Scopaeus Erichson, Revised Definition

Herman, Lee, 2023, Generic Revisions Of The Scopaeina And The Sphaeronina (Coleoptera: Staphylinidae: Paederinae: Lathrobiini), Bulletin of the American Museum of Natural History 2023 (460), pp. 1-195 : 61

publication ID	https://doi.org/ 10.1206/0003-0090.460.1.1
DOI	https://doi.org/10.5281/zenodo.8109740
persistent identifier	https://treatment.plazi.org/id/FA4F87B1-E12D-FFC7-0BB5-C658FDF6F9FA
treatment provided by	Felipe
scientific name	Scopaeus Erichson, Revised Definition
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Scopaeus Erichson, Revised Definition View in CoL

Figures 1–8 View FIGS , 14, 15 View FIGS , 23–239 View FIGS View FIGS View FIGS View FIGS View FIGS View FIGS View FIGS View FIGS View FIGS View FIGS View FIGS View FIGS View FIGS View FIGS View FIGS View FIGS View FIGS View FIGS View FIGS View FIGS View FIGS View FIGS View FIGS View FIGS View FIGS View FIGS View FIGS View FIGS View FIGS View FIGS View FIGS View FIGS

FIGS. 1–8. Scopaeus. Figs. 1–4. habitus. 1. S. chiriquensis sp. grp., S. sp. 2. S. frater. 3. nitidus sp. grp., S. laxus. 4. S. “reticulate” sp. grp., S. sp. 5, 6. S. opacus sp. grp., S. elaboratus. 5. Habitus. 6. Labrum, dorsal, setae omitted. 7, 8. S. nevermanni sp. grp., S. sp. 7. Head, dorsal, marginal setae omitted. 8. Habitus, prolegs not visible.

FIGS. 11–16. Scopaeina. Head, trichobothrial variation. 11. Hyperscopaeus sp. 12. Micranops sp. 13. Orus punctatus. 14. Scopaeus “reticulate” sp. grp. 15. Scopaeus chiriquensis sp. grp. 16. Trisunius spathulatus.

FIGS. 23–28. Compound eyes, corneal surface. Scopaeus. 23. S. chiriquensis sp. grp. 24. S. debilis sp. grp. (S. debilis). 25. S. debilis sp. grp. (S. filiformis). 26. S. elegans sp. grp. (S. cameroni). 27. S. elegans sp. grp. (S. persicus). 28. S. gracilis sp. grp. (S. gracilis).

FIGS. 29–34. Compound eyes, corneal surface. Scopaeus. 29. S. laevigatus sp. grp. (S. laevigatus). 30. S. limbatus sp. grp. 31. S. longicollis sp. grp. 32. S. minutus sp. grp. (S. chalcodactylus). 33. S. mutatus sp. grp. 34. S. nevermanni sp. grp.

FIGS. 35–40. Compound eyes, corneal surface. Scopaeus. 35. S. nitidus sp. grp. 36. S. obscuripes sp. grp. (S. likovskyi). 37. S. ooderes sp. grp. (S. ooderes). 38. S. opacus sp. grp. (S. opacus). 39. S. punctatellus sp. grp. (S. punctatellus). 40. S. “reticulate” sp. grp.

FIGS. 41–46. Compound eyes, corneal surface. Scopaeus. 41. S. rotundiceps sp. grp. 42. S. ryei sp. grp. 43. S. sericans sp. grp. 44. S. signifer sp. grp. (S. bicolor). 45. S. similis sp. grp. (S. similis). 46. S. sulcicollis sp. grp. (S. sulcicollis).

FIGS. 47–60. Scopaeus. Head, Prothorax, Mouthparts. 47–50. Head. 47. S. picipes. 48. S. chiriquensis sp. grp. (S. sp.). 49. S. opacus sp. grp. (S. sp.). 50. S. nitidus sp. grp. (S. sp.). 51–57. Prothorax. 51, 52. S. picipes. 51. Ventral. 52. Lateral. 53, 54. S. chiriquensis sp. grp. (S. sp.). 53. Ventral. 54. Lateral. 55–57. S. nitidus sp. grp. (S. sp.). 55. Ventral. 56. Lateral. 57. S. opacus sp. grp. (S. sp.), ventral. 58–60. S. chiriquensis sp. grp. (S. sp.). 58. Labrum. 59. Labrum. 60. Labium, ventral.

FIGS. 61–80. Scopaeus. Mandibles, Genital segments. 61–71. Mandibles, dorsal. 61, 62. S. picipes. 63, 64. S. chiriquensis sp. grp. (S. sp.). 65, 66. S. nitidus sp. grp. (S. sp.). 67–71. S. opacus sp. grp. (S. spp.). 72–77. Tergites IX, X. 72, 73. S. picipes. 72. Male. 73. Female. 74, 75. S. nitidus sp. grp. (S. sp.). 74. Male. 75. Female. 76–79. S. chiriquensis sp. grp. (S. sp.). 76. Male. 77. Female. 78. Lateral gonocoxal plates. 79. Sternite IX, male. 80. S. picipes, Lateral gonocoxal plates.

FIGS. 81–86. Scopaeus. S. chiriquensis sp. grp., (S. sp.). 81. Head, trichobothrium. 82. Mesoventrite. 83. Metaventrite. 84. Metaventrite, left file. 85. Mesofemur, left base, plectral ridge. 86. Sternites II and III.

FIGS. 87–92. Scopaeus. 87–88. S. chiriquensis sp. grp., (S. sp.), Sternite IV. 87. Midanterior margin. 88. Midanterior margin enlarged, glandular(?) depression. 89–92. S. debilis sp. grp., (S. filiformis). 89. Pteroventrites. 90. Metaventrite, right file. 91. Sternites II and III. 92. Mesofemur, right base, plectral ridges.

FIGS. 93–98. Scopaeus. 93–94. S. debilis sp. grp., (S. filiformis). 93. Mesoventrite and anterior metaventrite. 94. Metaventrite, midapex, metakatepisternal processes. 95–98. S. dissimilis sp. grp., (S. sp.). 95. Metaventrite, right file. 96. Mesoventrite and anterior metaventrite. 97. Metaventrite, posterior, metakatepisternal processes. 98. Prothorax, ventral.

FIGS. 99–103. Scopaeus. 99–100. S. dissimilis sp. grp., (S. sp.) 99. Head, dorsal. 100. Labrum. 101–103. S. elegans sp. grp., (S. cameroni). 101. Metaventrite. 102. Metaventrite, right file. 103. Mesoventrite.

FIGS. 104–109. Scopaeus. 104–106. S. elegans sp. grp., (S. cameroni). 104. Metaventrite. 105. Metaventrite, left file. 106. Mesofemur, left base, plectral ridges. 107–108. S. elegans sp. grp., (S. persicus). 107. Mesofemur, left base, plectral ridges. 108. Trichobothrium, head. 109. S. gracilis sp. grp., (S. gracilis), Mesoventrite.

FIGS. 110–115. Scopaeus. 110–112. S. gracilis sp. grp., (S. gracilis). 110. Metaventrite and apex of mesoventrite. 111. Metaventrite, right file. 112. Mesofemur, right base, plectral ridges. 113–115. S. laevigatus sp. grp., (S. laevigatus). 113. Mesofemur, right base, plectral ridges. 114. Pteroventrites. 115. Metaventrite, right file, enlarged.

FIGS. 116–122. Scopaeus. S. laevigatus sp. grp. 116–118. (S. laevigatus). 116. Sternites II and III. 117, 118. Sternite IV, midanterior margin. 117. Glandular(?) region. 118. Glandular(?) pore. 119–122. (S. subfasciatus). 119. Metaventrite. 120. Metaventrite, enlarged section of left file. 121. Mesofemur, right base, plectral ridges. 122. Mesoventrite.

FIGS. 123–128. Scopaeus. 123. S. laevigatus sp. grp. (S. subfasciatus), Sternite IV, midanterior enlarged, glandular(?) opening. 124–128. S. limbatus sp. grp., (S. sp.). 124. Mesoventrite. 125. Metaventrite. 126. Metaventrite, enlarged section of left file. 127. Mesofemur, right base, plectral ridges. 128. Sternites II and III.

FIGS. 129–135. Scopaeus. 129–130. S. limbatus sp. grp. (S. sp.). 129. Sternite IV, anterior. 130. Sternite IV, midanterior, enlarged glandular(?) depression. 131–133. S. longicollis sp. grp. (S. sp.). 131. Metaventrite. 132. Mesofemur, right base, plectral ridges. 133. Metaventrite, left file. 134–135. S. minimus sp. grp. (S. minimus). 134. Sternite IV, midanterior. 135. Sternite IV, midanterior enlarged, glandular(?) depression and pore.

FIGS. 136–141. Scopaeus. 136–140. S. minimus sp. grp. (S. minimus). 136. Mesoventrite. 137. Metaventrite. 138. Metaventrite, right file. 139. Mesofemur, left base, plectral ridges. 140. Sternites II and III. 141. S. minutus sp. grp. (S. pusillus), Mesoventrite.

FIGS. 142–147. Scopaeus. 142–145. S. minutus sp. grp. (S. pusillus). 142. Metaventrite. 143. Metaventrite, left file. 144. Mesofemur, right base, plectral ridge. 145. Sternites II and III. 146–147. S. mutatus sp. grp. (S. sp.). 146. Mesofemur, right base, plectral ridges. 147. Sternites II and III.

FIGS. 148–153. Scopaeus. S. mutatus sp. grp. (S. sp.). 148. Mesoventrite. 149. Metaventrite. 150. Metaventrite, anterior left file. 151. Metaventrite, posterior left file. 152. Sternite IV, midanterior. 153. Sternite IV, midanterior enlarged, glandular(?) depression and pore.

FIGS. 154–160. Scopaeus. S. nevermanni sp. grp. (S. sp.). 154. Metaventrite. 155. Metaventrite, right file. 156. Mesoventrite. 157. Mesofemur, right base, plectral ridges. 158. Aedeagus, dorsal. 159. Labrum. 160. Labium, hypopharynx, tripartite ligular lobe.

FIGS. 161–167. Scopaeus. S. nitidus sp. grp. (S. sp.). 161. Head. 162. Mesofemur and coxa, plectral ridges. 163. Pteroventrites. 164, 165. Metaventrite, right file. 166. Mesoventrite. 167. Sternites II and III.

FIGS. 168–173. Scopaeus. 168–170. S. nitidus sp. grp. (S. sp.). 168. Labium, hypopharynx. 169. Labrum, epipharynx. 170. Mesofemur, left base, plectral ridges. 171–173. S. obscuripes sp. grp. (S. likovskyi). 171. Mesofemur, right base, plectral ridges. 172. Metaventrite. 173. Metaventrite, left file.

FIGS. 174–180. Scopaeus. 174. S. obscuripes sp. grp. (S. likovskyi). Mesoventrite. 175–180. S. ooderes sp. grp., (S. ooderes). 175. Mesofemur, right base, plectral ridges. 176. Mesoventrite. 177. Metaventrite. 178. Metaventrite, left file. 179. Sternites II and III. 180. Sternite IV, midanterior enlarged, glandular(?) pore.

FIGS. 181–186. Scopaeus. S. opacus sp. grp., (S. opacus). 181. Mesoventrite. 182. Mesoventrite, median depression, enlarged. 183. Metaventrite. 184. Metaventrite, left file. 185. Mesofemur, right base, plectral ridges. 186. Labrum, epipharynx.

FIGS. 187–192. Scopaeus. 187. S. opacus sp. grp., (S. opacus). Labium, hypopharynx. 188–192. S. punctatellus sp. grp.; S. punctatellus. 188. Mesoventrite. 189. Metaventrite. 190. Metaventrite, right file. 191. Mesofemur, right base, plectral file. 192. Sternites II and III.

FIGS. 193–198. Scopaeus. S. “reticulate” sp. grp. (S. sp.). 193. Head. 194. Head, right, anterolateral, trichobothrium. 195. Metaventrite. 196. Metaventrite, left file. 197. Mesofemur, right base, plectral ridges. 198. Mesoventrite.

FIGS. 199–204. Scopaeus. 199–200. S. “reticulate” sp. grp. (S. sp.). 199. Sternite IV. 200. Sternite IV, enlarged midanterior, glandular(?) pore. 201–204. S. rotundiceps sp. grp. (S. sp.). 201. Metaventrite. 202. Metaventrite, left file. 203. Mesofemur, left base, plectral ridges. 204. Sternites II and III.

FIGS. 205–210. Scopaeus. S. rotundiceps sp. grp. (S. sp.). 205. Head. 206. Head, dorsal, left lateral, bothrium. 207. Mesoventrite. 208. Mesofemur, right base, plectral ridges. 209. Metaventrite. 210. Metaventrite, left file.

FIGS. 211–216. Scopaeus. S. ryei sp. grp. (S. sp.). 211. Mesoventrite. 212. Mesofemur, left basal, plectral ridges. 213. Metaventrite, anterior right side and middle. 214. Metaventrite, right file. 215. Sternites II and III. 216. Sternite IV, midanterior enlarged, glandular(?) depression.

FIGS. 217–222. Scopaeus. 217–221. S. sericans sp. grp. (S. sp.). 217. Mesoventrite. 218. Mesofemur, left base, plectral ridges. 219. Metaventrite. 220. Metaventrite, left file. 221. Sternites II and III, cropped. 222. Scopaeus signifer sp. grp. (S. bicolor). Mesoventrite.

FIGS. 223–228. Scopaeus. 223–225. S. signifer sp. grp. (S. bicolor). 223. Metaventrite. 224. Metaventrite, left file. 225. Mesofemur, left base, plectral ridges. 226–228. S. similis sp. grp., (S. similis). 226. Mesofemur, right base, plectral ridges. 227. Metaventrite. 228. Metaventrite, right file.

FIGS. 229–235. Scopaeus. 229–231. S. similis sp. grp. (S. similis). 229. Mesoventrite. 230. Sternites II and III. 231. Sternite IV, midanterior enlarged, glandular(?) pore. 232–235. S. sulcicollis sp. grp., (S. sulcicollis). 232. Sternite IV. 233. Sternite IV, midanterior enlarged, glandular(?) pore. 234. Mesoventrite. 235. Sternites II and III.

FIGS. 236–239. Scopaeus. S. sulcicollis sp. grp. (S. sulcicollis). 236. Metaventrite. 237. Metaventrite, right file. 238. Mesofemur, left. 239. Mesofemur, left base, plectral ridges.

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Taxonomy

Kingdom	Animalia
Phylum	Arthropoda
Class	Insecta
Order	Coleoptera
Family	Staphylinidae

Scopaeus Erichson, Revised Definition

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