Paranemastoma kalischevskyi (Roewer, 1951)

Modebadze, Naia, Martens, Jochen, Snegovaya, Nataly & Barjadze, Shalva, 2023, An annotated list of the Georgian harvestmen (Arachnida, Opiliones), Caucasiana 2, pp. 211-230 : 211

19. Paranemastoma kalischevskyi (Roewer, 1951)

Nemastoma kalischevskyi Roewer, 1951: 122 (plate 2, fig. 14, original description)

Nemastoma suchumium Roewer, 1951: 122 (plate 4, fig. 31, original description)

Nemastoma charitonovi Mkheidze, 1952a: 545-546 (fig. 1, original description)

Nemastoma charitonovi - Mkheidze, 1959: 111 (mention)

Nemastoma charitonovi - Mkheidze, 1964: 119 (fig. 2, mention)

Nemastoma kalischevskyi - Ljovushkin, 1966: 117 (mention; locality data)

Nemastoma kalischevskyi - Ljovushkin, 1972: 63 (mention)

Nemastoma kalischevskyi - Staręga, 1966: 390-391 (figs 3-4, redescription, new records)

Paranemastoma kalischevskyi - Staręga, 1978: 204 (mention)

Paranemastoma kalischevskyi - Martens, 2006: 196-199 (figs 28, 30a-d; 32, redescription, variation, new records, elevational records, relationships)

Paranemastoma kalischevskyi - Schönhofer, 2013: 41 (mention)

Paranemastoma kalischevskyi - Snegovaya, 2013: 185 (mention)

Paranemastoma kalischevskyi - Snegvaya, Pkhakadze and Inkirveli, 2014: 198 (mention; locality data from historical collection of the Georgian National Museum)

Type locality.

Georgia, Abkhazia, Sokhumi Municipality, village Lata.

Occurrence data in Georgia.

Abkhazia • Lata, Sokhumi Municipality; leg. Kalichevsky, September 1905 ( Roewer 1951). • Tsebelda, Sokhumi Municipality; leg. Y.V. Vornov, 9 September 1908 (Snegovaya et al. 2014). • Akhali Atoni, Sokhumi Municipality; leg. A. Riedel, 16 November 1958 ( Staręga 1966). • Mtsvane Kontskhi, Batumi Botanical garden, Kobuleti Municipality; leg. B. Pisarski, 10 August 1963 ( Staręga 1966). • Chamkhona cave, Arabika karst massif, Gagra Municipality; leg. S.I. Ljovushkin, 1963 (Ljovushkin 1966; Ljovushkin 1972). • Akhali Atoni, Gudauta Municipality; leg. S.I. Golovatch, 8 October 1979 ( Martens 2006). • Pskhu-Gumista Nature Reserve, Sokhumi Municipality; leg. J. Bohac, 6 June 1982 ( Martens 2006). • 1-3 km N of Duripsh, Gudauta Municipality; 300-400 m a.s.l.; leg. A. Ushakov, 9 September 1985 ( Martens 2006). • Ritsa Lake, Gudauta Municipality; leg. I.A. Ushakov, 11 September 1985 ( Martens 2006). • Ritsa Lake, Gudauta Municipality; 950-1000 m a.s.l.; leg. S.I. Golovatch, 13/14 August 1986; Fagus , Abies , Picea , Acer etc. forest, litter, under bark and stones ( Martens 2006). • Ritsa Lake, Gudauta Municipality; 1300-1450 m a.s.l.; leg. Golovatch, 14/16 August 1986; Fagus , Acer etc. forest on slope, under bark and stones ( Martens 2006). • Pskhu, Sokhumi Municipality; 700-950 m a.s.l.; leg. S.I. Golovatch, 15/16 August 1986; Fagus , Quercus , Castanea etc. forest, litter, under bark and stones ( Martens 2006). • Tsebelda, Gudauta Municipality; 300 m a.s.l.; leg. S.I. Golovatch, 19 August 1986; Carpinus , Acer and Buxus scrub, litter ( Martens 2006). • Sokhumi Botanical Garden; leg. S.I. Golovatch, 20 August 1986. • MFT Cave, Gudauta Municipality; leg. N. Myuge, 30 August 1986 ( Martens 2006). • Kvemo Tsumuri, Pskhu-Gumista State Reserve; leg. A. Ushakov, 25 September 1989 ( Martens 2006). Adjara • Zeraboseli, Kintrishi State Reserve, Kobuleti Municipality; 450-600 m a.s.l.; leg. S.I. Golovatch and J. Martens, 1 March 1981 ( Martens 2006). • Batumi Botanical Garden; 20-150 m a.s.l.; leg. S.I. Golovatch and J. Martens, 5 July 1981 ( Martens 2006). • Zeraboseli, Kintrishi State Reserve; 800 m a.s.l.; leg. S.I. Golovatch, 13 October 1981 ( Martens 2006). • Uchkho, Khulo Municipality; leg. T. Mkheidze, 5 March 1939 / 5 August 1939 (Mkheidze 1952; Mkheidze 1959; Mkheidze 1964) • 10 km E of Chakva, Chakvistavi, Kobuleti Municipality; leg. S.I. Kurbatov, 6 May 1987 ( Martens 2006). Samegrelo-Zemo Svaneti • Nenskra Canyon, Lukhi, Mestia Municipality; 800 m a.s.l.; leg. A. Ryvkin, 2 September 1986 ( Martens 2006). • Kherkhvashi, Mestia Municipality; 1250-1700 m a.s.l.; leg. S.I. Golovatch, 21 September 1986; Quercus , Fagus , Carpinus , Picea , Abies etc. forest, litter ( Martens 2006). • Mazeri, Mestia Municipality; 1660 m a.s.l.; leg. V. Brachat and H. Meybohm, 26 June 2017 (CJM). • Ushguli, Mestia Municipality; 2190 m a.s.l.; leg. V. Brachat and H. Meybohm, 30 June 2017 (CJM). • 25 km N Jvari, Tsalenjikha Municipality; 600 m a.s.l.; leg. J. and B. Martens, 20 July 2019; damp broad-leaved forest on a steep slope, ground moss covered (CJM). • Mestia; 1910-1650 m a.s.l.; leg. J. and B. Martens, 22 July 2019; mixed forest (CJM). • Latpari Pass, Mestia Municipality; 2100 m a.s.l.; leg. V. Brachat and H. Meybohm, 15 June 2021 (CJM). Guria • Vakijvari, Ozurgeti Municipality; 430 m a.s.l.; leg. V. Brachat and H. Meybohm, 12 June 2021 (CJM). Imereti • Kutaisi; leg. A. B. Shelkovnikov, 30 June 1911 (Snegovaya et al. 2014). • Bakhmaro, Chokhatauri Municipality; 1950-2020 m a.s.l.; leg. S.I. Golovatch and J. Martens, 8 June 1981 ( Martens 2006). • Baghdati; 290-630 m a.s.l.; leg. V. Brachat and H. Meybohm, 18 May 2018 (CJM). Racha-Lechkhumi and Kvemo Svaneti • Ghebi, Oni Municipality; 1310 m a.s.l.; leg. V. Brachat and H. Meybohm, 29 May 2018. • Shkedi, Lentekhi Municipality; 720 m a.s.l.; leg. J. and B. Martens, 20 July 2019 (CJM). Samtskhe-Javakheti • Kortaneti, Borjomi Municipality; leg. A. Riedel, 21 October 1958 ( Staręga 1966). • 8 km of Akhaldaba, Borjomi Municipality; 1000m a.s.l.; leg. S.I. Golovatch, 12 May 1983; Nedzura valley, Picea , Carpinus and Fagus forest, litter, logs ( Martens 2006). • Baniskhevi Valley, Borjomi Municipality; 800-900 m a.s.l.; leg. S.I. Golovatch, 16 May 1983; Picea , Fagus and Carpinus forest, litter, logs, under stones ( Martens 2006). • Timotesubani monastery, Borjomi Municipality; 1200 m a.s.l.; leg. J. and B. Martens, 15 July 2019; mixed forest along a stream (CJM). • Borjomi; 960 m a.s.l.; leg. J. and B. Martens, 17 July 2019; mixed broad-leaved forest (CJM). Shida Kartli • Surami, Khashuri Municipality; 1000 m a.s.l.; leg. S.I. Golovatch, 17 May 1983; Fagus , Alnus , Castanea , Rhododendron forest ( Martens 2006). • Holy Mountain above Kvishketi, Khashuri Municipality; 1200 m a.s.l.; 15 July 2019; leg. J. and B. Martens; Fagus orientalis forest (CJM). • Holy Mountain above Kvishketi, Khashuri Municipality; 1200 m a.s.l.; leg. J. and B. Martens, 16 July 2019; mixed forest, mainly Picea orientalis , on stream banks (CJM). Mtskheta-Mtianeti • Pasanauri, Dusheti Municipality; leg. K.A. Satunin, July 1913 (Snegovaya et al. 2014). • Natakhtash (?) Pass; leg. A.B. Shelkovnikov, 14 July 1917 (Snegovaya et al. 2014). • Kazbegi; 1900 m a.s.l.; leg. M.S. Khokia, 17 July 1980; pitfall traps ( Martens 2006). • Magalakhari Pass; 1200 m a.s.l.; leg. S.I. Golovatch and K. Eskov, 6 May 1987 ( Martens 2006). Kakheti • Batsaro State Reserve, Akhmeta Municipality; 800-850 m a.s.l.; leg. S.I. Golovatch and K. Eskov, 5/6 May 1987; Fagus and Castanea forest, litter ( Martens 2006). • Gombori Pass, Sagarejo Municipality; 1500-1600 m a.s.l.; leg. J. and B. Martens, 28 July 2019; bushes along a stream (CJM). • Birkiani, Akhmeta Municipality; 930-1110 m a.s.l.; leg. J. and B. Martens, 31 July 2019; side valley, river banks, mixed broad-leaved forest (CJM).

Global distribution.

Endemic to the Caucasian ecoregion. Records in Azerbaijan, Turkey, Georgia and Russian north-western Caucasus ( Staręga 1978; Martens 2006; Schönhofer 2013; Snegovaya 2013; Kurt 2014).

Remarks.

This is a widely distributed species occurring in a large variety of forest habitats and even in open patches far from forests; its elevational distribution presently known extends from 290 m a.s.l. to 2190 m a.s.l. (CJM, Martens 2006). In terms of morphology, this is a rather plastic species regarding the apophysis of the first member of the male chelicera and the armament of the dorsal scutum of the body (images in Martens 2006). Genetic analysis is needed to detect possible cryptic species.