Pupina lowi Morgan, 1885

Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak, 2022, Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia, ZooKeys 1119, pp. 1-115 : 1

Pupina lowi Morgan, 1885

Figs 37K , 40E

Pupina lowi Morgan, 1885: 414, pl. 7, fig. 3a-d. Type locality: Lahat, Kinta [Perak State, Malaysia]. von Möllendorff 1891: 345. Sykes 1903: 197, Gunong Inas, Perak. van Benthem Jutting 1949: 57, Larut Mills, Perak. van Benthem Jutting 1960: 13, limestone hill Kaki Bukit, near kampong Wang Tangga, Perlis [Malaysia]. Maassen 2001: 41.

Pupina (Tylotoechus) lowi - Kobelt 1902: 317. Laidlaw 1928: 34.

Pupina artata [non Benson]- Berry 1963: pl. 6, fig. 36. Foon et al. 2017: 40, fig. 15b, Ipoh, Perak.

Type material examined.

Syntype MNHN-IM-2000-35846 (1 shell; Figs 37K , 40E ) from Lahat, Perak.

Diagnosis.

Shell globose; last whorl ca. three quarters of shell height. Apertural lip slightly thickened but not expanded. Parietal tooth sharp, tooth-like, thickened; columellar tooth fin-shaped, thickened, located next to slit-like anterior canal. Posterior canal gradually widening like keyhole.

Differential diagnosis.

Pupina lowi is most similar to P. tchehelensis and P. brachysoma in having a sharp, tooth-like, thickened parietal tooth, a fin-shaped, thickened columellar tooth that is located next to a slit-like anterior canal, and a posterior canal that is gradually widening. However, P. lowi is different from P. tchehelensis by having a more globose shell shape, and different from P. brachysoma in that an apertural lip is not expanded.

Distribution.

Perak and Perlis States, Malaysia ( Maassen 2001).

Remarks.

By comparing with the type specimen, the specimens of P. artata figured in Berry (1963: pl. 6, fig. 36) and Foon et al. (2017: fig. 15b) from Bat Cave Hill Plot 2, Ipoh, Perak should belong to P. lowi (Foon, pers. comm.). The specimen of P. lowi figured in BEDO (2017: 91) should constitute a different species as it is different from the syntype figured here in having a smaller, sharper parietal tooth revealing a wide posterior canal and an earlobe-shaped columellar tooth covering the anterior canal. Thus, that specimen should belong to the P. arula species group instead (see above).

Figures

Figure 37. Shells of Pupina aureola species group from mainland Southeast Asia. A-C Pupina tchehelensis: specimens A SMF 109947 / 6 B CUMZ 12135, and C CUMZ 12136 D, E Pupina latisulci sp. nov. D holotype CUMZ 12146 / 1 and E paratype CUMZ 12146 / 2 F-I Pupina paviei F paralectotype MNHN-IM- 2000 - 35837 G paralectotype RBINS 525404, and specimens H CUMZ 12134 and I NHMUK ex. Dautzenberg coll. J Pupina tongupensis, syntype NHMUK 1906.4.4.38 K Pupina lowi, syntype MNHN-IM- 2000 - 35846 L Pupina anceyi, syntype MNHN-IM- 2000 - 35833 M Pupina laffonti, syntype MNHN-IM- 2000 - 9656 N Pupina solidula, lectotype SMF 109915 / 1 O Pupina douvillei, holotype MNHN-IM- 2000 - 35532. Photo: M. Caballer, P. Maestrati, MNHN (F, K-O), F. Trus, RBINS (G), J. Ablett, H. Taylor, NHM (J).

Figure 40. A, B Pupina latisulci sp. nov. A holotype CUMZ 12146 / 1 and B paratype CUMZ 12146 / 2 from Khao Ok Talu, Phatthalung C, D Pupina stoliczkai sp. nov. C holotype CUMZ 12147 / 1 and D paratype NHMUK 20210336 from Wat Ratburana School, Chumpon E Pupina lowi, syntype MNHN-IM- 2000 - 35846 from Lahat, Perak F Pupina dorri dorri, paralectotype MNHN-IM- 2000 - 35836 from Haiphong, Vietnam. Photo: P. Maestrati, MNHN (E, F).