Prothyma (Genoprothyma) confluenteangustata Horn, 1924

Wiesner, Jürgen & Anichtchenko, Alexander, 2023, The Philippines species of the genus Prothyma Hope, 1838 (Coleoptera: Cicindelidae) Part 1: Subgenera Prothyma Hope, 1838, Genoprothyma Rivalier, 1964 and Parasymplecthyma subgen. nov., Zootaxa 5315 (1), pp. 1-36 : 9-12

Prothyma (Genoprothyma) confluenteangustata Horn, 1924 , stat. nov. ( Figs 5 , 6 )

Prothyma hopkinsi confluente -angustata Horn, 1924: 91.

Type locality. “ Insula Negros , Cuernos Mts. ” .

Prothyma Hopkinsi confluente -angustata : Horn 1926: 99.

Prothyma confluente-angustata: Horn 1934: 145 , f. 2, 146.

Prothyma (Genoprothyma) Hopkinsi confluente-angustata: Rivalier 1964: 156 .

Prothyma (Genoprothyma) hopkinsi confluenteangustata: Wiesner 1980: 124 ; Wiesner 1992: 58; Cabras, Cabigas and Wiesner 2016: 192; Wiesner 2020: 138.

Type material. Lectotype (here designated) male: “ Cuernos Mts. / Negros Baker // Syntypus [printed, red] // Type / W. Horn // LECTOTYPE / Prothyma confluente - / angustata W. Horn, 1924 / design. Wiesner & / Anichtchenko 2022 [printed, red]” ( SDEI).

Other material examined. 1 female: “ Cuernos Mts. / Negros Baker” ( SDEI) ; 2 males: “ Philippines, Negros / Canlaon Mt. / 15–31.vii.2015 ” ( DUBC) ; 1 male and 1 female with same label data ( JWc) ; 1 male: “ Philippines, Negros / Canlaon Mt. / 900m, 15.vii.2015 ” ( DUBC) ; 1 female: “ Philippines, Negros / Canlaon Mt. / 15–31.vii.2015 ” ( DUBC) ; 1 female: “ Philippines / Negros / Mt. Canlaon / 600–950 m / July 2015 ” ( DUBC) ; 1 male: “ May 2018 / Don Salvador / Benedicto, Negros / Philippines ” ( JWc) ; 1 male with same label data (WJc); 1 female: “ April 2018 / Don Salvador / Benedicto, Negros / Philippines ” (WJc) ; 1 male: “ Philippines / Centra Visayas / Negros Oriental / Coll. DHEURLE // Dumaguete / vi–2014 / Lumawig leg. // P. hopkinsi / confluenteangu- / stata Horn, 1924 / DHEURLE det.” (CDc) ; 1 male: “ Philippines / Negros Occid. / Don Salvador / coll. DHEURLE // Benedicto / Avril 2018 / Lumawig leg. // P. hopkinsi / confluenteangu- / stata Horn, 1924 / DHEURLE det.” (CDc) . 1 male and 1 female: “ Philippines / Mindanao I. / Mt. Kitanglad / 28.VII.1990 // Coll. H. Sawada // Prothyma (Genoproth.) / banksi W. Horn, 1923 / Det. F. Cassola 1991” ( MCZR) ; 1 male “ Filippine Mindanao / Cabanglasan / Bukidnon VII.2014 ” (RSc) .

Diagnosis. Prothyma (Genoprothyma) confluenteangustata is similar to P.tenuipenis (both with evenly wrinkled pronotum and elongate apex of aedeagus), but is distinguished by: 1, its large size; 2, dorsally smooth protarsi of males; 3, unique endophallus structures.

Description. Size: TL (without labrum) 11.5–14 mm (mean = 11.8 mm, n = 2).

Head: Color shining black; with greenish or coppery reflections; orbital plates with reddish reflections; area behind the eye and near orbital plates with bluish reflections; genae blue violet; clypeus with greenish-blue reflections; surface glabrous, with two setigerous punctures next to eye, one along frontal margin and one dorsally at center; strong longitudinal striae on orbital plates and vertex; strong transversal striae behind the eyes and on neck; striae becoming obsolete on genae, frons, and clypeus. Head slightly smaller than elytra (mean EW/HW = 1.1). Labrum ochre-testaceous, margins slightly brownish, a slender blackish area basomedially. Labrum of male ( Fig. 5d ) wider than long (mean LW/LL = 1.3); with four setae located near the margin, two lateral and two apical; with two blunt anterolateral teeth and two anteriorly-directed thorn-like anterior teeth at the anteromedian lobe, space between slightly bulged. Labrum of female ( Fig. 5e ) slightly wider than long (mean LW/LL = 1.1), with four setae located near the margin, two lateral and two apical; with two blunt anterolateral teeth and two anteriorly-directed thorn-like anterior teeth at the anteromedian lobe, space between with a large triangular tooth. Mandibles dark brownish, with four teeth, small area of base testaceous. Labial and maxillary palpi testaceous, the last segment shiny black. Antennae slender, reaching posteriorly to the middle of the elytra in the male, shorter in the female; scape ochre-testaceous, in some specimens brownish dorsally; antennomere two, three and four brownish with metallic luster, two and three with lighter spot at apex; scape with one apical seta, the following three antennomeres nearly glabrous; antennomeres five to eleven brownish, dull, finely, and evenly pubescent.

Thorax: Pronotum slightly longer than wide (mean PW/PL = 0.9); sides not or barely rounded between the transversal constrictions, nearly parallel sided; glabrous, with transverse wrinkles on disc; black with greenish or reddish reflections dorsally; transversal constrictions with blue reflections; margins bluish; sternae and episternae glabrous, shiny black; mesepisternum of females with a shallow, short longitudinal furrow in upper part.

Elytra: ( Fig. 5b, c ) twice as long as wide (mean EL/EW = 2.0), parallel-sided, with a flattened basal hump, dorsal surface shining black, with greenish or coppery reflections; elytral testaceous maculation vary and consists of humeral, lateromedial and preapical dots; the humeral dot is located at the lateral edge and extends from there to the base or sometimes almost absent, in the lectotype it is connected with the lateromedial dot and the preapical dot; the lateromedial dot varies from triangular to narrow and oblique stripe; the preapical dot is rounded and has a small lateral line towards the apex in the lectotype; surface with dense and evenly spaced punctures, becoming confluent and forming grooves near the suture; these grooves covers about the apical third to half of the elytra, nearly reaching the tip of the lateromedial spot; apical margin distinctly rounded and restricted towards middle suture, with a tiny sutural spine. Epipleurae black.

Ventral aspect: Venter glabrous, blackish; trochanter glabrous, coxa, femur, tibia, and tarsi setose; trochanter, coxa, and basal half of femur variably testaceous, remainder of legs brownish, with or without metallic luster. Protarsomeres 1–3 in males laterally covered by sparse and medium length setae, ventrally covered by short white setae.

Aedeagus: ( Fig. 6 ) in lateral view, moderately slender in the basal quarter, then abruptly and voluminously expanded, then tapering again towards the apex, extending to elongate and slightly downturned apex (total length 2.9–3.1 mm).

Distribution. NEGROS—Cuernos Mt., Camlaon Mt., Don Salvador Benedicto, Dumaguete. MINDANAO:— Bukidnon: Cabanglasan, Kitanglad Mt.

Remarks. Based on its large size, dorsally smooth protarsi of males, and details of endophallus structure we here elevate this taxon to species status.

Figures

FIGURE 5. Prothyma (Genoprothyma) confluenteangustata Horn, 1924. (a) Dorsal habitus lectotype male. Scale = 5 mm. (b) left elytron, Lectotype male. (c) left elytron, female from Mindanao, Mt. Kitanglad. Scale = 5 mm. (d) Labrum, Lectotype male. (e) Labrum, female from Cuernos Mts. Scale = 1 mm.

FIGURE 6. Variability of aedeagus structure of Prothyma (Genoprothyma) confluenteangustata Horn, 1924. (a–b) Lectotype, Negros; (c–h) Negros, Mt. Canlaon; (i–j) Mindanao, Kitanglad. Scale = 1 mm.