Pamphilius planifrons Beneš, 1976

Shinohara, Akihiko, Kramp, Katja & Taeger, Andreas, 2022, The Pamphiliinae of the Russian Far East and Korea (Hymenoptera, Pamphiliidae), Zootaxa 5167 (1), pp. 1-251 : 93-94

Pamphilius planifrons Beneš, 1976

( Figs 111 , 112 ) (https://doi.org/10.6084/m9.figshare.11405274)

Pamphilius planifrons Beneš, 1976: 164 ; Shinohara, 1988b: 184; Shinohara, 2002b: 427; Shinohara, 2004: 264; Shinohara & Lelej, 2007: 932, 940; Taeger et al., 2010: 90; Sundukov & Lelej, 2012: 109; Sundukov, 2017: 105; Lee et al., 2019: 11.

Material examined. Eighteen specimens, including the holotype, all from the Russian Far East ( Beneš 1976; Shinohara 1988, see comments below about the males; present work). New collection data: RUSSIA: Primorskij Kraj: 1♂, Ussurijskij Reserve, 21–26. V. 1994, A. Shinohara ( Fig. 112 , NSMT); 6♀, same locality, 9–12. VI . 1995 , A. Shinohara ( Fig. 111 , NSMT); 2♀, same locality, 9–12. VI . 1995 , A. Lelej ( NSMT); 1♂, Okeanskaja , Vladivostok, 28. V . 1994 , A. Shinohara ( NSMT); 1♀, Anisimovka , 300m, 1. VI . 1994 , A. Shinohara ( NSMT); 1♀, Uglekamensk, 2. VI . 1994 , A. Shinohara ( NSMT); 2♀, Valley of Tigrovaja River , 180m, 12km N of Partizansk, 6. VI . 1994 , A. Shinohara ( NSMT); 1♀ (DEI-GISHym 32052), Arsenyev , Ski-Base Bodrost, 200m, 44.122°N 133.270°E, 25. V. 2016, K. Kramp, M. Prous & A. Taeger, RU017 ( SDEI); 2♀ (DEI-GISHym 12992 & 32040), Zolotaya Dolina E, 40m, 42.943°N 133.161°E, 10. VI. 2016, K. Kramp, M. Prous & A. Taeger, in alcohol, RU046 ( SDEI).

Distribution. Russia (Primorskij Kraj).

Host plant. Unknown.

Remarks. This species is a member of the P. thorwaldi complex of the P. stramineipes subgroup of the P. vafer group ( Shinohara 2002b). Shinohara (1988b) recorded this species from South Korea based on two male specimens as P. planifrons . These two males actually belong to the new species P. belokobylskiji described above, and South Korea is here excluded from the distribution of P. planifrons . Pamphilius planifrons is most closely allied to P. rhoae , as discussed under the latter species. In the molecular analyses using COI and NaK sequences ( Figs 149 , 161 ), the relationships between P. belokobylskiji , P. planifrons and P. rhoae were poorly resolved.

Figures

FIGURE 111. Pamphilius planifrons, female, Russia – (a) dorsally, (b) lateroventrally, (c) head & thorax dorsally, (d) head & thorax ventrolaterally, (e), (f) antenna, (g) sawsheath laterally, (h) claw.

FIGURE 112. Pamphilius planifrons, male, Russia – (a) dorsally, (b) lateroventrally, (c) head & thorax dorsally, (d) head & thorax ventrolaterally, (e) face, (f) antenna, (g) genitalia dorsally, (h) genitalia ventrally, (i) claw.

FIGURE 149. Condensed ML tree based on CO1 gene sequences, with details of Pamphilius vafer group, part 2.

FIGURE 161. Condensed ML tree based on NaK gene sequences, with details of Pamphilius vafer group, part 2.