Biocellata bifida, Davis & Gentili-Poole & Mitter, 2008

Davis, Steve R., Gentili-Poole, Patricia & Mitter, Charles, 2008, A revision of the Cossulinae of Costa Rica and cladistic analysis of the world species (Lepidoptera: Cossidae), Zoological Journal of the Linnean Society 154 (2), pp. 222-277 : 244-245

BIOCELLATA BIFIDA SP. NOV.

( FIGS 22 , 55, 56 , 90 , 136)

Male ( Fig. 22 ): Forewing length: 15–17 mm.

Head: Antenna with shaft and rami creamy white; labial palpus brown dorsally, creamy white with some light brown ventrally; frons creamy white; vertex creamy white, irrorated with brownish grey.

Thorax: Patagia creamy white, heavily irrorated with brownish grey; dorsum and tegula with scales creamy white and creamy white basally, greyish brown apically; pleura cream; venter mostly brown with some cream. Legs with femur and tibia cream, irrorated with brown; fore tarsi cream, light brown, and brown; mid and hind tarsi brown, banded creamy brown distally on each tarsomere. Forewing creamy white with numerous grey transverse striae throughout wing dorsally; the two most distinct and largest of these striae located basally and medially along wing; subcostal region creamy white and mostly lacking striae; numerous short, transverse, grey and dark brown striae along costal margin; two dark brown to fuscous spots subterminally; fringe banded light creamy gold and brownish grey; forewing mostly light brown ventrally with some cream; numerous short, transverse, dark brown striae along costal margin. Hindwing creamy white dorsally; some creamy light brown along apical margin; fringe can be slightly banded grey and creamy white, but mostly creamy white; hindwing mostly creamy white ventrally.

Abdomen: Mostly cream and creamy white dorsally; cream and brown ventrally; A8 cream with slight light brown.

Male genitalia ( Figs 55, 56 , 90 ): Valva subtrapezoidal, approximately 0.9¥ length of genital capsule; gnathos–uncus expanse approximately 0.5¥ length of genital capsule; saccular process absent, with sacculus heavily sclerotized and bearing flattened rugosities; juxta process small and slender; bridge of gnathos strongly bifurcate, with extending arms slightly upcurved and apex of each arm acute; each arm with a serrate ridge ventrally; uncus shallowly bifurcate, with apices acute. Aedoeagus with sclerotized plate formed mediad, where juxta attaches; rostellum long and slender, inwardly curving, with apex acuminate as viewed laterally; a patch of triangular spines on one side, along margin where vesica emerges from aedoeagus; rostellum becoming slightly more narrow than aedoeagus as viewed ventrally, with apex rounded.

Female: Unknown.

Distribution: Known only from Heredia and Limon Provinces, Costa Rica.

Figures

Figures 20–31. Adults: 20, male, Biocellata alfarae; 21, female, Biocellata alfarae; 22, male, Biocellata bifida; 23, male, Biocellata davisorum; 24, male, Spinulata acutipennis; 25, female, Spinulata acutipennis; 26, male, Spinulata manes; 27, female, Spinulata manes; 28, male, Spinulata oblongata; 29, male, Spinulata quasivinnea; 30, female, Spinulata quasivinnea; 31, male, Simplicivalva ampliophilobia. Scale bar = 10.0 mm.

Figures 53–64. Male genitalia. Biocellata alfarae: 53, genital capsule; 54, aedoeagus. Biocellata bifida: 55, genital capsule; 56, aedoeagus. Biocellata davisorum: 57, genital capsule; 58, aedoeagus. Spinulata acutipennis: 59, genital capsule; 60, aedoeagus. Spinulata manes: 61, genital capsule; 62, aedoeagus. Spinulata oblongata: 63, genital capsule; 64, aedoeagus.

Figures 89–94. Male genitalia, lateral view of genital capsule: 89, Biocellata alfarae; 90, Biocellata bifida; 91, Biocellata davisorum; 92, Spinulata acutipennis; 93, Spinulata manes; 94, Spinulata oblongata. Scale bar = 1.0 mm.