Xylocopa (Neoxylocopa) fimbriata Fabricius, 1804

Mérida-Rivas, Jorge A., Hinojosa-Díaz, Ismael A., Ayala-Barajas, Ricardo, Pozo, Carmen & Vandame, Rémy, 2022, Revision of carpenter bees of the subgenus Neoxylocopa Michener (Hymenoptera: Apidae) from Mexico and Mesoamerica, Zootaxa 5158 (1), pp. 1-67 : 7-16

publication ID

https://doi.org/10.11646/zootaxa.5158.1.1

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lsid:zoobank.org:pub:2191B88A-2CEB-496B-A255-5800EEFAEAFA

DOI

https://doi.org/10.5281/zenodo.6821864

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https://treatment.plazi.org/id/FE1487AC-AD0C-FFA9-FF42-9043FD4EF82D

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scientific name

Xylocopa (Neoxylocopa) fimbriata Fabricius, 1804
status

 

Xylocopa (Neoxylocopa) fimbriata Fabricius, 1804

( Figs. 1–10 View FIGURES 1–5 View FIGURES 6–10 , 102–104 View FIGURES 102–110 , 132 View FIGURES 132–135 )

Xylocopa fimbriata Fabricius, 1804: 340 . Holotype: University of Copenhagen Zoological Museum, Copenhagen, Denmark (♀). Type locality: ‘America meridionalis ’ [according to Moure (1960: 144).

Xylocopa corniger Westwood, 1840: 270 . Holotype: Oxford University Museum Oxford. England (♀). Type locality: ‘un- known’, synonymy by Smith (1874:284).

Xylocopa cajennae Lepeletier, 1841: 203 . Syntype: Oxford University Museum Oxford. England (♂). Type locality: French Guiana, Cayenne, synonymy by Smith (1874:284).

Xylocopa cornuta Lepeletier, 1841: 176 . Syntype: Istituto di Zoologia dell’Universita di Torino, Torino, Italy (♀). Type locality: French Guiana, Cayenne, synonymy by Erichson (1848:591).

Xylocopa virescens Lepeletier, 1841: 186 . Holotype: Istituto di Zoologia dell’Universita di Torino, Torino. Italy (♀). Type locality: French Guiana, Cayenne, synonymy by Moure (1960:144).

Xylocopa fimbriata var. motaguensis Cockerell, 1912 ; 556. Holotype: American Museum of Natural History , New York, USA (♀). Type locality: Guatemala, Gualán.

Diagnosis. Body length, female, 28.74 mm (27.62–30.60; n=5), male, 29.43 mm (28.45–30.97; n=5). Female: Head with elevated prominences adjacent to ocelli with appearance of horns ( Fig. 1 View FIGURES 1–5 ); middle ocellus clearly below level of lateral ocelli and upper margin of eyes; area between parapsidal line and mesoscutal disc with sparse punctation (punctures separated by three or more puncture diameters); wings brown to dark brown with blue-violet iridescence ( Fig. 3 View FIGURES 1–5 ); pubescence evenly black; integument of terga evenly black; T1 to T4 with scattered minute punctures; T2 to T4 with minute hairs, T5 hairs 0.49–1.02 (length respect MOD). Male: Terga yellowish-orange, with no bands ( Fig. 10 View FIGURES 6–10 ); metasoma with yellowish orange pubescence; posterior tibia internal distal extreme with a notable sharp projection but with apex rounded, anterior margin with a rounded prominence, width of projection two thirds the width of the tibia ( Fig. 9 View FIGURES 6–10 ); length of hairs on terga respect MOD, T2 0.71–1.59, T3 0.77–1.28, T4 0.79–1.42, T5 0.81–1.54; tip of the lateral process of the gonostylus ending broad, rhomboid shaped ( Fig. 103 View FIGURES 102–110 ).

Description. Female: Integument and pubescence black; wings brown to dark brown with blue-violet iridescence ( Fig. 3 View FIGURES 1–5 ); venation dark brown. Body length 28.74 mm (27.62–30.60; n=5); length of forewing 25.27 mm (24.63–26.29; n=5); head length 7.18 mm (6.78–7.58; n=5); intertegular distance 8.37 mm (7.66–8.78; n=5); width of T2 12.48 mm (11.69–12.82; n=5). Head: Frontal area with two large prominent elevations similar to horns that arise between the lateral ocelli and middle ocellus; middle ocellus below upper level of eyes ( Fig. 1 View FIGURES 1–5 ). Very wide inter-ocular space (wider than compound eye length). Clypeus with a large depression toward the middle of antertior submargin, with large dense punctures separated by less than their diameter, diminishing toward the inferolateral areas; inferior margin of the clypeus with punctures on its entire width, forming a margin with a width equal to the diameter of a puncture. Lateral ocelli located above tangent of eyes. Mesosoma: Mesoscutal disc unpunctate, area between parapsidal line and disc with some sparse punctation (punctures separated by three or more puncture diameters). Metasoma: T1 to T3 with scattered minute punctures. Length of hairs on terga respect MOD as follows: T2 to T4 with minute hairs, T5 0.49–1.02. Male: Pubescence and integument yellowish orange. Hyaline wings, venation reddish-brown. Body length 29.43 mm (28.45–30.97; n=5); length of forewing 23.75 mm (22.71–24.79; n=5); head length 4.98 mm (4.83–5.18; n=5); intertegular distance 8.47 mm (7.83–8.95; n=5); width of T2 13.74 mm (13.17–14.20; n=5). Head: Labrum with a triangular shaped area without punctures on the middle region; rest of labrum with scattered punctures. Clypeus with very small, scattered punctures on marginal, submarginal, and inferolateral regions; subapical region without punctures (some specimens with some very scattered punctures); clypeus disc and superior region with dense punctures, separated by three or more times their diameter. Lateral ocelli located on the same level as tangent of eyes. Mesosoma: Between the parapsidal line and discal area with dense punctures, separated by one to one and a half times their diameter. Posterior tibia: anterior margin with long light brown pubescence; posterior margin with long and short pubescence; internal surface with more abundant pubescence toward the basal area and short dense pubescence on the distal extreme; internal distal extreme with a notable sharp projection but with apex rounded; anterior margin with a rounded prominence; width of projection two thirds the width of the tibia ( Fig. 9 View FIGURES 6–10 ). Metasoma: T1 with dense, long, plumose hairs, T2 to T5 with plumose hairs restricted to submarginal area, mixed with simple hairs, T6 and T7 with a mixture of dense, long, simple and plumose hairs. Length of hairs on terga respect MOD as follows: T2 0.71–1.59, T3 0.77–1.28, T4 0.79–1.42, T5 0.81–1.54. Genitalia: Distal angle of the ventral section of the gonostylus, acute ( Fig. 104 View FIGURES 102–110 ); basal angle at the base of lateral process of the gonostylus, strong, very noticeable ( Fig. 103 View FIGURES 102–110 ); tip of the lateral process ending broad, rhomboid shaped ( Fig. 103 View FIGURES 102–110 ).

Variation. The elevations adjacent to the ocelli present variation in size among specimens collected in a single or different locality on the same date. Throughout its distribution, wings may have bluish iridiscence with violet or only bluish iridescence, with the papillate area with greenish iridescence. These variations have led to the fact that throughout history, different species have been described, which have been placed in synonymy. Metasoma: Some males have dark brown terga, perhaps due preservation issues in collections.

Distribution ( Fig. 132 View FIGURES 132–135 ). Present in Barbados, Brazil, Colombia, Costa Rica, Ecuador, El Salvador, French Guiana, Guadalupe, Guatemala, Guyana, Honduras, Martinique, Mexico, Nicaragua, Panama, Saint Kitts and Nevis Islands, Trinidad and Tobago, and Venezuela ( Gonzalez & Gonzalez 2009; Moure 2012; Mawdsley 2018). It has also been reported for Bolivia and Peru by Moure (2012), however according to Hurd (1978) the presence of this species in those countries is seemingly an error, and none of the supporting references giving distributional records in Moure’s compilation are posterior to Hurd’s assertion. Widely distributed in Mexico and Central America, in the ecoregions ( Dinerstein et al. 2017): Dry forests of Balsas, Dry forests of Central America, Pine-oak forests of Central America, Dry forests of the Central Depression of Chiapas, Seasonal humid forests of Costa Rica, Humid isthmus-Atlantic forests, Humid isthmus-Pacific forests, Dry forests of Jalisco, Mesoamerican mangroves of the Gulf and Caribbean, Mangrove of the Mesoamerican Pacific north, Dry forest of Panama, Humid forests of PetenVeracruz, Sierra of the Tuxtlas-Veracruz, Humid forests of the Sierra Madre of Chiapas, Pine-oak forests of the Sierra Madre of Oaxaca, Pine-oak forests of the Sierra Madre del Sur, Pine-oak forests of the Sierra Madre Occidental, Dry forests of Sinaloa, Mangroves of the South Pacific of Mesoamerica, Dry forests of the South Pacific, Mountainous forests of Talamanca, Pine-oak forests of the Trans-Mexican Volcanic Belt, Dry forests of Veracruz, and Humid forests of Veracruz. Broad altitudinal distribution, 0 to 2620 m.

Material examined. 968 ♀ 56 ♂: COSTA RICA : 1 ♀ ( MZUCR). Alajuela: Los Chiles: Caño Negro , 10.89381, -84.78884; 35 m, 14-XI-1992 GoogleMaps , K. Martínez, 1 ♀ ( MNCR). Naranjo: San Jerónimo de Naranjo , 10.1, -84.36; 1036 m, 15-IV-1965 , J. L. Corrales, 1 ♀ ( MZUCR). Cartago: Tapanti National Park, Biological Station , 9.761389, - 83.78472; 1200 m, 31-X-2001 GoogleMaps , R. Brooks , 1 ♀ ( SEMC). Guanacaste: Bagaces: Cañas , 14 km S, EJN, 10.30, - 85.09560; 36 m, 8-III-1988 , F. D. Parker, 1 ♀ ( BBSL). Entrada a Palo Verde , 10.38334, -85.31668; 15 m, 13-XI- 2001 GoogleMaps , A. Soto, 1 ♀, 17-XI-2001, F. Umaña, 1 ♀, I. Jiménez , 1 ♀ ( MNCR). Finca Montezuma , 3 km SE Río Naranjo, 10.66409, -85.09720; 452 m, 7-II-1992 GoogleMaps , F. D. Parker, 1 ♀; - IV-1992, F. D. Parker, 1 ♀ ( BBSL). Hacienda Palo Verde , 2 km E, 10.34990, -85.33169; 42 m, I-1974 GoogleMaps , K. H. Keeler, 1 ♀ ( BBSL). Palo Verde , 10.38333, - 85.31667; 10 m, - III-1988 GoogleMaps , A. Bernecker, 1 ♀; R. Lucking , 1 ♀ ( MZUCR). Palo Verde National Park , 10.31460, - 85.25320, 6 m, 17-II-1991 GoogleMaps , F. D. Parker, 1 ♀ ( BBSL). Parque Nacional Palo Verde, Estación Palo Verde , 10.34912, -85.35235; 5 m, 1-X-1990 GoogleMaps , U. Chavarria, 2 ♀ 1 ♂; 1-XI-1990, D. Acevedo, 1 ♂; 1-I-1991, D. Acevedo, 4 ♀; 10-I- 1991, J. Corrales, 1 ♀; 1-III-1991, D. Acevedo, 1 ♀; 1-IV-1991, D. Acevedo, 1 ♀; 1-VI-1991, D. Acevedo, 1 ♀; 25-III-1992, A. Gutiérrez, 3 ♀; M. Ortíz , 2 ♀; 1-IX-1993, U. Chavarria, 1 ♂; 5-V-1999, I. Jiménez, 1 ♀; 6-IX-1999, I. Jiménez, 1 ♀; J. Weng , 1 ♀; 17-XI-2004, Y. Cárdenas, 1 ♀ ( MNCR). Parque Nacional Palo Verde, Sector Palo Verde , 10.36667, -85.38326; 107 m, 2-XI-1999 GoogleMaps , I. Jiménez, 1 ♀ ( MNCR). R. Naranjo , 3 km SE, 10.66409, - 85.09720; 451 m, 25-I-1992 GoogleMaps , F. D. Parker, 1 ♂ ( BBSL). La Cruz: Estación Pitilla , 9 km S Santa Cecilia, 10.9926, -85.42947; 673 m, 1-IX-1989 GoogleMaps , C. Moraga, 1 ♀ ( MNCR). La Garita, Est. Los Almendros, 11.03385, -85.52479; 300 m, 28-III-1992 GoogleMaps , M. Reyes, 3 ♀; G. Gallardo , 2 ♀; 23-X-1992, E. López, 1 ♀ 1 ♂ ( MNCR). Mirador casa 1, 11.03846, -85.72889; 81, 9-IX-2007 GoogleMaps , M. Zumbado, 2 ♀ ( MNCR). Parque Nacional Guanacaste, Casa Oeste , Cerro El Hacha, 12 km SE La Cruz, 11.00568, -85.56495; 300 m, 1-I-1988 GoogleMaps , M. Espinoza, 8 ♀ ( MNCR). Parque Nacional Guanacaste, Cerro el Hacha, 10.89985, -85.57733; 400 m, 11-I-1988 GoogleMaps , A. Solís, 3 ♀; 4-II-1988, A. Solís, 1 ♀ ( MNCR). Parque Nacional Guanacaste, Estación Biológica Maritza , 10.96254, -85.49524; 600 m, 1-V-1989 GoogleMaps , 1 ♀; 1-V-1990, R. Blanco , 1 ♀; 1-II-1992, F. Araya, 1 ♀; 27-II-1992, A. Gutiérrez, 1 ♀ ( MNCR). Santa Elena, Cerro El Hacha, 12 km SE de La Cruz, 10.98316, -85.54106; 300 m, 1-I-1988 GoogleMaps , A. Solis, 6 ♀; 1-IX-1991, E. López, 1 ♀; 1-XI-1991, E. López, 2 ♂ ( MNCR). Santa Elena, Finca Jenny, 10.86550, -85.57353; 200m, 1-IX-1988 GoogleMaps , 2 ♀; 1-X-1988, 7 ♀; 1-XI- 1988, 14 ♀ 3 ♂; 1-XII-1988, 5 ♀; 1-III-1989, 1 ♀ 1 ♂; 16-X-1991, E. Araya, 1 ♀; 1-1-1992, E. Araya, 1 ♀; 19-I- 1993, E. Araya, 1 ♀; 5-VIII-1993, E. Araya, 1 ♀; 5-IX-1993, E. Araya, 3 ♀; 10-X-1993, E. Araya, 3 ♀ ( MNCR). Santa Elena, P.N. Santa Rosa, Estación Murciélago, 8 km SW de Cuajiniquil, 10.90193, -85.73101; 100 m, 1-II- 1989 GoogleMaps , 1 ♀; 10-X-1993, C. Cano, 1 ♀; F. Quesada , 1 ♀; 3-XII-1993, C. Cano, 1 ♀; 7-IV-1994, C. Cano, 1 ♀; 4-VIII- 1994, F. Quesada, 1 ♀ ( MNCR). Santa Rosa, Parque Nacional, 10.81666, -85.61666; 279 m, 11-II-1979 GoogleMaps , C. Starr, 1 ♀; 11-III-1986, W. Wcislo, 1 ♀ ( SEMC). Liberia : 16.4 km NE de Liberia, 10.72750, -85.39361; 360 m, 22-XI- 2001 GoogleMaps , R. Brooks , 1 ♀ ( SEMC). Curubandé , 10.71666, -85.38333; 360 m, 22-XI-2001 GoogleMaps , C. Godoy, 1 ♀; J. Weng , 1 ♀ ( MNCR). Liberia , 10.63564, -85.44495; 139 m, 1-IX-1964 GoogleMaps , Robert & Roberts, 1 ♀ ( SEMC). Nacascolo, Parque Nacional Santa Rosa, Estación Santa Rosa, 10.83641, -85.61549; 300 m, 1-I-1989 GoogleMaps , 5 ♀; 1-I-1991, 2 ♀; 1-XI-1991, O. Espinoza, 1 ♀; 1-II-1992, O. Espinoza, 3 ♀; D. Brenes , 1 ♀; G. Pereira , 1 ♀ 1 ♂; 1-VII-1992, G. Pereira, 1 ♂ ( MNCR). Sector Las Pailas , 4.5 km SW del Volcán Rincón de la Vieja, 10.77678, -85.35191, 800 m, 22-XI-1992 GoogleMaps , G. Rodríguez, 1 ♀, 6-I-1993, J. Sihezar, 1 ♀ ( MNCR). Nicoya: San Antonio, 3 km NO de Nacaome, Parque Nacional Barra Honda , 10.16821, -85.37365; 100 m, M. Reyes: 13-X-1992 GoogleMaps , 1 ♂; 13-XI-1992, 2 ♀; 1-XII-1992, 2 ♀; 1-I-1993, 1 ♀; 1-XI-1994, 1 ♂ ( MNCR). Tilarán: Santa Rosa Biological Station , 10.83641, -85.61549; 300 m, 10- III-1986 GoogleMaps , W. Wcislo, 2 ♀ ( SEMC). Heredia: Belén: La Ribera: San Antonio , 9.98905, -84.18284; 950 m, 1-VIII- 1991 GoogleMaps , M. A. Zumbado, 1 ♀ ( MNCR). Santo Domingo: Santo Domingo , 9.9992, -84.06239; 1243 m, 12-V-2017 GoogleMaps , I. Aguilar, 1 ♀ ( CINAT). Hotel Bougainvillea , 9.974999, -84.07666; 1135 m, 14-V-1999 GoogleMaps , D. Brzoska, 2 ♀ ( SEMC). Santa Rosa, 9.973625, -84.09439; 1100 m, 9-IX-2002 GoogleMaps , M. Solís, 1 ♀ ( MNCR). Limón: Talamanca: Amubri , cacao farm, 19-I-1979 , C. Starr, 1 ♀ ( SEMC). Puntarenas: Garabito: Reserva Biológica Carara, Estación Quebrada Bonita , 9.77423, -84.60812; 27 m, 1-III-1995 GoogleMaps , R. Guzmán , 1 ♀ ( MNCR). Isla San Lucas: Puesto Isla San Lucas , 9.94652, -84.90556; 0 m, 7-III-2005 GoogleMaps , J. Azofeita, 3 ♀; 14-IX-2005, J. Azofeita, 1 ♀ ( MNCR). Monteverde : 10.3069, -84.8097; 1430 m, 23-III-2018 GoogleMaps , 1 ♀ ( CINAT). Paquera: Isla Pan de Azúcar , 9.926427, -84.9041; 0 m, 29-XI-2005 GoogleMaps , J. Azofeita, 1 ♀ ( MNCR). San José: Acosta: San Ignacio de Acosta , 4 km E, 9.79145, -84.19698; 1220 m, 8-VII- 1963 GoogleMaps , C. D. Michener, 1 ♀ ( SEMC). Aserrí: Aserrí Jocotal , 9.77687, -84.13753; 850 m, - V-2006 GoogleMaps , J. Mora, 1 ♀ ( MZUCR). Escazú : 9.91670, -84.13330; 1100 m, F. D. Parker: 4-III-1988 GoogleMaps , 1 ♀; 22-IV-1989, 1 ♀ ( BBSL). San Antonio de Escazú , 9.65444, -84.15444; 1350 m, 26-II-1985 GoogleMaps , W. Wcislo, 1 ♀ ( SEMC). Montes de Oca: Lourdes , 9.93206, -84.04119; 1230 m, 14-IV-1975 GoogleMaps , R. Ochoa , 1 ♂ ( MZUCR). San José, Sabanilla , 9.94515, -84.03109; 1282 m, - IV-2014 GoogleMaps , S. Alvarado, 1 ♀ ( MZUCR). Mora: Ciudad Colón , 10.30, -84.416700; 800 m, - XII-1989 , L. Fourniev, P. Hanson, 1 ♀ ( BBSL). Z.P. El Rodeo, Upaz, Mora , 9.90933, -84.27888; 984 m, 22-II-2018 GoogleMaps , M. Fernández, Gliricidia sepium , 1 ♀ ( MZUCR). Puriscal: Mercedes Sur, Fila Altos de Carara , 9.77522, -84.47642; 1116 m, 6-II-1993 GoogleMaps , J. C. Saborio, 1 ♀ ( MNCR). Santiago de Puriscal , 9.88595, -84.34916, 700 m, - II-2016 GoogleMaps , M. Fernández, 1 ♀ ( MZUCR). San José : 9.91189, -84.07362; 1140 m, 24-VIII-1962 GoogleMaps , C. D. Michener, 1 ♀ ( SEMC). San José , 9.93835, -84.0898; 1137 m, - II-1988 GoogleMaps , A. Bernecker, 1 ♀ ( MZUCR). Santa Ana: 9.9336, -84.1841; 902 m, 27-III-2012 GoogleMaps , A. Chassoul, 1 ♀ ( CINAT). EL SALVADOR: La Libertad: Quetzaltepeque : 13.82634, -89.27047; 450 m, 13-VII- 1969 GoogleMaps , J. R. Quezada , 1 ♀ ( EBCH). Quezaltepeque , 3 mi S, 13.78198, -89.27908; 861 m, 19-VII-1961 GoogleMaps , M. Irwin, 1 ♂ ( SEMC). Quezaltepeque , 3 mi W, 13.83498, -89.31937; 496 m, 1-VIII-1961 GoogleMaps , M. E. Irwin, 1 ♂ ( BBSL). GUATE- MALA: Huehuetenango: Jacaltenango: Rancho Viejo , 15.62299, -91.78; 949 m, 7-V-2010 , 2 ♀ ( ECOAB). La Democracia: Cajuil , 15.63746, -91.78722, 1026 m, 8-V-2010 GoogleMaps , 1 ♀, 18-XI-2010, M. Hurtado, 1 ♀ ( ECOAB). Chaparral , 15.67407, -91.95515; 1007 m, J. Mérida: 20-II-2010 GoogleMaps , 1 ♀; 28-IV-2010, 1 ♀; 8-V-2010, 1 ♀ ( ECOAB). Nojoya , 15.66123, -91.82008; 1000 m, 29-IV-2010 GoogleMaps , P. Sagot, 9:30 h, 1 ♀ ( ECOAB). Pinalito , 15.6653, -91.8618; 963 m, 2-III-2010 GoogleMaps , C. Balboa, 10:50 h, 1 ♀; 17-XI-2010, O. Argüello, 1 ♀; 17-11-2010, M. Hurtado, 1 ♀ ( ECOAB). La Libertad: Camojaito , 15.61604, -91.87273; 940, 1-III-2010 GoogleMaps , D. Sánchez, 9:55 h, 1 ♀ ( ECOAB). San Antonio Huista: Miralvalle , 15.66789, -91.95520; 1127 m, 8-V-2010 GoogleMaps , 2 ♂ ( ECOAB). Chejbal , 15.68596, -91.78158, 1085 m, 23-II-2010 GoogleMaps , C. Balboa, 13:00 h, 1 ♂; M. Cigarroa , 12:00 h , 1 ♂. MEXICO : 5 ♀ 1 ♂ ( CNIN), 3-VIII-1937 , C.C. Hoffmann, 1 ♂ ( CNIN), 5-VII-1949 , J. Julia, 1 ♀ ( CNIN). Chiapas : 2 ♀ 2 ♂ ( ECOAB). Acacoyagua : 10 de Abril 15.44472, -92.69966; 596 m, 3-IX-2004 GoogleMaps , R. Ayala , 1 ♀ ( ECOAB). Las Golondrinas , 15.43271, -92.65260; 862 m, 8-XII-2006 GoogleMaps , C. Balboa, 1 ♀; 18-XII-2006, C. Balboa, 1 ♀; 20-V-2007, R. Vandame , 1 ♀ ( ECOAB). Nueva Reforma , 15.435, -92.7509; 508 m, 4-IX-2004 GoogleMaps , M. Rincón, 1 ♀ ( ECOAB). Rosario Zacatonales , 15.43365, -92.65051; 915 m, 31-V-2005 GoogleMaps , M. Guzmán, 1 ♀; 4-VIII-2005, M. Guzmán, 2 ♀; 10-XI-2006, C. Balboa, 1 ♀; 24-IV-2007, C. Balboa, 1 ♀; 10-V-2007, M. Guzmán, 1 ♀ ( ECOAB). Ángel Albino Corzo: Jerusalén , 15.77486, -92.64197; 885 m, 3-XI- 2017 GoogleMaps , P. Sagot, 1 ♀ ( ECOAB). Querétaro, 15.8344, -92.76275; 678 m, 2-III-2016 GoogleMaps , P. Sagot, 1 ♀ ( ECOAB). Arriaga: Carretera Tuxtla-Arriaga , 16.32649, -93.87328; 426 m, 10:45 h, P. Sagot GoogleMaps , 1 ♀ ( ECOAB). Ejido Adolfo López Mateos , 16.32131, -93.98319; 368 m, 20-I-2009 GoogleMaps , M. Guzmán, 7 ♀; 23-I-2009, C. Balboa, 3 ♀; 10-III-2009, C. Balboa, 3 ♀; 10-III-2009, M. Guzmán, 2 ♀; 13-III-2009, J. Florez, 1 ♀; 17-XI-2009, C. Balboa, 1 ♀; 20-XI-2009, C. Balboa, 4 ♀ 2 ♂ ( ECOAB). Paraje Poza Verde , 16.34207, -93.9544; 466 m, 24-I-2009 GoogleMaps , M. Guzmán, 1 ♀; 18-XI- 2009, C. Balboa, 4 ♀ ( ECOAB). Cacahoatán: 4 km al norte carr. a Unión Juárez , 15.035095, -92.151412, - 730 m, 6-III-2020 GoogleMaps , P. Sagot, 1 ♀ ( ECOAB). Chiapa de Corzo: Autopista a Chiapa de Corzo , 16.748, -92.974; 575 m, 28- XII-2011 GoogleMaps , 11:30 h, P. Sagot, 1 ♀ 1 ♂ ( ECOAB). Autopista a San Cristóbal de Las Casas , 16.73596, -92.95005; 730 m, 11:30 h, P. Sagot GoogleMaps 2 ♂ ( ECOAB). Carretera 28, Sur de la Providencia, 16.46988, -92.95717; 500 m, 9:15 h, P. Sagot GoogleMaps , 2 ♀ ( ECOAB). Cintalapa: Cintalapa , 16.68596, -93.72855; 582 m, - XI-1948 GoogleMaps , 3 ♀ ( CNIN). Comitán de Domínguez: Francisco Sarabia , 16.17231, -92.11812; 1572 m, 4-XII-2019 GoogleMaps , P. Sagot, 3 ♀ ( ECOAB). Frontera Comalapa: Rodulfo Figueroa , 15.851745, -92.069833; 595 m, 31-III-2016 GoogleMaps , 8:20 h, P. Sagot, 1 ♀. Huehuetán : 15.01789, -92.3939; 86 m, 14-V-1987 GoogleMaps , J. Mendoza, 1 ♀; L. Mendoza , 2 ♀; 14-IX-1987, J. Mendoza, 1 ♀ ( ECOAB). Ixtapa: Multajo, 2 km E, Carr 190, 16.71654, -92.85908; 1502 m, 15:00 h, P. Sagot GoogleMaps , 1 ♀. La Concordia: Carretera 157, 7 km sur de Benito Juárez, 15.97922. -92.81434; 581 m, 2-III-2016 , 13:20 h, P. Sagot, 1 ♀ ( ECOAB). Carretera 157, El Espinal, 15.94570, -92.77746; 613 m, 2-III-2016 GoogleMaps , 12:44 h, P. Sagot, 2 ♀ ( ECOAB). Carretera 157, Norte del Embarcadero , 16.03700, -92.55436, 587 m, 1-IV-2016 GoogleMaps , 8:35 h, P. Sagot, 1 ♀ ( ECOAB). Carretera 157, Oeste Niños Héroes , 16.116839, -92.653494; 616 m, 11:40 h, 1-IV-2016 GoogleMaps , P. Sagot, 2 ♀ ( ECOAB). La Tigrilla, 15.97605, - 92.81658; 586 m, 4-III-2020 GoogleMaps , E. Gómez, 10:35 h, 2 ♀ ( ECOAB). Niños Héroes , 16.11359, -92.62986; 606 m, 7:50 h, P. Sagot GoogleMaps , 1 ♀ ( ECOAB). La Trinitaria: San Francisco de Asís , 16.0118, -92.0267; 767 m, 16-XI-2011 GoogleMaps , 7:40, J. Mérida, 2 ♀; P. Sagot 1 ♀ ( ECOAB). Mapastepec: Camino a Santa Rosa, 15.4696, -92.7784; 233 m, 26-XI-2004 GoogleMaps , M. Guzmán, 1 ♀; 24-II-2005, M. Rincón, 1 ♀ ( ECOAB). La Palma, 15.53032, -92.80199; 498 m, 17-V-2007 GoogleMaps , C. Balboa, 1 ♀; M. Guzmán : 18-III-2005 , 4 ♀; 19-III-2005, 2 ♀; 22-VI-2005, 1 ♀; 23-VI-2005, 1 ♀; C. Balboa : 6-XII- 2005 , 3 ♀; 17-V-2007, 2 ♀ ( ECOAB). Unión Los Olivos , 15.52120, -92.80114; 316 m, J. Esponda: 5-5-2005 GoogleMaps , 1 ♀; 2-III-2006, 1 ♀; 29-IX-2006, M. Cigarroa, 3 ♀; 1-X-2006, J. Mérida, 1 ♀; 10-X-2006, J. Mérida, 4 ♀; C. Balboa : 10-X-2006 , 1 ♀; 18-X-2006 ,, 2 ♀; 19-X-2006, 1 ♀; 23-X-2006, M. Cigarroa, 3 ♀; 28-X-2006, J. Mérida, 6 ♀; 8- XI-2006, M. Guzmán, 4 ♀; 22-XI-2006, J. Esponda, 1 ♀; 27-XI-2006, C. Balboa, 2 ♀; 18-IV-2007, M. Guzmán, 2 ♀ ( ECOAB). Unión Los Olivos-orilla del río, 15.5219, -92.79967; 325 m, M. Guzmán: 23-10-2004 GoogleMaps , 1 ♀; 28-X- 2004, 1 ♀; 8-XI-2004, 1 ♀; 20-XI-2004, J. Esponda, 3 ♀; M. Guzmán : 28-XI-2004 , 11 ♀; 2-II-2005, 6 ♀; 2-III- 2005, 1 ♀; 26-V-2005, 1 ♀; 28-V-2005, 1 ♀; 1-VI-2005, 3 ♀; 27-VI-2005, J. Esponda, 1 ♀ ( ECOAB). Mazatán: Mazatán , 14.85849, -92.45159; 22 m, 25-V-1989 GoogleMaps , G. López, 1 ♀; 20-VII-1989, A. Aquino, 1 ♀ ( ECOAB). Motozintla: Belisario Domínguez , 15.3041, -92.3811; 791 m, 25-V-1998 GoogleMaps , D. Pérez, 1 ♀ ( ECOAB). Finca Belen , 15.21944, -92.34916; 1337 m GoogleMaps , 1 ♀ ( ECOAB). Salanueva , 15.31660, -92.34023; 1000 m, 3-II-2019 GoogleMaps , 11:20 h, P. Sagot, 1 ♀ ( ECOAB). Ocozocoautla: 8.1 km N Nuevo México S de Ocozocoautla , 16.69761, -93.41156; 773 m, 11-IV-1993 GoogleMaps , C. Mendoza, Lonchocarpus sp , 1 ♀ ( EBCH). 11.7 km S Ocozocoautla , 16.6688, 93.41589; 750 m, 11-IV-1993 GoogleMaps , A. Rodríguez, 1 ♀ ( EBCH). 15 km N Nuevo México S de Ocozocoautla , 16.59037, -93.43355, 777 m, 11-IV-1993 GoogleMaps , E. Tovar, Piscidia piscipula , 1 ♀; R. Ayala , Piscidia piscipula , 2 ♀ ( EBCH). Pijijiapan: Carr. 200, 15.71307, - 93.23767; 78 m, 14:00 h, P. Sagot GoogleMaps 1 ♀; I. Pérez , 1 ♀ ( ECOAB). Carr. 200, 15.70490, -93.22095; 58 m, 9-III-2020 GoogleMaps , E. Gómez, 2 ♀; P. Sagot , 7:40 h , 3 ♀ ( ECOAB). Joachim Amaro , 15.77766, -93.40933; 51 m, 1-II-2019 GoogleMaps , 9:30 h, G. Mera, 1 ♀; I. Pérez , 1 ♀ ( ECOAB). Solo Dios , 15.66695, -93.42535; 9 m, 11:25 h, P. Sagot GoogleMaps 1 ♀ ( ECOAB). Socoltenango : 16.22243, -92.35469; 741 m, 15-I-2017 GoogleMaps , 7:30 h, P. Sagot, 1 ♀ ( ECOAB). Tapachula : 36.7 km N Tapachula, 15.103, -92.3097; 440 m, 17-IV-1993 GoogleMaps , E. Tovar, Crotalaria acapulcensis , 5 ♀ ( EBCH). Joaquín Miguel Gutiérrez , 14.74328, -92.31274; 28 m, 8-VIII-2014 GoogleMaps , J. Ruíz et al, 2 ♀; 9-I-2015, J. Ruíz et al. 4 ♀ ( ECOAB). Joaquín Miguel Gutiérrez , 14.75482, -92.29233; 33 m, 7-VIII-2014 GoogleMaps , M. Guzmán et al, 1 ♀ ( ECOAB). Joaquín Miguel Gutiérrez , 14.76733, -92.29931; 38 m, 7-VIII-2014 GoogleMaps , M. Guzmán et al, 1 ♀ ( ECOAB). Rancho Santa Teresita , 14.9048, -92.3144; 102 m, 10-VIII-1987 GoogleMaps , L. Mendoza, 1 ♀ ( ECOAB). Tapachula , 14.88669, -92.28660; 127 m, 14-IX-1988 GoogleMaps , A. Aquino, 1 ♀; 19-VI-2005, A. Niño, 1 ♀ ( ECOAB). Tonala: Ejido Nueva Costa Rica , 16.1352, - 93.6824; 408 m, 19-I-2009 GoogleMaps , C. Balboa, 11 ♀; 22-I-2009, C. Balboa, 3 ♀; M. Guzmán , 1 ♀; P. Sagot , 1 ♀; 9-III-2009, M. Guzmán, 4 ♀; 9-III-2009, C. Balboa, 4 ♀; J. Mérida , 1 ♀; 12-III-2009, C. Balboa, 2 ♀; 21-V-2009, C. Balboa, 2 ♀; M. Guzmán , 1 ♀; 17-IX-2009, M. Guzmán, 1 ♀; 16-XI-2009, C. Balboa, 4 ♀ 2 ♂; 19-XI-2009, M. Guzmán, 4 ♀ ( ECOAB). Puerto Arista , 15.94282, -93.82458; 6 m GoogleMaps . 31-I-2019, 10:45 h, G. Mera, 2 ♀; P. Sagot , 2 ♀ ( ECOAB). Tuxtla Chico: 1a Sección de Medio Monte , 14.89049, -92.2094; 180 m, 8-X-2015 GoogleMaps , I. Hinojosa, 1 ♀ ( CNIN). Casa Jorge Mérida , 14.98054, -92.17292; 15-III- GoogleMaps 2008, 453 m, J. Mérida, 1 ♀ ( ECOAB). Rosario Izapa , 14.9733, -92.1546; 445 m, 24-VII-1987 GoogleMaps , R. Santos , 2 ♀; 25-VII-1987, R. Santos , 1 ♀; 8-X-1987, L. Godínez, 2 ♀; L. Mendoza , 2 ♀; R. Santos , 1 ♀; 13-V-2005, C. Pérez, 1 ♀ ( ECOAB). Tuzantán: Carr. Huixtla-Motozintla , 15.1628, -92.4329; 110 m, 3-2-2019 GoogleMaps , 8:45 h, P. Sagot, 5 ♀; I. Pérez , 3 ♀ 1 ♂ ( ECOAB). Unión Juárez: Talquián Viejo , 15.0881, -92.0916; 1748 m, 4-VI-2005 GoogleMaps , H. Colin 1 ♀ ( ECOAB). Unión Juárez , 15.0618, -92.08; 1310 m, 29-XII-19896, F. Pérez , 2 ♀ ( ECOAB). Venustiano Carranza: Carretera Pujiltic-Venustiano Carranza , 16.30446, -92.53471; 637 m, 14:00 h, 2-I-2017 GoogleMaps , P. Sagot, 1 ♀ ( ECOAB). Carretera V. Carranza-Acala , 16.401466, -92.730145; 532 m, 21-XII-2015 GoogleMaps , 12:40 h, P. Sagot. 1 ♀ ( ECOAB). Venustiano Carranza , 16.3166, -92.5666; 621 m, 21-IV-1993 GoogleMaps , C. Mendoza, 1 ♀; E. Tovar , 1 ♀; J. L. Neff , 1 ♀ ( EBCH). Villa Corzo: Carretera a Ejido Sierra Morena , 16.16865, -93.50988; 796 m, 24- XII-2016 GoogleMaps , P. Sagot, 1 ♀; R. Vandame , 1 ♀ ( ECOAB). El Parral, camino Oeste a Villaflores , 16.36631, -93.06750; 981 m, 20-X-2017 GoogleMaps , P. Sagot, 1 ♀ ( ECOAB). Villa Corzo , El Parral, carr. 157, 16.36959, -92.97258; 643 m, 4-I-2016 GoogleMaps , P. Sagot, 1 ♀ ( ECOAB). San Pedro Buena Vista, Belisario Domínguez , 16.0572, -93.1137; 823 m, 4-I-2016 GoogleMaps , 9:40 h, P. Sagot, 1 ♀ ( ECOAB). Villafores : 5 km W Domingo Chanona, 16.3167, -93.3332; 670 m, 11-IV-1993 GoogleMaps , F. A. Noguera, Wisteria sinensis , 1 ♀ ( EBCH). Carretera 230, 16.41139, -93.68287; 644 m, 30-I-2019 GoogleMaps , 12:20 h, G. Mera, 2 ♀; P. Sagot , 1 ♀ ( ECOAB). Carretera 230, 16.34958, -93.42621; 716 m, 30-I-2019 GoogleMaps , 9:15 h, P. Sagot, 3 ♀; I. Pérez , 1 ♀ ( ECOAB). Carretera 230, 16.35493, -93.52333; 693 m, 30-I-2019 GoogleMaps , 10:35 h, P. Sagot, 1 ♀ ( ECOAB). San Clemente , 16.4182, -93.71358; 608 m, 30-1-2019 GoogleMaps , 12:50 h, I. Pérez, 4 ♀ ( ECOAB). Colima: - VIII-1916 , C. C. Hoffmann, 1 ♀ ( CNIN). Armería: Highway 200- Armeria R., 18.93103, -103.96384; 20 m, 22-II-1992 GoogleMaps , D. Brzoska, 5 ♀ ( SEMC). Colima, 19.23879, -103.75923; 457 m, 9-II-1954 GoogleMaps , R. Dreisbach , 1 ♀, 9-XI-1954, R. Dreisbach , 1 ♀ ( SEMC). Comala: Glory Desert , 19.34572, -103.79819; 594 m, 15-II-2012 GoogleMaps , 10:00 h, E. Esquivel, 1 ♂; J. Mérida , 1 ♀ ( ECOAB). Hacienda San Antonio , 19.45192, -103.71394; 1190 m, 26-X-2015 GoogleMaps , R. Ayala , 2 ♀ ( EBCH). Manzanillo: Playa de Oro , 19.11991, -104.36848; 6 m, 7-I-1986 GoogleMaps , D. Zehr, 1 ♀ ( SEMC). Tecomán: Pascuales Malecón , 18.8494, -103.9536; 16 m, 3-X-2010 GoogleMaps , R. Ayala , 2 ♀ ( EBCH). Guerrero: L. Ancona , 2 ♀ ( CNIN). Acapulco : 16.87111, -99.89777; 87 m, 6-VIII-1954 GoogleMaps , Univ. of Kansas Mex. Exped . 5 ♀ ( SEMC). Atoyac de Álvarez : El Faisanal, 17.43333, -100.20000; 1200 m, 3-VIII-1983 GoogleMaps , R. Ayala , 1 ♀ ( MZFC). El Rincón de las Parotas , 17.24516, - 100.39135; 429 m, 8-VII-1985 GoogleMaps , I. Hinojosa, 1 ♀; G. Rodríguez , 1 ♀; 10-IX-1985, I. Hinojosa, 3 ♀; O. Morales , 3 ♀; 2-XI-1985, A. Morales, 1 ♀ ( MZFC). Puente de los Lugardo , 17.33833, -100.25500; 925 m, 30-III-1984 GoogleMaps , R. Ayala , 1 ♀ ( EBCH), 30-III-1984 , R. Ayala , 7 ♀ 2 ♂ ( MZFC), 1-XI-1985 , I. Hinojosa, 1 ♀ ( MZFC). Puente del Rey 9 km al sur de El Paraíso, 17.34907, -100.26303; 872 m, 10-IX-1985 GoogleMaps , V. Velarde , 1 ♀ ( MZFC). Río Santiago , 17.25333, -100.31833; 698 m, 2-V-1985 GoogleMaps , O. Morales, 1 ♀; 3-V-1985, O. Morales, 1 ♀; M. A. Luis , 1 ♂; 5-VII-1985, I. Hinojosa, 1 ♀; 8-VII-1985, I. Hinojosa, 1 ♀; M. A. Luis , 1 ♀; 9-VII-1985, I. Hinojosa, 1 ♀; M. A. Luis , 1 ♀, 5- VIII-1985, M. A. Luis, 1 ♀; 9-IX-1985, I. Hinojosa, 1 ♀; O. Morales , ♀; 9-XI-1985, G. Rodríguez, 1 ♀ ( MZFC). Azoyu: Carr. Cuanacaxtitlan a Azoyu , 16.77561, -98.59564; 318 m, 29-XII-2019 GoogleMaps , E. Caballero, 1 ♀ ( ECOAB). Carr. Cuanacaxtitlan a Azoyu , 16.75822, -98.60384; 340 m, 29-XII-2019 GoogleMaps , E. Caballero, 2 ♀, E. Gómez , 1 ♀ (ECO- AB). Chilpancingo de los Bravo: Chilpancingo , 2 mi E; 17.48477, -99.44462; 1243 m, 7-VIII-1962 GoogleMaps , R. Roberts , 1 ♀ ( SEMC). Copala: Carr. entre Car. 200 y Playa Ventura, 16.59204, -98.92507; 15 m, 25-XII-2019 GoogleMaps , E. Caballero, 1 ♀ ( ECOAB). Malinaltepec: Cocoyul , 17.02440, -98.77848; 1090 m, 26-XII-2019 GoogleMaps , E. Caballero, 1 ♀; P. Sagot , 1 ♀ ( ECOAB). Ometepec: Huajintepec , 16.61298, -98.24503; 343 m, 24-XII-2019 GoogleMaps , E. Gómez, 1 ♀ ( ECOAB). Pilcaya: Grutas , 2 km SE, hwy 55, 18.67007, -99.51193; 1163 m, 6-I-1989 GoogleMaps , R. Minckley, B. Danforth , 1 ♀ ( SEMC). Pilcaya, Taxco , 23 mi N, 18.71100, -99.67296; 1341 m, 8-VIII-1954 GoogleMaps , Univ. of Kansas Mex. Exped . 2 ♀ ( SEMC). San Luis Acatlán: Yoloxóchitl , 16.81583, -98.69027; 600 m, 1-I-2012 GoogleMaps , Amith & García, 1 ♀; 8-XI-2012, Amith & Bautista, 1 ♂ ( SEMC). Yoloxóchitl , 16.80523, -98.66190; 661 m, 29-XII-2019 GoogleMaps , P. Sagot, 1 ♀ ( ECOAB). San Marcos: Acapulco , 21 mi E, 16.77559, -99.53882; 92 m, 3-VIII-1963 GoogleMaps , M. Naumann, 6 ♀ ( SEMC). Taxco de Alarcón: Taxco , 20 mi S, 18.43419, -99.55960; 1051 m, 6-II-1954 GoogleMaps , R. Dreisbach , 1 ♀ ( SEMC). Taxco , 4 mi S, 18.49939, - 99.59972; 1326 m, 3-VIII-1951 GoogleMaps , Univ. of Kansas Mex. Exped . 1 ♀ ( SEMC). Técpan de Galeana : La Ola, 17.65103, -100.87428; 1859 m, 20-XI-2012 GoogleMaps , 8:45 h, E. Esquivel, 1 ♀ ( ECOAB). Rodesia , 17.20696, -100.69623; 24 m, 18- VII-1971 GoogleMaps , 1 ♀ ( CNIN). Técpan , 17.22670, -100.62684; 37 m, 18-7-1971 GoogleMaps , 1 ♀ ( CNIN). Jalisco: Cabo Corrientes: El Tuito , 20.31970, -105.3265; 604 m, F. D. Parker, T. L. Griswold GoogleMaps , 2 ♀ ( BBSL). El Tuito , 20.32405, -105.31912; 630 m, 22-X-1977 GoogleMaps , C. Beutelspacher, 1 ♀ ( CNIN). El Tuito , 20.30275, -105.34759; 692 m, 5-IX-2018 GoogleMaps , 12:30 h, J. Mérida, 2 ♀ ( ECOAB). Punta Tehualmixtle , 20.23896, -105.57492; 35 m, 24-XI-1991 GoogleMaps , J. Villa, E. Ramírez, 2 ♀ ( EBCH). Cihuatlán: Barra de Navidad , 19.21594, -104.68423; 14 m, 10-IV-1986 GoogleMaps , RAyala, 1 ♀ ( EBCH). Hostotipaquillo: Plan de Barrancas , 3 km SE, 21.05300, -104.19600; 766 m, 8-VII-1963 GoogleMaps , F. D. Parker, L. A. Stange, 1 ♀ ( BBSL). La Huerta: Chamela , 19.53000, -105.08166; 5 m, 5-VII-1984 GoogleMaps , R. Ayala , 3 ♀ ( MZFC), 26-IX-1985 , F. D. Parker, T. L. Griswold , 2 ♀ 1 ♂ ( BBSL). Estación de Biología Chamela , 19.49948, -105.04242; 70 m, 25-V-1980 GoogleMaps , S. H. Bullock, 1 ♀ ( CNIN), 21-23-VII-1980 , A.Pescador, 1 ♀ ( CNIN), 26-VII-1980 , A.Pescador, 1 ♀ ( CNIN), S. H. Bullock : 1-VIII-1981 , 1 ♀ ( EBCH), 4-VIII-1981 , 2 ♀ ( EBCH), 11-IX-1981 , 1 ♀ ( EBCH), 30-IX-1981 , A.Pescador, 1 ♀ ( CNIN), S. H. Bullock : 19-IX -1982 , 1 ♂ ( EBCH), 19-IX-1982 , 1 ♀ ( CNIN), 9-XI-1982 , 1 ♂ ( SEMC), 13-I- 1983 , 1 ♀ ( EBCH), 30-I-1983 , 1 ♀ ( EBCH), 20-II-1984 , 1 ♀ ( EBCH), 19-V-1984 , 1 ♀ ( EBCH), R. Ayala : 22-III- 1985 , 1 ♀ ( SEMC), 16-V-1985 , 2 ♀ ( EBCH), 1-VII-1985 , F. D. Parker, T. L. Griswold , 2 ♀ ( EBCH), 17-VII-1985 , R. Ayala , 2 ♀ ( EBCH), 25-IX-1985 , C. D. Michener, 1 ♀ ( EBCH), 26-IX-1985 , Radclife B. Roberts, 1 ♀; F. D. Parker, T. L. Griswold , 1 ♀ 1 ♂ ( EBCH), 13-X-1985 , M. Sánchez, 1 ♂ ( EBCH), 28-I-1986 , R. Ayala , 1 ♀ ( SEMC), 29-VI-1986 , R. A. Usela , 1 ♀ ( EBCH), R. Ayala : 5-VII-1986 , 1 ♀ ( EBCH), 5-IX-1987 , 1 ♂ ( EBCH), 4-XI-1987 , T. L. Griswold , 1 ♀ ( BBSL), 21-V-1988 , E.Ordway, Solanum ochraxceun-ferrugineum , 1 ♀ ( EBCH), 9-VI-1988 , R. Minckley , 1 ♀ ( SEMC), 2-IX-1988 , A. Rodríguez, 1 ♀ ( EBCH), 12-VII-1989 , C. D. Michener, 1 ♀ ( SEMC), 14-X- 1990 , G. Ortega, C. Mayorga, J. Martínez, S. Rodríguez, 1 ♀ ( CNIN), - I-1991 , C. D. Michener, 1 ♀ ( SEMC), 19- VIII-1995 , R. Ayala , 1 ♀ ( EBCH), 5-12-VII-2013 , S. Zaragoza, 1 ♀ ( CNIN), 1-7-I-2014 , S. Zaragoza, 1 ♀ ( CNIN), 7-IX-2018 , S. Kilpatrick, 2 ♀; 15-VI-2019, J. Mérida, 1 ♀ ( ECOAB). Playa Careyes, Chamela , 19.44282, - 105.02328; 16 m, C. D. Michener: 30-IX-1985 GoogleMaps , 1 ♀ ( SEMC), 30-IX-1985 , 1 ♀ ( EBCH). Playa Careyitos, 19.43766, -105.02605; 2 m, 10-IX-1985 GoogleMaps , R. Ayala , 2 ♀; 27-IX-1985, R. B. Roberts , 7 ♀ ( EBCH). Río Cuixmala , 19.41055, - 104.95638; 46 m, 27-II-1991 GoogleMaps , E. Ramírez, 2 ♀ ( EBCH). Mixtlán: Guadalajara , 65 mi W, 20.47589, -104.37009; 1685 m, 3-IX-1971 GoogleMaps , 1 ♂ ( BBSL). Pihuamo: Pihuamo , 19.25376, -103.36904; 665 m, 9-II-1954 GoogleMaps , R. Dreisbach , 1 ♀ ( SEMC). Tequila: Tequila , 3 mi NW, 20.91882, -103.86330; 1206 m, 19-VII-1953 GoogleMaps , Univ. of Kansas Mex. Exped . 1 ♀ ( SEMC). Volcán de Tequila , 20.82757, -103.85104; 1800 m, 12-IV-1995 GoogleMaps , H. Fierros, 1 ♀ ( EBCH). Volcán de Tequila , 20.86341, -103.84396; 1300, 15-VII-1995 GoogleMaps , 1 ♀; 7-XI-1995, 1 ♀ ( EBCH). Tomatlán: Carr. Barra de Navidad-Pto Vallarta (La Virgencita), 19.84972, -105.31777; 26 m, 18-IX-1985 GoogleMaps , R. A. Usela , 1 ♀ ( EBCH). Mexico: Camino a Ixtapantogo, San Nicolás , 12-I-1952 , 3 ♀ ( CNIN). Ixtapan de la Sal: San Pedro Tlacochaca , 18.85753, -99.64716; 1800 m, 10-XI-2015 GoogleMaps , 8:30, P. Sagot, 1 ♀ ( ECOAB). Ixtapan de La Sal , 18.84375, -99.67552; 1900 m, 30-X-1991 GoogleMaps , R. Ayala , Salvia sp , 1 ♀ ( EBCH). Michoaacán: Ario : El Cangrejo, 20 km N La Huacana, 19.04166, - 101.81138; 1090 m, 30-X-1987 GoogleMaps , 1 ♀ ( EBCH). Arteaga: Carr. Arteaga 2 km, 18.34803, -102.29194; 830 m, 18-VII- 1989 GoogleMaps , V. R. Castrejón , 1 ♀ ( EBCH). El Zorrillo , 18.28172, -102.271311; 758 m, 13-X-1994 GoogleMaps , 1 ♀ ( MZFC). Buenavista, 10 km W Apatzingán, La Huina , 19.10813, -102.52083; 256 m, 17-VII-1989 GoogleMaps , V. R. Castrejón , Chilopsis linearis , 1 ♀ ( EBCH). Coahuayana: Palos Marías , 18.81154, -103.54045; 240 m, 29-V-1988 GoogleMaps , L. Cervantes, 1 ♀ ( CNIN). Gabriel Zamora : 20 km S Uruapán, 19.25057, -102.05532; 944 m, 7-II-1990 GoogleMaps , S. H. Bullock, 1 ♀ ( EBCH). La Huacana : 5.8 km N de la Huacana, 19.00811, -101.82282; 766 m, 18-VII-1989 GoogleMaps , Brooks & Roig-Alsina, 1 ♀ ( SEMC). 17 km N La Huacana , 19.03256, -101.81486; 1010 m, 4-XI-1992 GoogleMaps , A. Rodríguez, 1 ♀ ( EBCH). Zicuirán , 18.87756, -101.96943; 200 m, 8-VII-1988 GoogleMaps , G. Rodríguez, 1 ♀ ( EBCH). Lázaro Cárdenas: Chuquiapan, Playa Azul , 32 km W, 18.04952, -102.60812; 6 m, L. Godínez: 3-XI-1987 GoogleMaps , 2 ♀ ( EBCH), 3-XI-1987 , 3 ♀ ( SEMC). Los Amates, Playa Azul , 26 km N, 18.14611, -102.27833; 250 m, L. Godínez: 2-XI-1987 GoogleMaps , 1 ♀ ( EBCH), 2-XI-1987 , 3 ♀ ( SEMC). Puente Mexcalhuacan , 41 km W Playa Azul, 18.05709, -102.6576: 4 m, 3-XI-1987 GoogleMaps , T. L. Griswold , 1 ♀ ( BBSL). Múgica : 36.7 km NE de Arteaga, 19.00476, -102.10223; 384 m, 16-VII-1989 GoogleMaps , Brooks & Roig-Alsina, 1 ♀ ( SEMC). Tzitzio: Guayabo de Derrumbadero , 19.29889, -100.91917; 986 m, A. Reyes GoogleMaps , 1 ♀ ( ECOAB). Tzitzio , 2 mi S, 19.56476, -100.92203; 1383 m, 29-VII-1962 GoogleMaps , Univ. of Kansas Mex. Exped . 1 ♀ ( SEMC). Uruapan : El Sabino, 19.29139, -101.97110; 929 m, 20-VII-1936 GoogleMaps , H. Thomas, 1 ♀ ( SEMC). La Tzararacua, 19.35413, -102.07173; 1533 m, 26-V-1988 GoogleMaps , V. Hernández , 1 ♀ ( CNIN). Morelos: Acteopan: km 48 carr. Atotonilco-Atlixco , 10-IX-2009 , H. Brailovsky, E. Barrera, 1 ♀ ( CNIN). Cuernavaca : 18.93402, -99.23148; 1568 m, - VII-1951 GoogleMaps , G. Pérez, 1 ♀ ( CNIN). Cuernavaca , 18.93481, -99.23082; 1561 m, 17-III-1959 GoogleMaps , H. E. Evans, D. M. Anderson, 1 ♀ ( BBSL). Huitzilac: Derrame del Chichinautzin , km 80.7, 19.02861, -99.19278, 2620 m, 11-XI-1985 GoogleMaps , O. Yañez, 1 ♀ ( MZFC). Derrame del Chichinautzin , km 85.6, 19.02333, -99.15111; 2470 m, 6-IV-1997 GoogleMaps , I. Hinojosa, 1 ♀ ( MZFC). Jojutla: Cerro del Higuerón , 18.57333, -99.18000; 905 m, 5-VII-1981 GoogleMaps , A. Cuauhtémoc, 1 ♀ ( MZFC). Tepalcingo: Tepalcingo , 18.59222, -98.84556; 1170 m, 19-IX-1984 GoogleMaps , J. Avila, 2 ♀ ( MZFC). El Limón , 18.53805, -98.9325; 1287 m, 30-VIII-2004 GoogleMaps , E. Delgado, 1 ♀; 31-VIII-2004, D.Reynoso, 7 ♀; O. Yañez , 3 ♀ ( MZFC). Tepoztlán: km 12, Autopista México-Cuautla , 18.97333, -99.08250; 1650 m, 18-XI-1995 GoogleMaps , I. Hinojosa, 1 ♀; J. L. Salinas , 1 ♀, 17-III- 1996, I. Hinojosa, 1 ♀; O. Yañez , 1 ♀; 14-IV-1996 I. Hinojosa, 3 ♀; O. Yañez , 3 ♀; 5-V-1996, I. Hinojosa, 10 ♀; O. Yañez , 2 ♀; 13-VII-1996, I. Hinojosa, 2 ♀; I. Hinojosa : 8-IX-1996 , 1 ♀; 10-XI-1996, 4 ♀; O. Yañez , 3 ♀; I. Hinojosa : 30-XI-1996 , 3 ♀; 25-I-1997, 2 ♀; 16-II-1997, 1 ♀; O. Yañez : 16-II-1997 1 ♀; 2-III-1997, 1 ♀; I. Hinojosa : 5-IV-1997 , 2 ♀; 11-V-1997, 3 ♀; 11-V-1997, J. L. Salinas, 5 ♀; I. Hinojosa : 8-VI-1997 , 1 ♀; 5-VII-1997, 1 ♀ 1 ♂ ( MZFC). Tepoztlán, km 19.5, Autopista México-Cuautla , 18.93806, -99.03833; 1350 m, 7-X-1995 GoogleMaps , I. Hinojosa, 2 ♀; 5-XI-1995, O. Yañez, 1 ♀; 10-II-1996, I. Hinojosa, 1 ♀; I. Hinojosa : 24-II-1996 , 3 ♀; 18-IV-1996, 2 ♀; 25-V- 1996, O. Yañez, 1 ♀; 16-VI-1996, I. Hinojosa, 7 ♀; O. Yañez , 6 ♀; 20-VII-1996, I. Hinojosa, 2 ♀; 21-VII-1996, O. Yañez, 2 ♀; 11-VIII-1996, I. Hinojosa, 7 ♀; O. Yañez , 2 ♀ 1 ♂; 1-IX-1996, I. Hinojosa, 3 ♀; O. Yañez , 3 ♀; 28-IX- 1996, I. Hinojosa, 1 ♀; O. Yañez , 1 ♀; 13-X-1996, I. Hinojosa, 1 ♀; 16-IX-1996, O. Yañez, 2 ♀; 12-I-1997, I. Hinojosa, 1 ♀; O. Yañez , 1 ♀; 9-II-1997, I. Hinojosa, 3 ♀; O. Yañez , 2 ♀; 1-III-1997, I. Hinojosa, 1 ♀; 17-VIII-1997, I. Hinojosa, 1 ♂; O. Yañez , 1 ♀ ( MZFC). Tepoztlán, km 20, Autopista México-Cuautla , 18.93556, -99.03417; 1320 m, 13-IV-1997 GoogleMaps , I. Hinojosa, 1 ♀ ( MZFC). Tepoztlán, km 20.5, Autopista México-Cuautla , 18.93472, -99.02944; 1280 m, 3-VIII-1997 GoogleMaps , I. Hinojosa, 2 ♀ ( MZFC). Tepoztlán, km 4.5, Autopista México-Cuautla , 19.00417, -99.12944; 1940 m, 16-XII-1995 GoogleMaps , I. Hinojosa, 1 ♀; 27-I-1996, I. Hinojosa, 1 ♀; 16-III-1996, I. Hinojosa, 3 ♀; O. Yañez , 2 ♀; 13-IV-1996, I. Hinojosa, 2 ♀; O. Yañez , 1 ♀; 28-IV-1996, O. Yañez, 1 ♀; 23.XI-1996, I. Hinojosa, 1 ♀; 15-XII- 1996, I. Hinojosa, 1 ♀; 26-I-1997, I. Hinojosa, 3 ♀; 15-II-1997, I. Hinojosa, 5 ♀; O. Yañez , 3 ♀; 9-III-1997, I. Hinojosa, 2 ♀ ( MZFC). Tepoztlán, km 8 Carr. Tepoztlán-Cuautla , 18.93917, -99.04500; 1357 m, 23-VIII-1981 GoogleMaps , R. Ayala , 1 ♀ ( MZFC). Tlaltizapán: Yautepec , 7.3 mi S, 18.80105, -99.09742; 1011 m, 17-VIII-1962 GoogleMaps , M. Naumann, 1 ♀ ( SEMC). Tlaquiltenango: Huaxtla , 18.393314, -99.050249; 1160 m, 20-IX-2015 GoogleMaps , I. Hinojosa, 3 ♀ ( CNIN). Sierra de Huautla, CEAMISH, Huatla , 2.5 km N, 4.0 km W, 18.44979, -99.03335; 1109 m, 7-IX-1996 GoogleMaps , R. Brooks , 1 ♀ ( SEMC). Yautepec: km 21, Autopista México-Cuautla , 18.93194, -99.02583; 1283 m, 15-III-1997 GoogleMaps , I. Hinojosa, Jacaranda mimosifolia , 2 ♀ ( MZFC). Oaxtepec , 18.90637, -98.97026; 1379 m, 22-V-1989 GoogleMaps , A. Rodríguez, 1 ♀; F. A. Noguera , 1 ♀ ( EBCH). Yautepec , 4 mi. SW, 18.85786, -99.11771; 1250 m, 2-VII-1961 GoogleMaps , C. D. Michener, 16 ♀ ( SEMC). Nayarit: Acaponeta : 22.50202, -105.35524; 31 m, 29-IX-1966 GoogleMaps , G. E. Bohar, A. S. Bohart, 1 ♀ ( BBSL). Río Quimichis , 22.36472, -105.54583; 7 m, 12-III-1981 GoogleMaps , R. López , 3 ♀ ( MZFC). Bahía de Banderas: Punta Mita , 3 km S, 20.76028, -105.47487; 22 m, 28-II-1992 GoogleMaps , D. Brzoska, 1 ♀ ( SEMC). Compostela: Guayabitos 4 km E, 21.02696, -105.22536; 70 m, 16-VII-1989 GoogleMaps , C. Cordero, 1 ♀ 1 ♂; S. Anaya , 1 ♀ ( EBCH). El Nayar: Tepic , 20 mi N, 21.80975, -104.87740; 344 m, 16-II-1974 GoogleMaps , G. E. Bohart, 1 ♀ ( BBSL). San Blas : La Bajada, 21.50499, -105.15666; 193 m, 13-X-1979 GoogleMaps , J. Llorente, 1 ♀, 8-XII-1980, M. T. Ortis , 1 ♀ ( MZFC). San Blas , 21.53333, -105.28333; 2 m, L. A. Navarrete: 30-III-1981 GoogleMaps , 1 ♀; 19-X-1982, 1 ♀ ( MZFC). San Blas , 21.53333, -105.28333; 2 m, 11-III-1972 GoogleMaps , F. D. Parker, D. Miller, 1 ♀ ( BBSL). San Blas 20 km S, 21.43396, -105.18722; 10 m, 7-IV-1994 GoogleMaps , D. Yanega, 1 ♀; W.E. LaBerge , 1 ♀ ( EBCH). San Blas , 6 mi E, 21.53859, -105.20945; 60 m, 27-VIII-1959 GoogleMaps , Menke & Stange, 1 ♀ ( SEMC). Santa María del Oro, Santa Isabella, 9 mi NW, 21.27489, -104.6485; 1291 m, 10-III-1972 GoogleMaps , F. D. Parker, D. Miller, 1 ♀ ( BBSL). Tecuala: Acaponeta , 16 mi S, 22.28710, -105.27176; 45 m, 12-VIII-1963 GoogleMaps , M. Naumann, 1 ♀ ( SEMC). Tepic : La Yerba, 21.53167, -105.06167; 1067 m, 24-III-1982 GoogleMaps , J. Llorente, 1 ♀ ( MZFC). Reserva de Biosfera Sierra de San Juan , 21.51785, -104.97235; 1080 m, 20-X-2013 GoogleMaps , 13:00 h, J. Mérida, 1 ♀ ( ECOAB). Sierra San Juan , 11-X-1989 , A. Cadena, 1 ♀ ( CNIN). Tepic , 16 mi NW, 21.68103, -105.04519; 270 m, 19-VII-1953 GoogleMaps , Univ. of Kansas Mex. Exped . 3 ♀ ( SEMC). Tepic , 17 mi NW, 21.68103, -105.04519; 266 m, 23-XI-1948 GoogleMaps , E. S. Ross, 1 ♂ ( BBSL). Venustiano Carranza , 21.52000, -104.98500; 1200 m, 24-XI-1980 GoogleMaps , J. Llorente, 1 ♀; 23-III-1982, M. A. Luis, 4 ♀; 24-III-1982, M. A. Luis, 8 ♀; 25-III-1982, J. Llorente, 3 ♀; M. A. Luis , 1 ♀; 28-III-1982, M. A. Luis, 1 ♀; 20-IV-1982, A. López, 1 ♀ ( MZFC). Xalisco: El Refilón, km 35 Carr. Tepic-Puerto Vallarta , 21.31667, - 104.90000; 902 m, 19-VIII-1982 GoogleMaps , R. López , 4 ♀ ( MZFC). Oaxaca: Candelaria Loxicha: Candelaria Loxicha , 15.9205, -96.5070; 490 m, 20-XII-2013 GoogleMaps , N. Arnold, 1 ♀ ( ECOAB). Carretera a Pochutla , 15.97210, -96.48030; 1250 m, 3-V-2012 GoogleMaps , P. Sagot, 1 ♀ ( ECOAB). Pluma Hidalgo: Cascada Mayor , 15.92584, -96.417702, 1341 m, O. Yañez: 19-IX-2007 GoogleMaps , 1 ♀; 20-IX-2007, 3 ♀ ( MZFC). Pluma Hidalgo-Vereda de la Bomba de agua, 15.92584, - 96.417702; 1349 m, 18-IX-2007 GoogleMaps , O. Yañez, 1 ♀ ( MZFC). Ranchería San Isidro , 15.940394, -96.400701; 1296 m, 19-IX-2007 GoogleMaps , O. Yañez, 1 ♀ ( MZFC). Salina Cruz : 16.18608, -95.19235; 93 m, 7-IX-1965 GoogleMaps , D. Janzen, 1 ♀ ( SEMC). San Dionisio del Mar : 16.33708, -94.76381; 7 m, 20-I-1992 GoogleMaps , L. Godínez, 1 ♀ ( SEMC). San Gabriel Mixtepec : 16.11053, -97.06561; 718 m, 10-XII-2014 GoogleMaps , 9:00 h, I. Pérez, 1 ♀ ( ECOAB). San Jacinto Tlacotepec : 16.51826, - 97.35996; 1129m, 13-XII-2018 GoogleMaps , 10:35 h, I. Pérez, 1 ♀; P. Sagot , 2 ♀ ( ECOAB). San Juan Lajarcia: Carretera 190, 16.52987, -95.93127; 1110 m, 14-I-2016 GoogleMaps , 14:50 h, P. Sagot, 2 ♀ ( ECOAB). San Miguel Chimalapa: Vista Hermosa , 16.67893, -94.79296; 140 m, 12-IV-2016 GoogleMaps , 11:00 h, N. Arnold, 1 ♀ ( ECOAB). San Pedro Comitancillo : 16.45254, -95.20089; 112 m, 15-XI-2018 GoogleMaps , 9:20 h, P. Sagot, 1 ♀ ( ECOAB). San Pedro Mixtepec: Puerto Escondido , 15.87646, -97.10498; 10 m, 2-IX-1990 GoogleMaps , L. Godínez, 3 ♀ ( SEMC). Puerto Escondido 10 km NW, 15.92442, - 97.14531; 12 m, 6-III-1989 GoogleMaps , A. Rodríguez, 1 ♀; R. Ayala , 3 ♀ ( EBCH). San Pedro Pochutla: Puerto Ángel , 15.6643, -96.4883; 20 m, 9-III-1989 GoogleMaps , R. Ayala , 1 ♀ ( EBCH). Toltepec , 15.8764, -96.4445; 380 m, 3-IX-1987 GoogleMaps , R. Barba , 1 ♀ ( CNIN). San Pedro Tututepec: Laguna Chacahua , 16.03333, -97.67444; 5 m, 20-VIII-1984 GoogleMaps , J. J. Pérez, 1 ♀ ( MZFC). San Pedro Tututepec , 15.9671, -97.1921; 246 m, 18-XII-2018 GoogleMaps , 11:45 h, O. Caballero, 1 ♀ ( ECOAB). San Pedro Tututepec , 15.9487, -97.2825; 64 m, 18-XII-2018 GoogleMaps , 13:40 h, P. Sagot, 1 ♀ ( ECOAB). Santa Lucía Monteverde: Peña Flor de Clavo , 16.90850, -97.77870; - 976 m, 9-IX-2019 GoogleMaps , E. Caballero, 1 ♀ ( ECOAB). Santa María Colotepec : 15.86330, -97.00450; 24 m, 19-XII-2018 GoogleMaps , 8:45 h, I. Pérez, 1 ♀ ( ECOAB). Santa María Huatulco: Aeropuerto , 15.75778, -96.25673; 115 m, 8-I-2010 GoogleMaps , P. Sagot, 1 ♀ ( ECOAB). Santa María Tonameca: carretera a Loxicha , 15.8054, -96.7603; 155 m, 20-XII-2018 GoogleMaps , 9:25 h, I. Pérez, 1 ♀ ( ECOAB). Santa María Yucuhiti: San José Zaragoza-Miramar , 16.98825, -97.7924; 1604 m, 5-XI-2015 GoogleMaps , I. Hinojosa, A. C. Martínez, 1 ♀ ( CNIN). Santiago Niltepec : Microondas Los Tules, Niltepec , 16.50, -94.58; 44 m, 23-XI-1983 , R. Torres , 1 ♀ ( EBCH). Santo Domingo de Morelos: 15.8756, -96.6941; 288 m, 20-XII-2018 GoogleMaps , 11:25 h, I. Pérez, 2 ♀; O. Caballero , 1 ♀; P. Sagot , 2 ♀ ( ECOAB). Santo Domingo Tehuantepec: Tehuantepec , 16.31853, -95.24061; 44 m, 2-VII-1957 GoogleMaps , Univ. of Kansas Mex. Exped . 1 ♀ ( SEMC). Tehuantepec , 12 mi W, 16.29339, -95.41169; 404 m, 16-IX-1974 GoogleMaps , G. E. Bohart, W. Hanson, 3 ♀ ( BBSL). Tehuantepec , 14 mi NW, 16.39868, -95.40926; 242 m, 26-VI-1961 GoogleMaps , Univ. of Kansas Mex. Exped . 1 ♂ ( SEMC). Tehuantepec , 17 km W, 16.34649, -95.39159; 176 m, D. Janzen GoogleMaps , 3 ♀ ( SEMC). Santo Domingo Zanatepec : 16.46121, -94.23886; 196 m, 20-XII-2019 GoogleMaps , 10:50 h, E. Gómez, 3 ♀; P. Sagot , 1 ♀ ( ECOAB). General Pascual Fuentes , 16.48069, -94.21259; 590 m, 10-IV-2016 GoogleMaps , 10:00 h, L. Reyes, 1 ♀ ( ECOAB). Tapanatepec , 8 mi W, 16.45387, -94.26291; 122 m, 16-VII-1951 GoogleMaps , Univ. of Kansas Mex. Exped . 1 ♀ ( SEMC). Puebla: Acatlán : 9 mi NW, 18.22280, -98.12673; 1380 m, 13-VI-1953 GoogleMaps , Univ. of Kansas Mex. Exped . 1 ♀ ( SEMC). Chinantla: Tehuitzingo , 8 mi SE, 18.25606, -98.23040; 1202 m, 29-VI-1961 GoogleMaps , Univ. of Kansas Mex. Exped . 1 ♀ ( SEMC). Huehuetlán el Grande: Bienes comunales, 18.75558, -98.17335; 1345 m, 2-IX-2018 GoogleMaps , 9:25, L. Marín, 1 ♀ ( ECOAB). Casa Esmeralda , 18.74948, -98.17328; 1346 m, 3-I-2018 GoogleMaps , L. Marín, 1 ♀ ( ECOAB). Izúcar de Matamoros : 7 km al SE de San Juan Raboso, 18.516, -98.412; 1483 m, 11-XI-2015 GoogleMaps , 13:55 h, N. Brasero, 1 ♀ ( ECOAB). Carr. Fed. Izúcar de Matamoros-Huajuapan de León , 18.45619, -98.37350; 1156 m, I. Hinojosa, 4-IX-1998 GoogleMaps , 1 ♀. Xicotepec : La Ceiba, 20.390896, -97.877357; 239 m, 29-III-1956 GoogleMaps , C. Bolivar, J. Ordoñez, 1 ♀ ( CNIN). Sinaloa: Badiraguato: Tameapa , 25.62674, -107.37219; 1058 m, 17-X-2013 GoogleMaps , 10:35 h, J. Mérida, 1 ♀ ( ECOAB). Concordia: Copala, 18.5 km E on hwy 40, at Chirimoyos , 23.43750, -105.81194; 1587 m, 17-III-1990 GoogleMaps , 3 ♀ ( SEMC). Mazatlán : 10 mi W Concordia, 23.28402, -106.22299; 24 m, 30-X-1965 GoogleMaps , G. E. Bohart, A. S. Bohart, 1 ♀ ( BBSL). Mazatlán , 23.22670, -106.40760; 15 m, 25-VIII-1976 GoogleMaps , 2 ♀ ( BBSL). Villa Unión , 6 mi N, 23.27230, -106.21729; 21 m, 9-IX-1970 GoogleMaps , W. J. Hanson, T. L. Whitworth , 1 ♀ ( BBSL). Veracruz: Acayucán : 17.94833, -94.91333; 108 m, 16-II-1954 GoogleMaps , R. Dreisbach , 1 ♀ ( SEMC). Actopan: Jose Cardel dunes, 20 km. N, 19.54961, -96.39262; 8 m, 7-XI-1991 GoogleMaps , T. L. Griswold , 1 ♀ ( BBSL). Catemaco : 18.42212, -95.11384; 364 m, 6-IX-1974 GoogleMaps , W. J. Hanson, G. E. Bohart, 1 ♀ ( BBSL). Cotaxtla: Exp. Sta. Cotaxtla , 18.83354, -96.39402; 67 m, 3-IX-1962 GoogleMaps , D. H. Janzen, 1 ♀ ( EBCH). Emiliano Zapata : 41 km SE Xalapa, 19.36085, -96.57331; 273 m, 7-IX-1991 GoogleMaps , R. Ayala , 1 ♀ ( EBCH). Medellín de Bravo: Cotaxtla Experimental Station , 18.93274, -96.19205; 22 m, 5-VII-1962 GoogleMaps , D. Janzen, 1 ♀ ( SEMC). Teocelo: Barranca de Cayoapa Tejería , 19.37333, -96.92667; 993 m, 28-II-1981 GoogleMaps , A. Garces, 1 ♂ ( MZFC). Teocelo , 19.3852, -96.97159; 1160 m, 21-III-1989 GoogleMaps , I. Hinojosa, 1 ♀; O. López , 1 ♀ ( MZFC). Tierra Blanca: km 1.5 a Jícaro , 18.54472, - 96.25000; 49 m, 20-III-1989 GoogleMaps , I. Hinojosa, 2 ♀; O. López , 2 ♀ ( MZFC). NICARAGUA: Granada: Reserva Domitila , 11.70833, -85.95333; 100 m, 8-VI-2002 GoogleMaps , R. Brooks, Z. Falin, S. Chatzimanolis , 1 ♀, 9-VI-2002, R. Brooks, Z. Falin, S. Chatzimanolis , 1 ♀; 8.VI-2011, C.W.Shin, 1 ♀ ( SEMC). Reserva Natural Volcán Mombacho , 11.84166, -85.98055; 1200 m, 2-VI-2002 GoogleMaps , R. Brooks, Z. Falin, S. Chatzimanolis , 1 ♀ ( SEMC). San Carlos airstrip, 11.13361, -84.80416; 100 m, 1-VI-2002 GoogleMaps , R. Brooks, Z. Falin, S. Chatzimanolis , 2 ♀ ( SEMC). Rivas: Cárdenas: Miguel Ángel Soto , 11.19744, -85.66385; 195 m, 1-II-2016 GoogleMaps , M. Calero, 4 ♀, R. Joyce , 1 ♂ ( ECOAB). San Juan del Sur: Hacienda la Flor , 11.14192, -85.78290; 16 m, 17-II-2016 GoogleMaps , M. Calero, 1 ♀ ( ECOAB). Río Ostional , 11.12598, -85.74980; 22 m, 11-II-2016 GoogleMaps , M. Calero, 1 ♀ ( ECOAB). PANAMÁ : 1 ♀ ( BBSL). Panamá: Canal Zone, Summit , 9.06516, - 79.64544; 90 m, 26-XI-1930 GoogleMaps , H. Schwarz, 1 ♀ ( SEMC). Chiriquí: Boquete : 8 km S, 8.69858, -82.45046; 820 m, 5-I-2012 GoogleMaps , F. D. Parker, T. D. Mcintyre , 4 ♀ 1 ♂; 14-I-2012, F. D. Parker, T. D. Mcintyre , 1 ♀; 21-I-2012, F. D. Parker, T. D. Mcintyre , 1 ♀ ( BBSL). Boquete , 8.76667, -82.43333; 1061 m, 17-I-1981 GoogleMaps , C. D. Michener, 2 ♀ ( SEMC). Gualaca: Jorge Arauz, 4 km N, IRHE Fortuna Station , 8.69999, -82.23333; 1200 m, 17-VI-1993 GoogleMaps , S. Lingafelter, 1 ♀ ( SEMC). Cocle: Penonomé : 8.54535, -80.35419; 103 m, 30-XI-1925 GoogleMaps , 1 ♀ ( SEMC). Panamá: Old Panamá, 9.00805, -79.48488; 10 m, 15-III-1945 GoogleMaps , C. D. Michener, 1 ♀; 19-IV-1945, C. D. Michener, 2 ♀ ( SEMC). Panamá: Balboa, Albrook Field , 8.96984, -79.56218; 5 m, 24-X-1937 GoogleMaps , L. Stannard, 1 ♀ ( SEMC). Balboa , 8.95049, -79.56610; 5 m, 20-XII-1959 GoogleMaps , W. J. Hanson, 1 ♀ ( BBSL). Cerro Azul, N. of Tecumén , 8.80000, -82.38333; 2180 m, 5-IX-1971 GoogleMaps , N. Lee, 1 ♂ ( BBSL). Panamá City, 8.99861, -79.51073; 30 m, 10-V-1945 GoogleMaps , C. D. Michener, 1 ♀ ( SEMC). Panamá Oeste: Chame : 8.57884, -79.87673; 18 m, 29-IV-1945 GoogleMaps , C. D. Michener, 5 ♀ ( SEMC) .

Commentary. In areas where species of the frontalis group are sympatric, X. fimbriata may be confused with X. frontalis and X. nautlana , from which it may be distinguished as it presents frontal elevational characteristics, the prominences adjacent to the ocelli in the form of horns; males are distinguished because they do not present black integument on any of the terga of the metasoma.

MNCR

Museo Nacional de Costa Rica

R

Departamento de Geologia, Universidad de Chile

SEMC

University of Kansas - Biodiversity Institute

BBSL

USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research

CNIN

Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico

V

Royal British Columbia Museum - Herbarium

MZFC

Museo de Zoologia "Alfonso L. Herrera"

T

Tavera, Department of Geology and Geophysics

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Hymenoptera

Family

Apidae

Genus

Xylocopa

Loc

Xylocopa (Neoxylocopa) fimbriata Fabricius, 1804

Mérida-Rivas, Jorge A., Hinojosa-Díaz, Ismael A., Ayala-Barajas, Ricardo, Pozo, Carmen & Vandame, Rémy 2022
2022
Loc

Xylocopa cajennae

Smith, F. 1874: 284
Lepeletier, A. L. M. 1841: 203
1841
Loc

Xylocopa cornuta

Erichson, W. F. 1848: 591
Lepeletier, A. L. M. 1841: 176
1841
Loc

Xylocopa virescens

Moure, J. S. 1960: 144
Lepeletier, A. L. M. 1841: 186
1841
Loc

Xylocopa corniger

Smith, F. 1874: 284
Westwood, J. O. 1840: 270
1840
Loc

Xylocopa fimbriata

Moure, J. S. 1960: 144
Fabricius, J. C. 1804: 340
1804