Paraleucopis corvina Malloch,

Wheeler, Terry A. & Sinclair, Bradley J., 2019, Systematics of Paraleucopis Malloch with proposal of Paraleucopidae, a new family of acalyptrate Diptera, Zootaxa 4668 (3), pp. 301-328: 314-316

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Paraleucopis corvina Malloch


Paraleucopis corvina Malloch 

( Figs 10View FIGURES 8–10, 37–39View FIGURES 37–42, 44View FIGURES 43–44)

Paraleucopis corvina Malloch, 1913: 149  . Type locality: Kerr Ranch, New Mexico.

P. corvina: Wheeler, 1959: 154  [biology]; Steyskal, 1971: 1 [redescription, terminalia figured]; Steyskal, 1981: 405 [key]; Smith, 1981: 411 [biology].

Type material examined. HOLOTYPE Ƌ, labelled: “Hunter/ No 1891”; “Kerr Ranch/ 4-23-[19] 10 N. Mex.”; “On/ Yucca sp.”; “JD Mitchell/ Collector”; “Type/ No. 15755/ U.S. N.M. [red label]”; “ Holotype / Paraleucopis  / corvina Malloch  [red label]”  . PARATYPES: Same data as holotype (4 Ƌ, 4 ♀, USNM)  .

Additional material examined. MEXICO: 28 mi SW Las Arrastras , B.C. [Baja California], 29.v.1962, R.A.E. Ryckman, C.P. Christianson (1 ♀, USNM); Baja N., 9 km NW Rancho Santa Inés, 6.iv.1985, 29°46′N 114°46′W, 550 m, Clark & Blom (3 ♀, LACM)GoogleMaps  . USA. Arizona: Graham Mtns, Marijilda Wash , 4025 ft, 32°42.6′N 109°46.5′WGoogleMaps  , 28–29.v.2013, Malaise, J.E. O’Hara (1 Ƌ, CNC); S.W. Research Station , 5 mi SW Portal, 5400 ft  , 23.v.–, C.W. Sabrosky (1 Ƌ, CNC). California : Blythe  , 1.v.1955, W.R. Richards (1 ♀, CNC); San Bernardino Co., Mitch- ells Caverns   ,, T. Griswold (1 ♀, LACM). New Mexico: Lincoln Co., Jicarilla Mtns, nr. Jicarilla, T 5N, R12E, 23  , 27.v.1991, sweeping Gramma grass, evening, J. Gelhaus, C. Nelson #508 (1 ♀, ANSP); Valencia Co., Grants , 10 mi E  ,, G.E. Bohart (2 Ƌ, LACM). Texas : Balley Co  ., 29.iv.1959, M.R. Wheeler; ex. Great Horned Owl’s nest (1 ♀, AMNH); Sanderson   , 27.iv.1959, Becker & Howden (1 ♀, CNC). Washington : Chelan Co., Lake Wenatchee  , 4.viii.1951 (2 Ƌ, 1 ♀, AMNH)  .

Recognition. This species is characterized by fronto-orbital setae not distinct; antenna mostly yellow, postpedicel darker dorsoapically; face with pruinescence restricted to antennal foveae; parafacial shiny; fore coxa and all femora dark; hind femur without anteroventral preapical setae; surstylus triangular, strongly tapered to narrow apex.

Description. Frons dark to reddish-brown band above antennae; gena dark. Face mostly shiny, with pruinescence restricted to antennal fovea; parafacial shiny. Fronto-orbital setae not distinct. Antenna mostly yellow, with dorsoapical margin of postpedicel darkened; antennal bases separated by slightly more than length of pedicel. Clypeus shiny.

Katepisternum with several stout ventral setae, anterior to mid coxa; short setulae on anterior half. Scutum shiny with dense setulae. Coxae and femora dark, with apex of femora pale; fore tibia and apical half of mid tibia pale; hind tibia mostly dark; tarsomeres on all legs pale; mid tibia with dark ventroapical seta, less than one-quarter length of tarsomere 1.

Male postabdomen ( Fig. 37View FIGURES 37–42): epandrium rounded, evenly sclerotized, setose. Hypandrium broad, with pair of short anterior setae; with rounded postgonite apically bearing strong marginal setae. Surstylus long, strongly tapered to narrow apex; apex arch posteromedially; setae confined mostly to posterior margin. Distiphallus tapered to narrow apex. Cercus elongate, longer than dorsal length of epandrium; apex rounded.

Female postabdomen ( Fig. 10View FIGURES 8–10): tergite 6 ( Fig. 38View FIGURES 37–42) with broadly bilobed anteriorly margin, slightly narrowed medially with sinuous posterior margin; posterior corners slightly prolonged with pair of long setae, more than half length of tergite; spermatheca ( Fig. 39View FIGURES 37–42) with long pigmented denticles or papillae; without end apparatus.

Distribution. This is the most widely distributed species, ranging as far north as Washington, south to Texas and Baja California ( Fig. 44View FIGURES 43–44).

Remarks. Several specimens have been taken from birds’ nests ( Malloch 1913; Wheeler 1959). Malloch (1913) included 14 specimens in the type series, but only nine were identified. The whereabouts of the remaining five specimens remains unknown.


Smithsonian Institution, National Museum of Natural History


Natural History Museum of Los Angeles County


Canadian National Collection of Insects, Arachnids, and Nematodes


Academy of Natural Sciences of Philadelphia


American Museum of Natural History














Paraleucopis corvina Malloch

Wheeler, Terry A. & Sinclair, Bradley J. 2019

P. corvina:

Steyskal, G. C. 1981: 405
Smith, R. L. 1981: 411
Steyskal, G. C. 1971: 1
Wheeler, M. R. 1959: 154

Paraleucopis corvina

Malloch, J. R. 1913: 149