Habetia lalibu, Ingrisch, 2021

Ingrisch, Sigfrid, 2021, Revision of the genera Habetia Kirby, 1906 and Parahabetia gen. nov. from New Guinea (Orthoptera, Tettigoniidae, Conocephalinae, Agraeciini), Zootaxa 5020 (2), pp. 201-256 : 238-239

Habetia lalibu sp. nov.

Figs 4G , 9F , 11J , 16J–L , 19G , 20D , Map 1(11).

Holotype (male): Papua New Guinea: New Guinea NE, Lalibu, elev. 2300 m (6°16’S, 143°59’E), 8–14.iv.1968, leg. J.L. Gressitt & Mea —depository: Bernice B. Bishop Museum, Honolulu ( BPBM).

Other specimens studied: same data as holotype 1 male (paratype, BPBM) .

Diagnosis. H. lalibu sp. nov. belongs to a group of species that have the male cerci with a simple, rounded projection on the dorsal margin just before the cercus is curved mediad into the compressed apical internal process instead of a prolonged, wavy projection as in H. bivittata sp. nov. It differs from the other three species with similar basic structure of the cerci ( H. kondiu sp. nov., H. simbai sp. nov., and H. wau sp. nov.) by the dorsal projection that is swollen instead of flattened and the internal projection is almost parallel sided with a pair of tubercles of equal size at tip instead of with distinctly converging margins and with a pair of acute teeth or with a curved process as in H. wau . The male tenth abdominal tergite has a narrow and short apical incision as in H. simbai but different from the long and wide apical incision as in H. kondiu and H. wau ; however, the tenth tergite of H. lalibu has the apical margin concave on both sides of the incision and with a short obtuse projection on both sides, while in H. simbai the apical margin on both sides of the apical incision is obliquely truncate. The male titillators of H. lalibu have the proximal projections subfused and with upcurved lateral margins, forming together a wide furrow, and the distal projections are widened and strongly curved, while in the two species with similar titillators, H. simbai and H. kondiu , the proximal projections are separate, and the distal projections forming moveable arms or are separated from the titillator stem.

Description. Coloration of face of general color; mandibles darkened at base, otherwise of general color; in one male part of antennal scrobae in external half of ventral margin black ( Fig. 4G ). Femora with the following number of spines on ventral margins: (1) a 6, p 6–9; (2) a 5–6, p 3; (3) a 9–11, p 10–11 (n=2).

Male. Stridulatory file on underside of left tegmen 1.6 mm long, with 87–88 teeth that are at both ends very narrow with 15–16 basal and 23–28 apical teeth, in central 1 mm of file with 48–49 teeth, of which are 21–22 largest teeth in central 0.5 mm, n = 2 ( Fig. 19G ). Tenth abdominal tergite in about basal half with sub-parallel but little wavy, down-curved lateral areas, afterward with approaching lateral margins, forming a pair of obtuse triangular lobes, in middle of concave hind margin between both triangular lobes roundly incised ( Fig. 9F ). Epiproct with medial furrow and apical margin rounded. Cerci elongate, rounded with slightly approaching lateral margins; at end suddenly bent in a roughly 90°-angle mediad into compressed internal projections that end into a pair of very short, obtuse tubercles; cercus stem just before apical curvature with a semi-circular but little swollen dorsal crest; internal surface of cercus little excavated around curvature ( Fig. 9F ). Subgenital plate in lateral view curved throughout; in basal area elevated and provided with a large oval swelling that is in situ probably hidden by the ninth tergite, afterward with dorsal rim and a lateral furrow in about mid-length of plate; ventral surface from base with a large triangular and membranous incision, lateral margins little swollen; in about apical third subgenital plate deep roundly, in middle little angularly incised and up-curved, in apical area consisting only of the stiffened lateral margins that are faintly approaching towards tip; styli short, inserted at underside of the narrow apical extensions of the subgenital plate ( Fig. 11J ). Titillators at base slightly widened and faintly curved, basal branches subfused, separated by a fine membranous suture; in upper area divided into proximal and distal branches; proximal branches stick together, curved proximo-ventrad, have lateral areas upcurved, and tip shortly bi-lobate; distal branches around dividing point fully fused and black, after a short distance divided from each other and running more dorsad and laterad; afterward they are strongly curved ventrad and little approaching each other while becoming markedly wider and reddish brown, at end they are compressed, becoming roughly circular plates with granular surface and margin and are slightly projecting distad ( Figs 16J–L ). There is also a pair of large elongate lateral sclerites with three flattened expansions from a little wavy central spindle, and the membranes covering the titillators dorsally are provided with a pair of small, rounded caps.

Measurements (2 males).—Body w/wings: 39–41; body w/o wings: 32–33; pronotum: 8.2–8.5; tegmen: 28.5– 29.5; hind femur: 20.0–21.5; tegmen width: 4.5–5.0 mm.

Etymology. The new species is named after the type locality, noun in apposition.

Figures

FIGURE 4. Frons of Habetia species: A–C, H. bivittata sp. nov., variation of the black marks (A Goroka male, B Mt. Wilhelm male, C Keltiga female); D, H. dentata sp. nov. female (Saidor); E, H. kondiu sp. nov. female (Kubor Range); F, H. pinnigera sp. nov. female (Madang); G, H. lalibu sp. nov. male (Lalibu); H, H. simbai sp. nov. female (Simbai Valley); I, H. nigricauda sp. nov. female (Popondetta); J, H. quatrispina sp. nov. female (Pionierlager); K, H. wau sp. nov. male (Nami Creek); L, H. sororcula sp. nov. female (Ifar). Scales 5 mm.

FIGURE 9. Male abdominal apex in apical (A), dorsal (B, D–E), and apico-dorsal view (F–H); left cercus (C).—A–C, Habetia bivittata sp. nov. (A Goroka, B Daulo Pass, C Mt Wilhelm); D–E, Habetia kondiu sp. nov. (D Kubor Range, E Wabag); F, Habetia lalibu sp. nov. (Lalibu); G, Habetia wau sp. nov. (Nami Creek); H, Habetia simbai sp. nov. (Simbai Valley).—Scales 1 mm.

FIGURE 11. Male subgenital plate and cerci of Habetia species in ventral view (J in ventro-apical view).—A, H. tuta sp. nov. (Rosensee); B, H. pallida sp. nov. (Ifar); C, H. pedala sp. nov. (Toricelli Mts); D, H. curvata sp. nov. (Albatros Bivak); E, H. imitatrix Karny, 1912 holotype (Bongu); F, H. dentata sp. nov. (Saidor); G, H. bivittata sp. nov. (Goroka); H, H. simbai sp. nov. (Simbai Valley); I, H. kondiu sp. nov. (Kondiu); J, H. lalibu sp. nov. (Lalibu); K, H. wau sp. nov. (Nami Creek). Scales 5 mm (A–E), 1 mm (F–K).

FIGURE 16. A–C, Habetia simbai sp. nov. (Simbai Valley); D–I, Habetia kondiu sp. nov. (D–E Kubor Range, F–I Wabag); J–L, Habetia lalibu sp. nov. (Lalibu).—Male titillators in distal view (A, D, G, J), in proximal view (B, E, H, K), in oblique distal view (C), in ventral view (F), and in oblique lateral view (I, L).—Abbreviations: aa apical arm, cb curved base, dp dorsal projection, ls lateral sclerite, pp proximal projection, sa separate apical arm. The arrow points at the hinge between the central and apical area of the titillator. Scales 1 mm.

FIGURE 19. Stridulatory files on underside of left tegmen of males of Habetia species: A–D, H. bivittata sp. nov. (A Sinofi, B Daulo Pass, C–D Goroka); E–F, H. kondiu sp. nov. (E Kubor Range, F Wabag); G, H. lalibu sp. nov. (Lalibu); H, H. simbai sp. nov. (Simbai Valley). Scales 1 mm.

FIGURE 20. Habetia sp. males: Habitus in dorsal view with left wings spread.—A, H. bivittata sp. nov. (Moife); B, H. kondiu sp. nov. (Wabag); C, H. dentata sp. nov. (Saidor); D, H. lalibu sp. nov. (Lalibu); E, H. pilleata sp. nov. (Wau); F, H. simbai sp. nov. (Simbai Valley); G, H. wau sp. nov. (Wau); H, H. tuta sp. nov. (Lake Chambri).—All scales 10 mm.