Ungla banksi Tauber, new replacement name

Tauber, Catherine A., Sosa, Francisco, Albuquerque, Gilberto S. & Tauber, Maurice J., 2017, Revision of the Neotropical green lacewing genus Ungla (Neuroptera, Chrysopidae), ZooKeys 674, pp. 1-188 : 12-18

Ungla banksi Tauber, new replacement name Figs 11, 12, 13, 14, 15, 16

Nothochrysa tibialis Banks, 1914. Canad. Entomol. 46: 26-27; "Rio Longo, Bolivia, 750 m. (Fassl.)". Penny 1977: 28 (list); Oswald 2015 [catalog listing as "Available, invalid, species, unreplaced junior homonym …”]. Chrysopa tibialis (Banks), Brooks and Barnard 1990: 280 [as " ' Chrysopa ' incertae sedis (? Leucochrysa )"]. Junior homonym, preoccupied by Nothochrysa tibialis Navás 1913 [now Italochrysa tibialis ( Navás, 1913)]. Holotype (Figs 11-15). In accordance with Recommendation 60A of the International Code of Zoological Nomenclature, we retain Banks’ type specimen (MCZ Type #12021, Male) as the holotype. Label data: (1) "Rio Zongo 750 m / Bolivia"; (2) "Fassl / coll"; (3) "Collection / N. Banks"; (4) “type” [red]; (5) "Type / 12021" [red]; (6) " Nothochrysa / tibialis Bks / type" [large, white, red border]; (7) "HOLOTYPE / Ungla banksi / Tauber et. al 2017 " [red]. We could not find the locality "Rio Longo" in Bolivia. According to Dr. Newton of the Field Museum of Natural History, Chicago (quoted in an email from A. Contreras-Ramos, Universidad Nacional Autónoma de México), the term probably is a misspelling of "Rio Zongo", a river in La Paz province, 15°43'S, 67°41'W. Thus, we presume that Banks misread Fassl’s handwritten “Z” as an “L”. Justification for name change. The species is a valid biological entity within the genus Ungla ; its original name ( Nothochrysa tibialis Banks, 1914) was preoccupied ( Oswald 2015). Because the name is a primary homonym, Articles 57.2 and 60.1 of the International Code of Zoological Nomenclature require the designation of a replacement name. In recognition of Nathan Banks’ discovery and original description of the species, we offer the name Ungla banksi as the replacement.

Diagnosis.

This species is distinguished by pronounced, dark brown body markings; dark brown or black marking on frontal base of foretibia; tall wings with rounded apices, forewing with very dark venation, hindwing venation less dark; stark, shiny, dark brown to black markings on dorsum of scape and vertex; moderately enlarged abdominal spiracles; distal setae of S8+9 larger than those on ventral or proximal region, but not exceptionally robust; those on narrow strip of distolateral margin flanged. Forewing 13.0 mm long, 5.5 mm wide; hindwing 12.1 mm long, 4.4 mm wide.

Redescription.

Head cream-colored, with vertex smooth, shiny, with inverted U-shaped marking prominent, dark brown, broad, separated anteromesally, not extending anteriorly to area between scapes; dorsal antennal fossa pale; area between eyes and posterior half of vertex cream-colored, unmarked; frons, clypeus cream-colored, without markings; gena with large, brown mark from eye onto clypeus; tentorial pits amber. Antenna with scape cream-colored, unmarked ventrally, laterally, entirely dark brown dorsally; pedicel, flagellum dark brown to black; maxillary palp, labial palp with basal segments pale, ultimate segment dark brown.

Prothorax with pair of wide, brown stripes laterally, extending mesally and becoming reddish along transverse furrow, pale green mesally; transverse furrow in posterior region, not reaching lateral margins of segment, golden to dark brown setae throughout. Mesothorax, metathorax with pair of broad, dark brown stripes laterally, pale green mesally. Legs pale, unmarked except for small dark brown spot near anterior base of foretibia. Measurements: head width: 1.5 mm; ratio head width: eye width: 3.0-3.2: 1; prothorax width: 1.2 mm, length: 0.9 mm.

Forewing, hindwing broad, with round apices. Forewing with venation heavy, especially at base of R, Rs, M, Cu; stigma lightly opaque, with four dark brown subcostal crossveins below, area surrounding subcostal crossveins marked with dark brown; longitudinal veins mostly green, with brown at intersections; transverse veins, crossveins mostly brown or dark brown, most with dark brown suffusion on surrounding membrane. Forewing 13.4-13.9 mm long, 5.3-5.5 mm wide (ratio, L: W = 2.4-2.6: 1); height of tallest costal cell 1.2 mm (cell number 8); length of first intramedian cell 1.1-1.2 mm; 11-12 radial cells (closed cells between R and Rs); third gradate cell 2.3 mm long, 0.4 mm wide (ratio, L: W = 5.5: 1); fourth gradate cell 2.3 mm long, 0.4 mm wide (ratio, L: W = 5.1: 1); 3-4 Banksian cells (b cells), 4 b’ cells; 7 inner gradates, 6-8 outer gradates, sometimes a middle gradate vein. Hindwing with venation delicate, not swollen, light green, except C, Sc, costal veinlets brown; 12.1-12.4 mm long, 4.4 mm wide (ratio, L: W = 2.7-2.8: 1), 11-12 radial cells, 2-3 Banksian (b) cells, 4 b’ cells, 5-6 inner gradates, 6-7 outer gradates.

Male: Abdomen with slightly enlarged spiracles (e.g., A7: spiracle diameter ~0.10 × length of sternite); T9+ectoproct relatively long (~two thirds times length of T7), with dorsal invagination deep (~0.6 × dorsal length of T9+ect), margins of invagination almost straight, base rounded; dorsal margin of T9+ect rounded distally (above anus), often compressed, thus appearing straight; posterior margin of ectoproct short, slightly concave; dorsal apodeme extending along ventral margin of entire segment, lightly sclerotized, posterior corner extending, bending mesally to form small, rounded knob; callus cerci large, ovate, with entire margin lightly sclerotized. S8+9 fused, with line of fusion hardly perceptible; dorsum tapering abruptly to shallow platform at ¾ distance to tip of segment, dorsal margin regular, ventral apodeme well sclerotized; terminus concave to flat, extending distally, slightly beyond tip of T9+ect, flat, distal margin upturned, heavily sclerotized; terminal setae enlarged, with flange-line protrusions basally. Gonarcus arcuate, with bridge slightly angled dorsally, arms robust, elongate, slightly curved inward, rounded distally, mesal section with digitiform process extending posteriorly and inward toward gonosaccus; mediuncus with heavy base, ridged dorsally, tapering to elongate, blunt terminus; gonosaccus bilobed, each lobe with elongate patch of gonosetae; gonosetae arising from enlarged setal bases, dense, large at tip of lobe, becoming smaller, scarcer basally near base of mediuncus; hypandrium internum not found.

Variation.

The thickness and the depth of the brown coloration of the veins on the forewing were considerably more robust in the male specimen than in the female.

Known distribution.

BOLIVIA: Departments of La Paz, Santa Cruz.

Specimens examined

(in addition to type above). Bolivia. Santa Cruz, Florida, 11 km. N.E. Achira, 1800 m, 3/XI/1999, cloud forest, Malaise trap, C. Porter & L. Stange (1F, FSCA).