium frontis; face long conical. Pronotum longer than wide with disc roundly merging into paranota; anterior margin subtruncate to faintly concave, posterior margin little convex or nearly truncate, with two little suspicious transverse furrows; apical area of disc little elevated with sub-rugose surface and sub-angular lateral margins; paranota rather low, swollen above auditory spiracle, ventral margin very faintly concave, anterior angle rounded, posterior angle nearly rectangular but with rounded tip, humeral sinus only slightly indicated. Prosternal spines short but distinct; mesosternal lobes triangular with a tubercle at tip or with obtuse tip; metasternal lobes with sub-truncate, oblique apical margin. Fore coxa with a long, mid coxa with a short spine at anterior margin. Femora of all legs with spines on both ventral margins; hind knee lobes bi-spinose. Tibial tympana conchate on both sides. Anterior tibiae with anterior surface convex and with indistinct rounded angles that are indicated by rows of few minute spinules. Wings little surpassing abdomen, in female hardly so; tegmen markedly narrowing from base toward subapical area; tip rounded.
Male. Tenth abdominal tergite vaulted with posterior area curved distad, flattened and divided into a pair of triangular lobes. Cerci rather long and narrow, curved and provided with two or three internal projections and modified tip. Male subgenital plate at end provided with a pair of short styli and with or without a pair of narrow apical projections that are longer than the styli. Titillators separate from each other, with basal area curved laterad, distal area long-oval or band-shaped, with a compressed crest at external side of swollen apical area.
Female. Subgenital plate with a medial furrow and lateral swellings. Ovipositor curved, but in apical area slightly re-curved, with acute tip.
Coloration in the preserved specimens almost uniformly light yellowish, ochre, or brownish, but might be discolored and a mainly green when alive. Face green. Antennal scapus and abdominal tergites in one male from Bulolo Gorge also green. In that male head and pronotum ochre with a greenish shine, and tibia with remnants of green flecks. Tegmen with light veins and darkened cells, especially in anterior area.
Etymology. The name of the new genus is built from the genus name Habetia adding the prefix para which is used to change the meaning of a word.
Key to the species and subspecies of Parahabetia View in CoL gen. nov. (males only)
1. Male cercus with three projections and a swollen, globular or oval tip ( Figs 21C, 21F, 21H View FIGURE 21 ). Male subgenital plate apart from the styli with a pair of narrow apical projections ( Figs 21D, 21G, 21I View FIGURE 21 ). Male paraproctes not strongly enlarged, rather inconspicuous. Stridulatory file on underside of left tegmen even in central area with teeth very densely packed ( Figs 22H–I View FIGURE 22 ).................................................................................................. 2.
- Male cercus with two projections and a compressed apical lobe ( Figs 22D–F View FIGURE 22 ). Male subgenital plate at end without apical projections, with styli only ( Fig. 22C–E View FIGURE 22 ). Male paraprocts strongly enlarged ( Fig. 22F View FIGURE 22 ). Stridulatory file on underside of left tegmen with teeth in central area clearly separated from each other ( Fig. 22G View FIGURE 22 )..................... P. bispinosa View in CoL sp. nov.
2. Male tenth abdominal tergite roundly incised from apical margin with incision and resulting apical lobes rather short, pointing oblique apico-mediad ( Fig. 21H View FIGURE 21 ). Male subgenital plate more elongate, with elevated disc and with flattened lateral areas that do not reach the bases of the styli; apical margin markedly incised before bases of styli and then prolonged into long, narrow projections that are markedly longer than the styli; styli inserted latero-ventrally ( Fig. 21I View FIGURE 21 ). Cerci less strongly curved, basal projection inserted somewhat away from base and with club-shaped, long-oval end; end of cercus globular ( Fig. 21H View FIGURE 21 )............................................................................. P. pictifrons pictifrons Karny, 1911 View in CoL
- Male tenth abdominal tergite long and narrowly incised and terminating into a pair of long, dorso-ventrally compressed projections with acute or obtuse tips. Male subgenital plate more semi-circular with compressed lateral areas reaching bases of styli or nearly so; apical margin before bases of styli only slightly incised with curved or with widened tips. Cerci with basal projection inserted closer to base; basal projections at end suddenly widened, with apical area either globular but little compressed or disc-shaped, in both cases with a spinule at tip; end of cercus globular or oval.........................3.
3. Apical lobes of tenth abdominal tergite with acute tip and separated by a narrow interspace from each other ( Fig. 21A View FIGURE 21 ). Male cercus with second and third internal projection inserted closely together, apical widening of cercus somewhat compressed and roughly oval ( Fig. 21C View FIGURE 21 ). Apical projections of subgenital plate at tip compressed and roundly widened ( Figs 21B, D View FIGURE 21 ). Male titillators only little curved in about mid-length, apical area narrower and apical cap less extended ( Fig. 23D View FIGURE 23 ).......................................................................................... P. pictifrons acuta ssp. nov.
- Apical lobes of tenth abdominal tergite with obtuse tip and separated by a wider interspace ( Fig. 21E View FIGURE 21 ). Male cercus with second and third internal projections inserted more away from each other; apical widening of cercus globular ( Fig. 20F View FIGURE 20 ). Apical projections of subgenital plate at tip twisted and narrow ( Fig. 21G View FIGURE 21 ). Male titillators curved in about mid-length, apical area wider and apical cap more extended ( Fig. 23E–G View FIGURE 23 ).................................... P. pictifrons obtusa ssp. nov.
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