Xylocopa (Neoxylocopa) gualanensis Cockerell, 1912

Mérida-Rivas, Jorge A., Hinojosa-Díaz, Ismael A., Ayala-Barajas, Ricardo, Pozo, Carmen & Vandame, Rémy, 2022, Revision of carpenter bees of the subgenus Neoxylocopa Michener (Hymenoptera: Apidae) from Mexico and Mesoamerica, Zootaxa 5158 (1), pp. 1-67 : 39-46

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https://doi.org/10.11646/zootaxa.5158.1.1

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DOI

https://doi.org/10.5281/zenodo.6821841

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Xylocopa (Neoxylocopa) gualanensis Cockerell, 1912
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Xylocopa (Neoxylocopa) gualanensis Cockerell, 1912

( Figs. 66–74 View FIGURES 66–70 View FIGURES 71–74 , 120–122 View FIGURES 120–125 , 138 View FIGURES 136–140 )

Xylocopa wilmattae gualanensis Cockerell, 1912: 555 . Holotype: American Museum of Natural History , New York, USA (♀). Type locality: Guatemala, Gualan.

Diagnosis. Body length, female, 21.56 mm (17.19–23.86; n=5); male, 21.53 mm (20.29–21.93; n=5). Female: Head without elevated ridges or prominences on frons; ocelli below tangent of the eyes; area between parapsidal line and mesoscutal disc with scattered punctures (separated by two or more times their diameter); wings bluish with purple iridescence basally and on middle area, papillate region with dark green iridescence ( Fig. 69 View FIGURES 66–70 ); pubescence evenly black; T1 densely punctate on discal area (punctures separated by one diameter or less), T2 and T3 less densely punctate (punctures separated by two or three diameters) ( Fig. 70 View FIGURES 66–70 ); T4 with punctures separated by three or more times their diameter; length of hairs on terga respect MOD, T2 0.25–0.37, T3 0.29–0.60, T4 0.44–0.88, T5 0.57– 1.08. Male: Terga yellowish-orange, without bands; metasoma with yellowish-orange pubescence; Posterior tibia internal distal extreme with prominent subapical projection, recurved distally defining a evident concave area; posterior margin of this process with a well-defined laminar margin of almost a third of the internal posterior margin of the tibia ( Fig. 74 View FIGURES 71–74 ); length of hairs on terga respect MOD, T2 0.27–0.51, T3 0.72–1.17, T4 0.67–1.58, T5 0.90–1.58. Tip of the lateral process of gonostylus ending oval-shaped (as broad as shaft of the process).

Description. Female: Integument and pubescence usually black; wings bluish with purple iridescence on basal and middle areas; papillate region with dark green iridescence ( Fig. 69 View FIGURES 66–70 ); venation black. Body length 21.56 mm (17.19–23.86; n=5); length of forewing 18.31 mm (17.13–20.19; n=5); length of head 5.31 mm (4.94–5.67; n=5); intertegular distance 6.56 mm (5.95–6.97; n=5); width of T2 9.62 mm (8.46–10.26; n=5) Head: Clypeus, genal area and vertex with densely punctate. Mesosoma: between the parapsidal line and discal area with scattered punctures, separated by two or more times their diameter; scutellum convex in profile. Metasoma: T1 with dense punctures on discal area, separated by one time or less their diameter. T2 and T3 with dense punctures, separated by two or three times their diameter; T4 with punctures separated by three or more times their diameter; T5 and T6 with dense appressed punctures. Length of hairs on terga respect MOD as follows: T2 0.25–0.37, T3 0.29–0.60, T4 0.44–0.88, T5 0.57–1.08. Male: Integument and pubescence yellowish-orange. Head: Clypeus with a dark brown middle longitudinal line that reaches the discal region ( Fig. 71 View FIGURES 71–74 ). Wings light brown (orange), venation brown light to reddish. Body length 21.53 mm (20.29–21.93; n=5); length of forewing 17.19 mm (16.28–17.86; n=5); head length 3.94 mm (3.82–4.02; n=5); intertegular distance 6.03 mm (5.68–6.63; n=5); width of T2 9.75 mm (9.35–9.94; n=5). Head: Labrum with narrow triangular middle unpunctate area; scattered punctures on the submargin half of the labrum; marginal areas unpunctate. Clypeus: lower margin unpunctate; disc with very scattered punctures; rest of clypeus with scattered punctures, separated one and a half times or more their diameter. Supraclipeal area, paraocular areas, and genal area with dense punctures, similar in size to those on clypeus. With an unpunctate area posterior to the ocelli, of half diameter of the ocelli. Vertex with unpunctate area on the middle; rest of vertex with densely punctate. Mesosoma: Between the parapsidal line and discal area densely punctate, separated by one to one and a half times their diameter. Hind tibia: internal distal extreme with a prominent subapical projection, with recurved distal extreme definining a very evident concave area; posterior margin of this process with a well-defined laminar margin of almost a third of the internal posterior margin of the tibia; internal surface hairless; anterior margin with long hairs, denser and shorter in the apical region; posterior margin with shorter pubescence with long and short hairs ( Fig. 74 View FIGURES 71–74 ). Metasoma: T2 to T6 with a narrow middle longitudinal line unpunctate; T1 to T7 with densely uniform punctate, separated by approximately twice their diameter. Length of hairs on terga respect MOD as follows: T2 0.27–0.51, T3 0.72–1.17, T4 0.67–1.58, T5 0.90–1.58. Genitalia: Distal angle of the ventral section of the gonostylus, acute ( Fig. 122 View FIGURES 120–125 ); basal angle at the base of lateral process of the gonostylus, obtuse, barely noticeble ( Fig. 121 View FIGURES 120–125 ); tip of the lateral process ending on an oval as broad as shaft of the process ( Fig. 121 View FIGURES 120–125 ).

Variation. Throughout the distribución, both sexes present variation in body size and wing iridescence, in the female from dark blue with greenish, to blue and violet with greenish iridescence; the iridescence in the papillate region is green; or wings magenta similar to those of X. griswoldi sp. nov. may be a complex of cryptic species, to demostrate this would require molecular studies as well as studies of their distribution.

Distribution ( Fig. 138 View FIGURES 136–140 ). Costa Rica, Guatemala, Mexico, Nicaragua, in the following ecoregions ( Dinerstein et al. 2017): Dry forests of the Balsas, Dry forests of Central America, Mountainous forests of Central America, Pine-oak forests of Central America, Central Mexican Scrubland, Dry forests of the depresion of Chiapas, Seasonal humid forests of Costa Rica, Pacífic isthmus Humid forests, Dry forests of Jalisco, Mesoamerican-Caribbean Gulf, Spiny scrubland of the Motagua valley, Mangroves of the Mesoamerican Pacific north, Swamps of Centla, Humid forests of Petén-Veracruz, Humid forests of the Sierra Madre of Chiapas, Pine-oak forests of the Sierra Madre del Sur, Pine-oak forests of the Sierra Madre Occidental, Dry forests of Sinaloa, Mangroves of the South Pacífic of Mesoamerica, Dry forests of the South Pacific, Pine-oak forests of the Transmexican Volcanic Belt, and Scrubland of the Tehuacán Valley. Broad altitudinal distribution: 0 to 2132 masl.

Material examined. 680 ♀ 28 ♂: COSTA RICA : 3 ♀ ( MZUCR); Alajuela : Palmares: Zaragoza , 10.03955, -84.42252; 998 m, 6-V-2007 GoogleMaps , F. Ruíz, 1 ♀ ( MZUCR). Guanacaste: Finca Montezuma , 3 km SE Río Naranjo, 10.66409, -85.09720; 452 m, 19-V-1993 GoogleMaps , F. D. Parker, 1 ♀ ( BBSL). Hacienda Palo Verde , 2 km E, 10.34990, - 85.33169; 42 m, -1-1974, K. H. Keeler GoogleMaps , 1 ♀ ( BBSL). Bagaces: Guanacaste 2.5 km E de Bagaces, 10.51667, - 85.23334; 40 m, 22-XI-2001 GoogleMaps , J. Weng, 2 ♀ ( MNCR). Lomas de Barbudal , 10.46667, -85.35001; 90 m, 18-XI-2001 GoogleMaps , F. Umaña, 1 ♀; A. Soto, 1 ♀ ( MNCR). P. N. Palo Verde, Sector Palo Verde, 10.36667, -85.38326; 107 m, 8-II-2000 GoogleMaps , I. Jímenez, 1 ♀ ( MNCR). Cañas: 10.42670, -85.09590; 85 m, 14-XI-1971 GoogleMaps , F. Cordero, 2 ♀ 2 ♂ ( BBSL). Cañas , 14 km S, EJN, 10.30, -85.0956, 36 m, 8-III-1988 , F. D. Parker: 15 ♀, 9-VII-1988, 1 ♀; 24-III-1989, 1 ♀; 7-V-1989, 2 ♀; 28-X-1989, 1 ♀; 11-I-1990, 1 ♀; 15-II-1990, 1 ♀ 1 ♂; 1-IV-1990, 2 ♀; 20-VII-1990, 1 ♂; 1-VIII-1990, 5 ♀; 1- X-1990, 1 ♀; 16-III-1991, 1 ♀; 12-II-1991, 2 ♀ ( BBSL). Cañas , 4 km N, 10.46554, -85.09310; 97 m, 15-II-1966 GoogleMaps , D. Janzen, 1 ♀ ( SEMC). La Pacifica , 4 km NW Cañas , 10.48349, -85.13790; 75 m, 20-VI-1971 GoogleMaps , F. Cordero, 1 ♂; 15-XI-1971, F. Cordero, 1 ♀; P. A. Opler, 1 ♀; 1-III-1974, P. A. Opler, 1 ♀ ( BBSL). Carrillo: Playas del Coco , 10.54694, -85.69388; 15 m, 23-IV-1961 GoogleMaps , A. Wille, 1 ♀ ( MZUCR), 10-VII-1965 , S. J. Arnold, 1 ♀, 14-VII-1965, S. J. Arnold, 1 ♀; 21-VIII-1965, S. J. Arnold, 1 ♀ ( EBCH). La Cruz: P. N. Guanacaste, Agua Buena, 11.03366, - 85.57695; 220 m, 1-II-1992 GoogleMaps , 1 ♀ ( MNCR). P. N. Guanacaste, Cerro el Hacha, 10.89985, -85.57733; 400 m, 11-I- 1988 GoogleMaps , A. Solís, 1 ♀ ( MNCR). Santa Elena, Finca Jenny , 10.86551, -85.57354; 200 m, 1-IX-1988 GoogleMaps , 2 ♀; 1-X-1988, 19 ♀; 1-XI-1988, 4 ♀; 1-XII-1988, 9 ♀; 1-III-1989, 2 ♀ ( MNCR). Liberia: Curubandé , 16.4 km NE Liberia, 10.71667, -85.38334; 360 m, 22-XI-2001 GoogleMaps , C. Godoy, 1 ♀; J. Weng, 2 ♀ ( MNCR). Mayorga, Río Góngora , 10.88623, -85.47209; 600 m, 1-VI-1992 GoogleMaps , 1 ♀ ( MNCR). Nacascolo, P.N.S. R., Playa Naranjo, 10.80271, -85.67479; 1 m, 1-XII- 1990 GoogleMaps , E. Alcazar, 1 ♀ ( MNCR). P. N. Sta. Rosa, Estación Santa Rosa, 10.83641, -85.61549; 300 m, 1-II-1992 GoogleMaps , R. Moraga , 3 ♀; D. Brenes, 1 ♀; 1-VI-1992, Hayata , 3 ♀ ( MNCR). Sector Las Pailas , 4.5 km SW del Volcán Rincón de la Vieja, 10.77678, -85.35191; 800 m, 23-IX-1992 GoogleMaps , C. Cano, 1 ♀; 5-VIII-1994, D. García, 1 ♀ ( MNCR). Nicoya: San Antonio , 3 km NO de Nacaome, P. N. Barra Honda, 10.16822, -85.37366, 100 m, 1-II-1997 GoogleMaps , M. Reyes, 1 ♀ ( MNCR). Santa Cruz : Cuajiniquil W, 10.07560, -85.66249; 250 m, 12-II-1995 GoogleMaps , T. L. Griswold , 2 ♀ ( BBSL). Santa Cruz , R. V. S. Bosque Diriá, Poza Redonda, 10.17374, -85.59589; 150 m, 24-XI-1988 GoogleMaps , R. Bartón , 1 ♀ ( MNCR). Tilarán: Santa Rosa Biological Station , 10.83641, -85.61549; 300 m, 10-III-1986 GoogleMaps , W. Wcislo, 2 ♀ ( SEMC). Santa Rosa National Park , headquarters area, 10.83409, -85.61329; 280 m, 12-II-1995 GoogleMaps , T. L. Griswold , 1 ♀ ( BBSL). Limón : Pococí: Colorado, Estación Cuatro Esquinas , 10.53955, -83.50648; 7 m, 1-IV-1989 GoogleMaps , R. Aguilar , 1 ♀ ( MNCR). Puntarenas: Estación San Miguel , 3 km. NW de Cabo Blanco, 9.58121, -85.13671; 50 m, 1-X-1993 GoogleMaps , 1 ♀ ( MNCR). Golfito , 8.63920, -83.16240; 130 m, 10-IV-1991 GoogleMaps , F. D. Parker, J. B. Welch, 1 ♀ ( BBSL). Isla San Lucas , Puesto Isla San Lucas, 9.94652, -84.90556; 0 m, 7-III-2005 GoogleMaps , Y. Cárdenas, 2 ♀; J. A. Azofeita, 4 ♀ ( MNCR). Paquera, Isla Pan de Azúcar , 9.92643, -84.90411; 0 m, 29-XI-2005 GoogleMaps , J. A. Azofeita, 2 ♀ ( MNCR). San José : Escazú: 9.91670, -84.13330; 1100 m, 31-I-1988 GoogleMaps , F. D. Parker, 1 ♀; 14-I-1989, F. D. Parker, 1 ♀ ( BBSL). Hatillo 3, 9.91649, -84.10458; 1100 m, 4-IV-2000 GoogleMaps , M. Law, 1 ♀ ( MZUCR). San Francisco de dos Ríos, 9.9166, -84.04989; 1100 m, - IV-1988 GoogleMaps , A. Bernecker, 1 ♀; P. Wiesner, 1 ♀; 30-VIII-1994, 1 ♀ ( MZUCR). Montes de Oca , 9.93651, -84.04882; 1200 m, 3-I-1984 GoogleMaps , 1 ♀; - V-1996, B. Roberto, 1 ♀ ( MZUCR). GUATEMALA; Chimaltenango: Finca el Recuerdo, 14.475, -91.122; 483 m, 23-II-2010 GoogleMaps , C. L. Yurrita, 1 ♀ ( CANG). Guatemala: Universidad de San Carlos , 14.58441, -90.55345; 1487 m, 2016, G. Quiñones GoogleMaps , 1 ♀ ( ECOAB). Huehuetenango : La Democracia: Cajuil , 15.63746, - 91.78722; 1026 m, 2-III-2010 GoogleMaps , J. Mérida, 1 ♂ ( CANG), 18-XI-2010 , 9:00 h, O. Argüello, 1 ♀ ( ECOAB). Pinalito , 15.6653, -91.8618; 963 m, 2-III-2010 GoogleMaps , 8:50 h, M. Cigarroa, 1 ♀ ( ECOAB). San Antonio Huista: Miralvalle , 15.66789, -91.95520; 1127 m, 8-V-2010 GoogleMaps , 2 ♀ ( ECOAB). Jutiapa : El Progreso: Hacienda el Tabacal, 14.90908, - 89.99188; 293 m, E. Enriquez, 2-III-2005 GoogleMaps , 1 ♀; 5-XII-2007, 1 ♀ ( CANG). Retalhuleu : Champerico: 1.3 mi E, 14.29017, -91.88639; 10 m, 17-VII-1966 GoogleMaps , 1 ♀ ( SEMC). Retalhuleu , 5 mi W, 14.57312, -91.76502; 232 m, 19-VII- 1966 GoogleMaps , D. Janzen, 1 ♂ ( SEMC). San Marcos: Nuevo Progreso, 14.80588, -91.907277; 770 m, 25-II-2010 GoogleMaps , O. Martínez, Vernonia deppeana, 1 ♀ ( CANG). Zacapa: Río Hondo , El Senegal, 15.220832, -89.632628; 1077 m, 1-III- 2005 GoogleMaps , E. Enriquez ( CANG) . Teculután: 7 km W, 14.96972, -89.77520; 261 m, 25-VI-1993 GoogleMaps , J. Ashe, R. Brooks , 1 ♀ ( SEMC). HONDURAS: Francisco Morazán : Zamorano: 14.06600, -86.38352; 461 m, 5-V-1964 GoogleMaps , G. A. Axtell, 1 ♀ ( BBSL). MEXICO: Chiapas : 3 ♀ 3 ♂. Acacoyagua: Rosario Zacatonales , 15.43365, -92.65051; 915 m, 28- V.2005, R. Ayala GoogleMaps , 1 ♀ ( ECOAB). Amatenango de la Frontera: 15.4380, -92.1212; 1069 m, 1-XI-2018 GoogleMaps , P. Sagot, 1 ♂ ( ECOAB). Ángel Albino Corzo: Jerusalén , 15.77486, -92.64197; 885 m, 3-XI-2017 GoogleMaps , P. Sagot, 1 ♀ ( ECOAB). Arriaga: Ejido Adolfo López Mateos , 16.32131, -93.98319; 368 m, 20-I-2009 GoogleMaps , M. Guzmán, 1 ♀; 23-I-2009, C. Balboa, 1 ♀; 10-III-2009, C. Balboa, 3 ♀; M. Cigarroa, 3 ♀; M. Guzmán, 3 ♀; 13-III-2009, J. Mérida, 1 ♀; M. Guzmán, 1 ♀; 22-V-2009, M. Guzmán, 2 ♀; 17-VII-2009, C. Balboa, 1 ♀; 17-XI-2009, C. Balboa, 1 ♀; M. Guzmán, 1 ♀; 20-XI-2009, C. Balboa, 1 ♀ ( ECOAB). Paraje Poza Verde , 16.34207, -93.95440; 466 m, 24-I-2009 GoogleMaps , C. Balboa, 1 ♀; M. Guzmán, 1 ♀; 20-V-2009, C. Balboa, 1 ♀; 18-XI-2009, M. Guzmán, 1 ♀ ( ECOAB). Chiapa de Corzo: Nicolás Bravo , 16.50111, -92.98249; 440 m, 10-III-2016 GoogleMaps , 8:40 h, P. Sagot, 1 ♀ ( ECOAB). Comitán de Domínguez: 16.25111, -92.13416; 1643 m, 31-III-2016 GoogleMaps , 7:25 h, P. Sagot, 1 ♀ ( ECOAB). Frontera Comalapa: Los Laureles , 15.88592, -92.11727; 575 m, 31-III-2016 GoogleMaps , 9:35 h, P. Sagot, 1 ♀ ( ECOAB). Huehuetán: 15.01789, - 92.39390; 86 m, 14-I-1988 GoogleMaps , L. Godínez, 1 ♀ ( ECOAB). La Concordia: Carretera 157, El Diamante, 16.10692, - 92.87352, 579 m, 2-III-2916 GoogleMaps , 15:08 h, P. Sagot, 3 ♀ ( ECOAB). La Trinitaria: 15.9302, -91.9860; 773 m, 3-XI- 2018 GoogleMaps , 10:05 h, P. Sagot, 3 ♂ 1 ♀ ( ECOAB). Carretera La Campana-Malpaso , 16.0271, -92.05389; 769 m, 16-XI-2011 GoogleMaps , 9:15 h, J. Mérida, 1 ♀ ( ECOAB). El Progreso, 16.1143, -91.8704; 1470 m, 21-V-2011 GoogleMaps , E. Pineda, 1 ♀ ( ECOAB). San Francisco de Asís , 16.0118, -92.0267; 767 m, 16-XI-2011 GoogleMaps , 8:00 h, J-Mérida, 2 ♀ ( ECOAB). Mapastepec: Unión Los Olivos-orilla del río, 15.5219, -92.7996; 315 m, 28-V-2005 GoogleMaps , M. Guzmán, 1 ♀, 23-X-2006, M. Cigarroa, 2 ♀ ( ECOAB). Mazapa de Madero: 15.4111, -92.1619; 890 m, 1-XI-2018 GoogleMaps , 15:05 h, P. Sagot, 1 ♂ ( ECOAB). Mazatán: Barra San José , 14.92230, -92.62698; 6 m, 2-II-2019 GoogleMaps , 9:20 h, G. Mera, 2 ♀; I. Pérez, 3 ♀; P. Sagot, 2 ♀ ( ECOAB). Mazatán , 14.85849, -92.45159; 22 m, 20-VI-1988 GoogleMaps , M. Cigarroa, 1 ♀ ( ECOAB). Motozintla de Mendoza: Río de Motozintla , 15.3827, -92.2208; 1170 m, 2-XI-2018 GoogleMaps , 8:00 h, P. Sagot, 1 ♂ ( ECOAB). 3 km SW de Motozintla , 15.35, -92.25; 1470 m, 20-IV-1993 , J. L. Neff, Jacaranda mimosifolia , 1 ♀ ( EBCH). Ocozocoautla: 8.1 km N Nuevo México S de Ocozocoautla , 16.69761, -93.41156; 773 m, 11-IV-1993 GoogleMaps , C. Mendoza, Lonchocarpus sp , 2 ♀ ( EBCH). Palenque: 17.51000, -91.98149; 78 m, 10-IX-1974 GoogleMaps , G. E. Bohart, W. Hanson, 1 ♀ ( BBSL). Pijijiapan: Carr. 200, 15.71307, -93.23767; 78 m, 1-II-2019 GoogleMaps , 14:00 h, P. Sagot, 1 ♀ ( ECOAB). Solo Dios , 15.66695, - 93.42535; 9 m, 1-II-2019 GoogleMaps , 11:25 h, I. Pérez, 5 ♀ ( ECOAB). Suchiate: Cosalapa , 14.63969, -92.26749; 10 m, 16-I- 1990 GoogleMaps , J. Sibaja, 1 ♀; 17-I-1990, J. Sibaja, 3 ♀; 24-I-1990, A. Aquino, 3 ♀; J. Sibaja, 2 ♀; 23-XI-1990, J. Sibaja, 2 ♀ ( ECOAB). Tapachula: 36.7 km N Tapachula , 15.103, -92.3097; 440 m, 17-IV-1993 GoogleMaps , A. Rodríguez, 1 ♀ ( EBCH). Carretera Jaritas , 14.78299, -92.36790; 30 m, 17-I-1990 GoogleMaps , J. Sibaja, 1 ♀; 19-I-1990, A. Aquino, 1 ♀; 24-I-1990, A. Aquino, 1 ♀; J. Sibaja, 1 ♀ ( ECOAB). Col. Seminarista, 14.89279, -92.24909; 152 m, 26-IV-1992 GoogleMaps , A. Aquino, 1 ♀; 23-I-1997, A. Aquino, 1 ♀; 18-II-1997, A. Aquino, 1 ♀; 23-III-1997, A. Aquino, 1 ♀; 8-IV-1997, A. Aquino, 1 ♀; 8-VII-1997, M. Vargas, 1 ♂ ( ECOAB). Ecosur 14.88669, -92.28660; 127 m, 4-XI-1992 GoogleMaps , 12:00 h, A. Aquino, 1 ♀ ( ECOAB). Finca Pumpuapa , 14.92638, -92.36361; 48 m, 25-II-1987 GoogleMaps , 9:15 h, L. Godínez, 1 ♀ ( ECOAB). Joaquín Miguel Gutiérrez, 14.74328, -92.31274; 28 m, 3-VII-2014 GoogleMaps 9:00 h, M. Guzmán 1 ♀; 8-V-2015, 9:00 h, J. Ruíz, 1 ♀ ( ECOAB). Lagartero , 14.8277, -92.2638; 80 m, 3-XII-1986 GoogleMaps , R. Balbuena , 1 ♀ ( ECOAB). Panteón Jardín , 14.8942, -92.2472; 156 m, 1-VII-1997 GoogleMaps , A. Aquino, 1 ♀ ( ECOAB). Rancho Santa Teresita , 14.9048, -92.3144; 102 m, 10- VIII-1987 GoogleMaps , L. Mendoza, 4 ♀ ( ECOAB). Tapachula , 14.88669, -92.28660; 127 m, 29-VIII-1986 GoogleMaps , 1 ♀ ( ECOAB). Teopisca: 16.53045, -92.46939; 1776 m, 13-IV-1993 GoogleMaps , C. Mendoza, Wigandia kunthii , 1 ♀ ( EBCH). Tonalá: Ejido Nueva Costa Rica , 16.1352, -93.6824; 408 m, 19-I-2009 GoogleMaps , C. Balboa, 5 ♀; 22-I-2009, C. Balboa, 1 ♀; 9-III-2009, P. Sagot, 2 ♀; 18-V-2009, M. Guzmán, 1 ♀; 14-IX-2009, M. Guzmán, 1 ♀; 16-XI-2009, C. Balboa, 4 ♀ ( ECOAB). El Madresal, 15.80021, -93.60083; 7 m, 19-XI-2017 GoogleMaps , 10:00 h, P. Sagot, 1 ♀ ( ECOAB). Manglar , 15.87592, -93.71470; 5 m, 31-I-2019 GoogleMaps , 13:20 h, P. Sagot, 1 ♀ ( ECOAB). Puerto Arista , 15.94839, -93.83723; 5 m, 31-1-2019 GoogleMaps , G. Mera, 1 ♀; I. Pérez, 4 ♀; P. Sagot, 6 ♀ ( ECOAB). Puerto Arista , 15.94282, -93.82458; 6 m, 9-X-2015 GoogleMaps , I. Hinojosa, 1 ♀ ( CNIN). Tuxtla Chico: El Manguito, 14.8903, -92.2314; 161 m, 13-V-1992 GoogleMaps , A. Aquino, 1 ♀; M. Rincón, 4 ♀ ( ECOAB). Rosario Izapa , 14.9733, -92.1546; 445 m, 24-VI-2005 GoogleMaps , H. Gallardo, 1 ♀; 8-VII-2005, M. Méndez, 1 ♀ ( ECOAB). Tuxtla Gutiérrez: 16.72786, -93.13071; 752 m, 10-VI-1991 GoogleMaps , J. Ashe, 1 ♀ ( SEMC). Tuxtla Gutiérrez, 16.75826, -93.10615; 524 m, 27-V-1984 GoogleMaps , O. Morales, 1 ♀ ( EBCH). Tzimol: 25 km SE S. Fco. Pujiltic, 16.16257, -92.26355; 590 m, 21-IV-1993 GoogleMaps , F. A. Noguera, Tecoma stans , 1 ♀; G. C. Eickwort, Tecoma stans , 1 ♀; W. E. LaBerge, 1 ♀ ( EBCH). Unión Juárez: 15.0618, -92.08; 1310 m, 28-VII-2001 , 1 ♀ ( ECOAB). Venustiano Carranza: Carretera Pujiltic-Venustiano Carranza , 16.30446, -92.53471; 637 m, 2-I-2017 GoogleMaps , P. Sagot, 1 ♀ ( ECOAB). Venustiano Carranza, 16.3166, -92.5666; 621 m, 21-IV-1993 GoogleMaps , C. Mendoza, Astianthus viminalis , 3 ♀, E. Tovar, Astianthus viminalis , 4 ♀ ( EBCH). Villaflores : 24.5 km NW, 16.35666, -93.44499; 730 m, 16-VI-1999 GoogleMaps , D. Brzoska, 1 ♀ ( SEMC). Colima: Armeria: Highway 200— Armeria R., 18.93103, -103.96384; 20 m, 22-II-1992 GoogleMaps , D. Brzoska, 1 ♀ ( SEMC). Comala: 19.34572, -103.79819; 594 m, 15-XI-2012 GoogleMaps , 10:00 h, J. Mérida, 2 ♀ ( ECOAB). Minatitlán: 33 km NW Colima , 19.40693, -103.96889; 867 m, 19-VI-1989 GoogleMaps , C. Everaert, 1 ♀ ( EBCH). Tecomán: Pascuales Malecón , 18.8494, -103.9536; 16 m, 3-X-2010 GoogleMaps , R. Ayala , 3 ♀ ( EBCH). Guerrero: Acapulco: 16.87111, -99.89777; 87 m, 6-VIII-1954 GoogleMaps , Univ. of Kansas Mex. Exped ., 8 ♀ ( SEMC). Acuitlalpán: 18.601287, -99.543012; 1584 m, 21- IX-1976 GoogleMaps , 1 ♀ ( CNIN). Benito Juárez: San Jerónimo , 17.138669, -100.470568; 39 m, 18-IV-1988 GoogleMaps , A. Cadena, 1 ♀; A. Cadena, L. Cervantes, 1 ♀ ( CNIN). Chilpancingo: 5 mi S 2.5 mi E, 17.48477, -99.44462; 1222 m, 5-VIII-1962 GoogleMaps , Univ. of Kansas Mex. Exped ., 1 ♀ ( SEMC). Chilpancingo , 5.5 mi S, 17.48020, -99.4726; 1287 m, 31-VII-1969 GoogleMaps , Univ. of Kansas Mex. Exped ., 1 ♀ ( SEMC). Copala: Entre Carr. 200 y Playa Ventura, 16.59204, -98.92507; 15 m, 25-XII-2019 GoogleMaps , 11:15 h, E. Caballero, 1 ♀; E. Gómez, 2 ♀ ( ECOAB). Playa Ventura , 16.56021, -98.97338; 4 m, 25- XII-2019 GoogleMaps , 9:00 h, E. Caballero, 1 ♀ ( ECOAB). José Azueta: Zihuatanejo , 17.64740, -101.54559; 19 m, 12-XII- 1966 GoogleMaps , G. E. Bohart, 3 ♀ ( BBSL). Zihuatanejo , 17.64546, -101.5271; 128 m, 9-VI-1984 GoogleMaps , H. Brailovsky, 1 ♂ ( CNIN). Malinaltepec: Cocoyul , 17.02440, -98.77848; 1090 m, 26-XII-2019 GoogleMaps , E. Gómez, 1 ♀; P. Sagot, 3 ♀ ( ECOAB). Cocoyul , 17.03001, -98.77689; 1074 m, 27-XII-2019 GoogleMaps , 11:00 h, P. Sagot, 1 ♀ ( ECOAB). Ometepec: Entrada sur de Huixtepec, 16.64547, -98.30962, 391 m, 24-XII-2019 GoogleMaps , 12:20 h, E. Caballero, 2 ♀ ( ECOAB). Pilcaya: 3 km SE Tonatico , 18.7641, -99.6633; 1610 m, 29-X-1991 GoogleMaps , F. A. Noguera, 1 ♀ ( EBCH). Michapa , 18.698805, -99.483348; 1190 m, 25-X-1942 GoogleMaps , B. Pelaez, 1 ♂ ( CNIN). Taxco , 23 mi N, 18.71100, -99.67296, 1341 m, 8-VIII-1954 GoogleMaps , Univ. of Kansas Mex. Exped ., 13 ♀ ( SEMC). San Luis Acatalán: Yoloxóchitl , 16.81583, -98.69027; 600 m, 8-XI-2012 GoogleMaps , Amith & Guadalupe 1 ♀; 14-XI-2012, J. Amith, 1 ♀ ( SEMC). Taxco de Alarcón: 13 km NE Taxco , 18.56685, - 99.48243; 1580 m, 29-X 1991 GoogleMaps , F. A. Noguera, 1 ♀; R. Ayala , 1 ♀ ( EBCH). Taxco 20 mi S, 18.43419, -99.55960; 1051 m, 6-II-1954 GoogleMaps , R. Dreisbach , 3 ♀ 1 ♂ ( SEMC). Taxco , 8 mi NE, 18.60451, -99.55129; 1662 m, 8-VIII-1954 GoogleMaps , Univ. of Kansas Mex. Exped ., 3 ♀ ( SEMC). Tepecoacuilco de Trujano: San Agustín Oapan, Housesite of P. Pantaleón, 17.94972, -99.43583; 516 m, 13-VIII-2009 GoogleMaps , Amith & Pantaleón, 1 ♀ ( SEMC). Jalisco: Autlán: 13 km N, camino mina San Francisco, 19.81639, 104.31684; 930 m, F. A. Noguera, A. Rodríguez GoogleMaps , 1 ♀ ( EBCH). Autlán , 6.6 km NE, 19.81869, -104.33993; 964 m, 20-VII-1994 GoogleMaps , A. Rodríguez, 1 ♀ ( EBCH). Autlán , 19.73153, -104.34541; 898 m, 15-VII-1989 GoogleMaps , R. Ayala , 1 ♀ ( EBCH). Ahuacapán , 19.68589, -104.33239; 934 m, 17-XI-2012 GoogleMaps , 11:30 h, E. Esquivel, 1 ♀ ( ECOAB). Cabo Corrientes: El Turco, km 54, carr. 200, 20.2016, -105.3429; 250 m, 21-X-2015 GoogleMaps , R. Ayala , 1 ♀ ( EBCH). Pino Suárez , 19.96128, -105.3230; 32 m, 7:55 h, J. Mérida GoogleMaps , 1 ♀ ( ECOAB). Cihuatlán: Barra de Navidad , 19.21594, -104.68423; 14 m, 10-II-1986 GoogleMaps , R. Ayala , 1 ♀ ( EBCH). El Limón: 10 km NE El Grullo , 19.84474, -104.14769; 979 m, 8-VIII-1991 GoogleMaps , F.A. Noguera, 1 ♀ ( EBCH). La Huerta: 18 km SO La Huerta , 19.34936, -104.71717; 370 m, 15-VII-1993 GoogleMaps , A. Rodríguez , F. A. Noguera , 1 ♀ ( EBCH). Careyes , 19.43766, -105.02605; 3 m, 12-II-1997 GoogleMaps , F. D. Parker, 4 ♀ ( BBSL). Chamela , 19.53000, -105.08166; 5 m, F. D. Parker, T. L. Griswold, 1-I-1985 GoogleMaps , 3 ♀; 26-IX-1985, 7 ♀ ( BBSL). Chamela, Río , 19.52789, -105.08100; 30 m, 11-VII-1989 GoogleMaps , T. L. Griswold , 1 ♀ ( BBSL). Cuiztmala Playa , 8 km S Careyes, 19.42689, -105.01340; 50 m, 4-X-1985 GoogleMaps , F. D. Parker, T. L. Griswold , 2 ♀ ( BBSL). Estación de Biología Chamela , 19.49948, -105.04242; 70 m, 8-III-1973 GoogleMaps , M. Sousa, 4 ♀ ( CNIN), S. Borrego , 1 ♀ ( EBCH), 12-V-1980 , S. H. Bullock: 1 ♀ ( EBCH), 17-V-1980 , 1 ♀; 29-V-1980, ♀ ( CNIN), 29-V-1980 , 1 ♀ ( EBCH), 21-VII-1980 , A. Pescador, 11 ♀ ( CNIN), 18-VI-1981 , S. H. Bullock: 2 ♀ ( EBCH), 11-IX-1981 , 1 ♀ ( EBCH), 4-X-1981 , 1 ♀ ( EBCH), 13-I-1982 , 2 ♀ ( EBCH), 5-V-1982 , 1 ♀ ( CNIN), 22-IX-1982 , 1 ♀ ( EBCH), 23- X-1982 , 1 ♀ ( EBCH), 17-I-1983 , 1 ♂ ( EBCH), 17-II-1983 , 1 ♀ ( EBCH), 17-IV-19833 ♀ ( EBCH), 9-VI-1983 , 2 ♀ ( CNIN), 12-VI-1983 , 1 ♀ ( EBCH), 23-X-1983 , 1 ♀ ( EBCH), 20-II-1984 , 1 ♀ ( EBCH), 5-VI-1984 , 1 ♀ ( EBCH), 5-III-1985 , R. Ayala : 1 ♂ ( EBCH), 14-III-1985 , 1 ♀ ( EBCH), 16-V-1985 , 2 ♀ ( EBCH), 8-VI-1985 , 1 ♂ ( EBCH), 1-VIII-1985 , F. D. Parker, T. L. Griswold , 1 ♀ ( EBCH), 6-VIII-1985 , R. Ayala , Prockia crucis , 1 ♀ ( EBCH), 17- VIII-1985 , R. Ayala , 1 ♀ ( EBCH). 20-IX-1985 , M. Sánchez, 1 ♀ ( EBCH), 25-IX-1985 , C. D. Michener, 1 ♀ ( EBCH), 26-IX-1985 , C. D. Michener, Apoplanesea paniculata , 2 ♀ ( EBCH), R. B. Roberts , 4 ♀ ( EBCH), F. D. Parker, T. L. Griswold , 9 ♀ ( EBCH), 26-IX-1985 , C. D. Michener, 1 ♀ ( SEMC), 27-IX-1985 , A. Rodríguez, 4 ♀ ( EBCH), 30-IX-1985 , R. Ayala , 1 ♀ ( EBCH), 1-X-1985 , C. D. Michener, 1 ♀ ( SEMC), 1-X-1985 , F. D. Parker, T. L. Griswold , 2 ♀ ( EBCH), 11-X-1985 , R. Ayala , 1 ♀ ( EBCH), 14-X-1985 , R. Ayala , 1 ♀ ( EBCH), 23-X-1985 , R. Ayala , 1 ♀ ( EBCH), 23-XI-1985 , A. Rodríguez, Bidens reptans , 1 ♀ ( EBCH), 17-XII-1985 , R. Ayala , 1 ♀ ( EBCH), 18-II-1986 , R. Ayala , 1 ♀ ( EBCH), 15-IV-1986 , F. D. Parker, 1 ♀ ( BBSL), 11-VIII-1986 , S. H. Bullock, 1 ♀ ( EBCH), 4-XI-1987 , T. L. Griswold , 2 ♀ ( BBSL), 8-IX-1988 , A. Rodríguez, 1 ♀, E. Ramírez, 1 ♀ ( EBCH), 10-VII- 1989 , T. L. Griswold , 3 ♀ ( BBSL), 6-XI-1989 , A. Cadena, 1 ♀; G. Ortega, C. Mayorga, 1 ♀ ( CNIN), 3-III-1990 , A. Chemas, 1 ♀ ( EBCH), 6-VII-1990 , G. Ortega, C. Mayorga, 1 ♀ ( CNIN), 14-X-1990 , G. Ortega, C. Mayorga, J. Martínez, S. Rodriguez, 1 ♀ ( CNIN), 25-X-1990 , G. Ortega, C. Mayorga, J. Martínez, S. Rodriguez, 1 ♀ ( CNIN), 5-VII-1991 , F. A. Noguera, Caesaria sp, 2 ♀ ( EBCH), 14-VIII-1991 , G. Ortega, C. Mayorga, 1 ♀ ( CNIN), 19-VIII- 1995 , R. Ayala , 1 ♀ ( EBCH), 13-VI-1996 , S.Borrego, 1 ♀ ( EBCH), 17-XI-2004 , M.Bateman, G.Gónzalez, 1 ♀ ( CNIN), 11-III-2000 , F. D. Parker, 1 ♀ ( BBSL), 26-IX-2016 , M.Argueta, 1 ♀ ( EBCH), 27-IX-2016 , E. López, Mimosa leptocarpa , 1 ♀ ( EBCH), 15-IX-2018 , J. Mérida, 1 ♀ 1 ♂ ( ECOAB), 8-IX-2018 , 11:30 h, J. Mérida, 1 ♀ ( ECOAB). Isla Cocinas , 19.54883, -105.10908; 45 m, 29-XI-1983 GoogleMaps , S. H. Bullock, 1 ♀ ( EBCH). Playa Careyes, nr Chamela , 19.44282, -105.02328; 16 m, 30-IX-1985 GoogleMaps , C. D. Michener, 3 ♀ ( SEMC), 30-IX-1985 , C. D. Michener, 1 ♀ ( EBCH). Playa Careyitos , 19.43766, -105.02605; 2m, R. B. Roberts GoogleMaps , 2 ♀; R. Ayala , 3 ♀ ( EBCH). Río Cuixmala , 19.41055, -104.95638; 46 m, 27-II-1991 GoogleMaps , E. Ramírez, 2 ♀, J. Villa, 1 ♀ ( EBCH). Río San Nicolás , 25 km N Chamela, 19.65947, -105.18005; 21 m, 5-XI-1987 GoogleMaps , T. L. Griswold , 1 ♀ ( BBSL). Río San Nicolás , 19.651, -105.181; 12 m, 10-XI-1985 GoogleMaps , M. Sánchez, 1 ♀; 18-XI-1985, R. Ayala , 3 ♀; 9-XII-1985, M. Sánchez, 1 ♀ ( EBCH). Magdalena : 20.92371, -103.99493; 1454 m, 17-VI-1988 GoogleMaps , R. Minckley , 1 ♂ ( SEMC). Pihuamo: 12 km N de Pihuamo , 19.32949, -103.38794; 760, 30-X-1992 GoogleMaps , R. Ayala , 1 ♀ ( EBCH). Puerto Vallarta: 20.62141, -105.22601; 14 m, G. E. Bohart: 25-I-1984 GoogleMaps , 16 ♀; 8-XII-1984, 6 ♀ ( BBSL), 29-IX-1985 , C. D. Michener, 1 ♀ ( SEMC), 29-IX-1985 , C. D. Michen- er, 1 ♀ ( EBCH). Tequila: Volcán de Tequila , 20.86341, -103.84396; 1300 m, 4-X-1995 GoogleMaps , H. Fierros, 3 ♀ ( EBCH). Tomatlán: José María Morelos , 19.6667, 105.1784; 26 m, 19-VII-1989 GoogleMaps , S. Anaya, 1 ♀ ( EBCH). José María Morelos 6 km N, 19.71976, -105.21425; 22 m, 19-VII-1989 GoogleMaps , C. Cordero, 1 ♀; S. Anaya, 2 ♀ ( EBCH). Tomatlán , 2 km N, 19.95696, -105.24828; 93 m, 19-VII-1989 GoogleMaps , C. Cordero, 1 ♀ ( EBCH). Tonaya: 22 km E El Grullo , 19.80784, - 104.00546; 880 m, 17-VII-1989 GoogleMaps , L. Godínez, Acacia angustissima, 1 ♀ ( EBCH). Zapotlán el Grande: Los Depósitos , 19.65319, -103.53162; 1721 m, 16-XI-2012 GoogleMaps , 9:40 h, J. Mérida, 1 ♀ ( ECOAB). Michoacán: Ario: 9 km N La Huacana , 19.04395, -101.80853; 1098 m, 4-XI-1992 GoogleMaps , A. Rodríguez, 1 ♀; R. Ayala , 2 ♀ ( EBCH). 17 km SO Ario de Rosales , 19.11145, -101.73965; 1470 m, 3-XI-1992 GoogleMaps , R. Ayala , 2 ♀ ( EBCH). El Cangrejo, 20 km N La Huacana, 19.04166, -101.81138; 1090 m, 30-X-1987 GoogleMaps , 3 ♀ ( EBCH). Morelia: 20 mi E, 23 mi S, 28-VII-1962 , Univ. of Kansas Mex. Exped ., 1 ♀ ( SEMC). Arteaga: 18.34803, -102.29194; 830 m, 8-V-1989 GoogleMaps , R. Aguirra , 1 ♀ ( SEMC). Arteaga , 3 km N, 18.38394, -102.29201; 984 m, 1-XI-1987 GoogleMaps , T. L. Griswold , 3 ♀ ( BBSL). La Lajita, 26 km N Arteaga, 18.40935, -102.14797; 660 m, 1-XI-1987 GoogleMaps , L. Godínez, 1 ♀ ( SEMC), 1-XI-1987 , R. Ayala , 1 ♀ ( EBCH). Buenavista: 10 km W Apatzingán, La Huina , 19.10813, -102.52083; 256 m, 17-VII-1989 GoogleMaps , V. R. Castrejón, Chilopsis linearis , 2 ♀ ( EBCH). Gabriel Zamora: 20 km S Uruapán , 19.25057, -102.05532; 944 m, 7-II-1990 GoogleMaps , S. H. Bullock, 1 ♀ ( EBCH). Gabriel Zamora, 19.0948, -102.05795; 522, 4-X-2011 GoogleMaps , R. Ayala , 2 ♀ ( EBCH). Lombardía , 19.15668, - 102.05352; 656 m, 5-X-2011 GoogleMaps , R. Ayala , 1 ♀ ( EBCH). Jungapeo: 19.49119, -100.48779; 1584, -5-1953, O. Hecht GoogleMaps , 2 ♀ ( BBSL). La Huacana: 2 km NE Zicuirán , 18.89107, -101.95805; 248 m, 10-VII-1988 GoogleMaps , G. Rodríguez, 2 ♀ ( EBCH). El Descansadero, Arteaga , 80 km N, 18.68201, -102.00509; 266 m, 1-XI-1987 GoogleMaps , L. Godínez, 1 ♀ ( SEMC). Zicuirán , 18.87756, -101.96943; 200 m, 8-VII-1988 GoogleMaps , G. Rodríguez, 3 ♀ ( EBCH). Lázaro Cárdenas: La Manzanilla, 18.06361, -102.69083; 11 m, 7-VII-1988 GoogleMaps , G. Rodríguez, 1 ♀ ( EBCH). Madero: Los Fresnos , 19.48527, - 101.05722; 2132 m, 2-XII-2014 GoogleMaps , 1 ♂ ( ECOAB). Múgica: Cuatro Caminos , 29.8 km S, on hwy 37, 18.98805, - 102.10305; 358 m, 18-VII-1989 GoogleMaps , 1 ♀ ( SEMC). Tacámbaro: Joyas Altas , 19.21441, -101.42350; 1403 m, 20-XI-2018 GoogleMaps , A. Reyes, 1 ♀ ( ECOAB). Morelos: Cuernavaca: 18.93481, -99.23082; 1561 m, F. D. Parker: 8-XI-1987 GoogleMaps , 1 ♀; 13- XI-1987, 1 ♀; 19-XI-1987, 1 ♀ ( BBSL). Tepoztlán: Cuernavaca , 6 mi E, 18.93414, -99.13841; 1472 m, 12-IX- 1973 GoogleMaps , W. J. Hanson, B. A. Haws, 1 ♀ ( BBSL). Tepoztlán , 6 mi W, 18.97416, -99.16429; 1647 m, 30-VII-1963 GoogleMaps , M. Naumann, 1 ♀ ( SEMC). Tepoztlán , 18.9848, -99.09305; 1673 m, 30-X-1983 GoogleMaps , O. Morales, 1 ♀ ( EBCH). Tlaquiltenango: Huautla , 2.5 km N, 4 km W, Estación CEAMISH, 18.46120, -99.04130; 940 m, 7-IX-1996 GoogleMaps , R. Brooks , 1 ♀ ( SEMC). Huaxtla , 18.393314, -99.05025; 1160 m, 9-IX-2015 GoogleMaps , I. Hinojosa, 1 ♀ ( CNIN). Yautepec: Cuernavaca , 12 mi E, 18.91371, -99.04437; 1237 m, 14-VIII-1954 GoogleMaps , Univ. of Kansas Mex. Exped ., 1 ♂ ( SEMC). Oaxtepec , 18.90637, -98.97026; 1379 m, 22-V-1989 GoogleMaps , A. Rodríguez , F. A. Noguera , 4 ♀ ( EBCH). Nayarit: Bahía de Banderas: Punta Mita , 3 km S, 20.76028, -105.47487; 22 m, 28-II-1992 GoogleMaps , D. Brzoska, 2 ♀ ( SEMC). San Blas: San Blas 20 km S, 21.43396, -105.18722; 10 m, 7-IV-1994 GoogleMaps , A. Rodríguez, 1 ♀ ( EBCH). Tecuala: Acaponeta , 16 mi S, 22.28710, -105.27176; 45 m, 12-VIII-1963 GoogleMaps , M. Naumann, 1 ♀ ( SEMC). Oaxaca: Ometepec: 11 km N, 16.76985, -98.40548; 708 m, 4-IV-1989 GoogleMaps , R. Ayala , Vitex sp , 1 ♀ ( EBCH). Putla Villa de Guerrero: San Marcos Malpica, 16.99897, -97.9; 744 m, 6-XI-2015 , N. Arnold, 1 ♀ ( ECOAB). Salina Cruz : 16.18608, -95.19235; 93 m, 7-IX- 1965 GoogleMaps , D. Janzen, 18 ♀ ( SEMC). San Dionisio del Mar: 16.33708, -94.76381; 7 m, 20-I-1992 GoogleMaps , L. Godínez, 2 ♀ ( SEMC). San Juan Colorado: Nuevo Progreso, 16.52503, -97.8862; 566 m, 10-XI-2015 GoogleMaps , 12:00 h, N. Arnold, 1 ♀ ( ECOAB). San Miguel Tlacamama: 16.41474, 98.09691; 235 m, 9-XI-2015 GoogleMaps , 13:00 h, N. Arnold, 1 ♀; M. López, 1 ♀ ( ECOAB). San Pedro Comitancillo: 16.45254, -95.20089; 112 m, 15-XI-2018 GoogleMaps , 9:20 h, P. Sagot, 1 ♀ (ECO- AB). San Pedro Huamelula: Playa Grande, 15.85964, -95.90502; 97 m, E. Gómez GoogleMaps , 1 ♀ ( ECOAB). San Pedro Mixtepec: 15.9013, -97.1454; 14 m, 18-XII-2018 GoogleMaps , 9:30 h, I. Pérez, 3 ♀; P. Sagot, 1 ♀ ( ECOAB). Puerto Escondido , 15.87646, -97.10498; 10 m, 30-IX-1990 GoogleMaps , L. Godínez, 1 ♀ ( SEMC). Puerto Escondido 10 km NW, 15.92442, - 97.14531; 12 m, 6-III-1989 GoogleMaps , R. Ayala , 1 ♂ ( EBCH). San Pedro Pochutla: Puerto Ángel , 15.6643, -96.4883; 20 m, 9-III-1989 GoogleMaps , R. Ayala , 2 ♀ ( EBCH). San Pedro Tututepec: Faisán , 16.14982, -97.72474; 56 m, 22-XII-2019 GoogleMaps , E. Gómez, 1 ♀ ( ECOAB). Laguna Chacahua , 15.96808, -97.68804; 6 m, 20-III-1984 GoogleMaps , J. Pérez, 1 ♀ ( EBCH). Santa María Colotepec: Juan Diego , 15.8233, -96.9078; 119 m, 19-XII-2018 GoogleMaps , 13:20 h, P. Sagot, 1 ♀ ( ECOAB). Santa María Colotepec, 15.8633, -97.0045; 24 m, 19-XII-2018 GoogleMaps , 8:45 h, I. Pérez, 1 ♀ ( ECOAB). Santa María Colotepec, 15.8490, -97.0190; 59 m, 19-XII-2018 GoogleMaps , 10:35 h, I. Pérez, 1 ♀; P. Sagot, 2 ♀ ( ECOAB). Santa María Colotepec, 15.80027, -96.98567, 6 m, 19-XII-2018 GoogleMaps , 11:00 h, P. Sagot, 1 ♀ ( ECOAB). Santa María Huatulco: Coyula , 15.68949, -96.30446; 11 m, 11-XII-2014 GoogleMaps , 9:00 h, I. Pérez, 1 ♀ ( ECOAB). Santa María Tonameca: carr. a Loxicha, 15.80540, -96.76030; 155 m, 20-XII-2018 GoogleMaps , 9:25 h, O. Caballero, 1 ♀ ( ECOAB). San Agustinillo bosque, 15.67065, -96.53800; 30 m, 29-I-2012 GoogleMaps , 11:00 h, P. Sagot, 7 ♀; 9-I-2016, 9:00 h, P. Sagot, 3 ♀ ( ECOAB). San Agustinillo Carretera , 15.66679, -96.54062; 19 m, 28-I-2012 GoogleMaps , 10:30 h, P. Sagot, 8 ♀ ( ECOAB). Santa María Zoquitlán: Las Casas , 16.63029, -96.45168; 1435 m, 21-XI-2018 GoogleMaps , 9:30 h, I. Pérez, 1 ♀ ( ECOAB). Santiago Niltepec : Microondas Los Tules , Niltepec, 16.5, -94.58; 44 m, 13-XII-1985 , R. Torres , 1 ♀ ( EBCH). Santo Domingo Tehuantepec: Tehuantepec , 17 km W, 16.34649, -95.39159; 176 m, 8-IX-1965 GoogleMaps , D. Janzen, 21 ♀ ( SEMC). Villa Sola de Vega: Carr. 131 a la costa, 16.43087, -97.06787; 1160 m, 14-XII-2018 GoogleMaps , 14:25 h, N. Arnold, 1 ♀ ( ECOAB). Puebla: Petlalcingo: 1 km SW Petlalcingo , 18.08047, -97.92819; 1280 m, 1-XI-1991 GoogleMaps , A. Rodríguez, 1 ♀ ( EBCH). Tepexco: 14 km NW Izúcar de Matamoros , 18.63578, 98.58635; 1306 m, 31-X-1991 GoogleMaps , F. A. Noguera, 1 ♀ ( EBCH). Tepexco , 4 km NW Tepexco, 18.663, -98.72239; 1273 m, 31-X-1991 GoogleMaps , T. L. Griswold , 1 ♀ ( EBCH). Sinaloa: Concordia: Copala , 23.39819, -105.93727; 548 m, 12-X-2013 GoogleMaps , 10:30 h, J. Mérida, 1 ♀ ( ECOAB). Copala , 16 km E, 23.43870, - 105.84376; 1168 m, 18-III-1990 GoogleMaps , R. Minckley , 2 ♀ ( SEMC). Mazatlán: 23.49106, -106.48155; 27 m, 29-X-2014 GoogleMaps , 9:50 h, J. Mérida, 1 ♀ ( ECOAB). Tabasco: Balancán: Capulín , 17.70578, -91.09616; 38 m, 10-V-2014 GoogleMaps , L. Marín, 1 ♀ ( ECOAB). NICARAGUA: Rivas: Cárdenas: Playa Isla Vista, 11.23401, -85.55012; 33 m, 29-I-2016 GoogleMaps , R. Joyce , 1 ♀, 26-IX-2016, M. Calero, 1 ♀ ( ECOAB). San Juan del Sur: Hacienda La Flor , 11.14192, -85.78289; 16 m, M. Calero: 28 I-2016 GoogleMaps , 5 ♀, 17-II-2016, 1 ♀ ( ECOAB). La Flor / Hda La Flor , 11.14059, -85.78459; 7 m, 18-V- 2016 GoogleMaps , M. Calero, 1 ♀ ( ECOAB). Las Parcelas , 11.13064, -85.47305; 24 m, 16-V-2916 GoogleMaps , M. Calero, 1 ♀ ( ECOAB). Río Ostional , 11.12598, -85.74979; 22 m, 8-II-2016 GoogleMaps , R. Joyce , 1 ♀; 11-II-2016, M. Calero, 1 ♀ ( ECOAB). Tortuga , 11.14057, -85.78368; 12 m, 11-VI-2015 GoogleMaps , E. Dorgay, 1 ♀; 4-VII-2015, E. Dorgay, 1 ♀ ( ECOAB) .

Commentaries. X. gualanensis was described as a subspecies of X. wilmattae by Cockerell in 1912. This author did not mention number of specimens analyzed, as Moure (2012) points out in his catalogue; in this last work the type is mentioned to be in the AMNH collection, however it is currently at the USNM on loan.

BBSL

USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research

MNCR

Museo Nacional de Costa Rica

SEMC

University of Kansas - Biodiversity Institute

R

Departamento de Geologia, Universidad de Chile

T

Tavera, Department of Geology and Geophysics

V

Royal British Columbia Museum - Herbarium

CNIN

Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico

AMNH

American Museum of Natural History

USNM

Smithsonian Institution, National Museum of Natural History

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Hymenoptera

Family

Apidae

Genus

Xylocopa

Loc

Xylocopa (Neoxylocopa) gualanensis Cockerell, 1912

Mérida-Rivas, Jorge A., Hinojosa-Díaz, Ismael A., Ayala-Barajas, Ricardo, Pozo, Carmen & Vandame, Rémy 2022
2022
Loc

Xylocopa wilmattae gualanensis

Cockerell, T. D. A. 1912: 555
1912