Paraleucopis boydensis Steyskal,

Wheeler, Terry A. & Sinclair, Bradley J., 2019, Systematics of Paraleucopis Malloch with proposal of Paraleucopidae, a new family of acalyptrate Diptera, Zootaxa 4668 (3), pp. 301-328: 314

publication ID

publication LSID

persistent identifier

treatment provided by


scientific name

Paraleucopis boydensis Steyskal


Paraleucopis boydensis Steyskal 

( Figs 13View FIGURES 11–14, 33–36View FIGURES 30–36, 43View FIGURES 43–44)

Paraleucopis boydensis Steyskal, 1971: 2  .

P. boydensis: Steyskal, 1981: 405  [key]; Smith, 1981: 411 [biology].

Type material examined. HOLOTYPE Ƌ, labelled: P.L. Boyd Des[ert]. Res[earch]. Center/ 3.5mi.S.Palm Desert, Riv[erside]/ Co.Calif./ 26 APR.1970 Suzy &/ Saul Frommer”; “UCR.ENT.MUS./ Specimen #/ 24871”; “ HOLO- TYPE / PARALEUCOPIS  / BOYDENSIS/ Steyskal [red label]” ( USNM). ALLOTYPE: Same data as holotype, UCR. ENT. MUS., specimen #22082 (1 ♀, USNM).

Additional material examined. USA. California: Andrews Canyon, Palm Springs , 10.iv.1955, W.R. Richards (2 ♀, CNC)  ; Borrego , 3.iv.1960, A.L. Melander (2 ♀, USNM)  ; Imperial Co., Fish Creek Mts , 300 ft, 20.iv.1955, W.R.M. Mason (1 ♀, CNC)  ; Palm Desert, 2100 ft, 4.iv.1955, W.R.M. Mason (1 Ƌ, CNC; 1 Ƌ DEBU); Riverside Co., Sec. 19; T. 6S; R.6E, elev. Ca. 2800 ft, NE of Black Hills, 9.v.1973, S. Frommer (5 Ƌ, 3 ♀, CNC)  ; San Ber- nardino Co., Mitchells Caverns ,, T. Griswold (1 ♀, LACM)  .

Recognition. This species is characterized by fronto-orbital setae not distinct; antenna yellow; face with pruinescence on upper half; parafacial with pruinescence; clypeus shiny; gena mostly shiny, pruinosity not extended posteroventrally below eye; fore coxa and all femora dark; hind femur without anteroventral preapical setae; surstylus L-shaped, with broad rectangular base bearing several strong setae.

Description. Frons dark to reddish-brown band above antennae; gena dark. Face with pruinescence on upper half ( Fig. 13View FIGURES 11–14); parafacial pruinose. Fronto-orbital setae not distinct. Antenna mostly yellow, with apical half of postpedicel darkened; antennal bases separated by more than length of pedicel. Clypeus shiny.

Katepisternum with several stiff, dark ventral setae, anterior to mid coxa; short setulae on anterior half. Scutum shiny with dense setulae. Coxae and femora dark, with apex of femora pale; fore tibia mostly pale; mid and hind tibiae darkened in middle, pale basally and apically; tarsomeres on all legs pale, except darkened tarsomere 5 dorsally; mid tibia with dark ventroapical seta, less than one-quarter length of tarsomere 1.

Male postabdomen ( Figs 33, 34View FIGURES 30–36): epandrium rounded, evenly sclerotized, setose. Hypandrium broad, with pair of long anteriorly setae; with narrow and rounded postgonite apically. Surstylus L-shaped, with broad rectangular base bearing several strong marginal setae; apex slightly curved medially. Distiphallus tapered to narrow apex. Cercus rounded, width subequal to length, shorter than dorsal length of epandrium.

Female postabdomen: tergite 6 ( Fig. 35View FIGURES 30–36) bilobed anteriorly, slightly narrowed medially with slightly sinuous to straight posterior margin; with 2 pairs of long setae, longer than half length of tergite; spermatheca ( Fig. 36View FIGURES 30–36) with long pigmented denticles or papillae; without end apparatus.

Distribution. This species is known only from California, USA ( Fig. 43View FIGURES 43–44).


Smithsonian Institution, National Museum of Natural History


Canadian National Collection of Insects, Arachnids, and Nematodes


Ontario Insect Collection, University of Guelph


Natural History Museum of Los Angeles County














Paraleucopis boydensis Steyskal

Wheeler, Terry A. & Sinclair, Bradley J. 2019

P. boydensis: Steyskal, 1981: 405

Steyskal, G. C. 1981: 405
Smith, R. L. 1981: 411

Paraleucopis boydensis Steyskal, 1971: 2

Steyskal, G. C. 1971: 2