Paratelius Kazantsev, 1992

Malohlava, Vladimír & Bocak, Ladislav, 2009, A revision of Paratelius Kazantsev, 1992 (Coleoptera: Lycidae), Zootaxa 2306 (1), pp. 44-50 : 45

Genus Paratelius Kazantsev, 1992

Diagnosis: Paratelius resembles Scarelus in general appearance and has similar long setae between dense, short pubescence on elytra. Paratelius , like Atelius , bears four longitudinal costae on each elytron. In all known species the phallus is very long, slender and slightly curved ( Figs 1A–1J ).

Redescription: Similar to Scarelus in general appearance. Small-bodied, slender, 5.8–7.0 mm long. Most species brown to reddish brown colored, some uniformly testaceous; when dark colored, humeral part of elytra lighter and apical half of elytra dark. Head small, free, hypognathous, covered with short pubescence, surface mat, with fine structure. Frons convex; eyes small; antennal tubercles inconspicuous, divided by shallow depression; antennal cavities very close each to other, separated by narrow cranial bridge. Antennae 11-segmented, compressed, slightly serrate, long, surpassing elytral apex by 3 to 5 antennomeres, covered with dense, recumbent pubescence; apical antennomere longest, narrowed to the apex, pointed. Mandibles stout, quite long, slightly curved, shining, with smooth internal margin. Other mouthparts tiny. Labium and maxillae clearly visible without dissecting, reduced. Labium reaching from gular margin of mouth cavity to one third of its diameter: maxillae reaching to a half of diameter. Labial palpi 2-segmented, maxillary palpi 4- segmented, apical palpomere stoutest in both palpi. Labrum projected forwards, triangular. Pronotum trapezoidal, flat, posterior margin wider than frontal one, frontal margin slightly rounded, lateral margins convex, posterior angles acutely projected, posterior margin almost straight. Pronotal disc with single median longitudinal carina with very deep and narrow depression in middle part. Scutellum flat, trapezoidal, shiny, sparsely pubescent, usually emarginate at apex. Elytra flat, with four longitudinal costae. Costae connected by regular transverse ridges, whole elytra densely and shortly pubescent, also bearing sparse, stout, long, erect setae. Legs slender, compressed, densely pubescent. Tarsus with 5 tarsomeres. Male genitalia trilobate, uniform in general shape ( Figs 1 A-J). Phallus very long, widest at base, paramerae very short, globular in shape, phallobase longer than paramerae.

Distribution: Malaysia: Sabah State. Distribution is limited to mountain ranges in northeastern Borneo. These mountains were produced by a counterclockwise rotation of Borneo since the Late Miocene ( Hall & Blundell, 1996). Based on the endemic occurence in a single mountain range, we may suppose speciation of all Paratelius species occurred in this region. Paratelius has not been recently collected in the lowlands of Borneo, and only the type specimen of P. diversicornis is recorded from Brunei without precise locality data. Although Borneo was connected during the last glacial maximum to Indochina ( Voris, 2000) and the generic structure of both faunas is similar, neither Paratelius from Borneo nor related Atelius from Vietnam expanded their ranges over the subaerial shelf of the Asian continent.

Figures

FIGURES 1A–J. Male genitalia in the ventral and lateral views. A, B Paratelius snizeki n. sp.; C, D P. emasensis n. sp.; E, F P. nigricornis n. sp.; G, H P. diversicornis (Pic).; I, J. P. juvencus (Kleine). Scales 0.5 mm.