Pseudoplatylabus violentus (GRAVENHORST)

M, Alexander, 2015, Ichneumon flies of the tribe Joppocryptini of Palaearctic. Description of new genus Londokia nov. gen. and two new species L. kasparyani nov. sp. and L. leleji nov. sp. (Hymenoptera, Ichneumonidae, Ichneumoninae), Linzer biologische Beiträge 47 (2), pp. 1865-1884 : 1869-1871

Pseudoplatylabus violentus (GRAVENHORST) (Plate 1)

Ichneumon violentus: GRAVENHORST 1829 - Ichn. Eur. 1: 613,.

Hoplismenus infaustus: WESMAEL 1844 - Nouv. Mem. Acad. Sci. Brux. 18: 110,,.

Pseudoplatylabus caudatus: SMITS van BURGST 1931 - Konowia 10: 30.

Pseudoplatylabus violentus: HEINRICH 1936 - Polskie Pismo Entomol. 13 (1934): 25.

Pseudoplatylabus violentus: HEINRICH 1962 - Canad. Ent. Suppl. 27: 685-686,,.

Pseudoplatylabus violentus: RASNITSYN 1981 - Opr. Faun. SSSR 3 (3): 562,,.

M a t e r i a l e x a m i n e d: Byelorussia: Beresina Nat. Res.: Domzhericy , 54°46'N- 28°16'E, meadow dry, Malaise trap, 29.06.198 9, 1, A. Tereshkin leg. ; Postrezh'e , 54°38'54.74"N- 28°20'44.34"E, Pinetum pleuroziosum, Malaise trap: 02.06.198 9, 1 ; 29.06.198 9, 5; 15.07.198 9, 1; 15.05.199 0, 2; 13.07.199 0, 4; 15.06.199 0, 1,1.; 54°38'45.90"N- 28°21'39.16"E, 29.06.199 0, 1; 54°38'N- 28°21'E, 26.09.199 1, 1, A. Tereshkin leg.; Postrezh'e , 54°38’N- 28°21’E, Pinetum sphagnosum, 25.05.199 3, 1, A. Tereshkin leg. ; Krajcy, 54°39'N- 28°16'E, Piceetum, net, 03.09.198 3, A. Tereshkin leg. Minsk, Krupki, Osecheno , 54°36’N- 29°17’E, meadow wet, Malaise trap, 02.09.198 9, 1, A. Tereshkin leg. ; Brest, Baranovichi, Molchad' , 53°18’N- 25°45’E, personal plot, Malaise trap, 07.06.198 8, 1, A. Tereshkin leg. ; Soligorsk, Dubei , personal plot, Malaise trap, 27.07.198 9, 1, A. Tereshkin leg. ; Pripiat Nat. Res., pos. Khvoensk : 52° 2'11.56"N- 27°56'43.14"E, meadow, Malaise trap: 09.06.198 7, 1, 07.07.198 7, 2 ; 52°02’N- 27°57’E, Alnetum urticosum, net, 16.05.198 7, A. Tereshkin leg.; Khojniki, Polesskij zap., ur. Dron'ki : 51°44’N- 29°52’E, Alnetum urticosum, Malaise trap: 30.05.198 9, 1, A. Tereshkin leg. ; personal plot, Malaise trap: 04.08.199 2, 1; 25.05.199 3, 3; 25.06.199 3, 1,2; 03.08.199 3, 6,4; 24.08.199 3, 1,6; 24.08.199 3, 1; 07.10.199 3, 2; 29.06.199 4, 1,4; 03.08.199 5, 2; 22.09.199 5, 3,1; 24.07.199 6, 1,1; 20.06.199 7, 6; 20.07.199 8, 4; Piceetum pleuroziosum: 09.07.199 2, 1 ; 08.09.199 2, 1; ur. Krasnosel'e : 51°33’N- 29°53’E, personal plot, Malaise trap: 09.07.199 2, 1 ; 25.06.199 3, 5; ur. Orevichi: 51°35’N- 29°50’E, 05.10.199 4, Querceetum flood plain, 1,1, A. Tereshkin leg. Russia: SW Caucasus, Krasnaja Poljana, Mzymta riv. val., 43°40'N- 40°12'E, personal plot, Malaise trap, 30.07.198 7, 2, A. Tereshkin leg. ; Primorskij Kraj: Kamenushka at Ussurijsk , 27.07.199 2, 1 ; Kajmanovka : 27.07.199 2, 1 ; 31.07.199 2, 1; 01.08.199 2, 1 A. Tereshkin leg.

F e m a l e

F l a g e l l u m: Slender, bristle-shaped, only just widened, with 32-33 segments, first segment moderately long, 3,3 times longer than width at apex, rust-colored with more darkened apex and white semiannulus on segments (8)9-12, flattened ventrally but not widened behind middle (only segment 20 square from lateral). Flagellum equal by length to front wing and 1,6 times shorter than body length.

H e a d: Head contour from front strongly narrowed downwards, transversal, 1,2 times wider than height; eyes large, temples from front visible only to 1/8 of lower part of an eye, genae long, from front 3,3 times shorter than height of an eye; head from above transversal, 2,2 times wider than length in middle. Vertex from lateral behind hind ocelli smoothly, almost straightly slanting down to occipital carina; temples long, 1,4 times longer than longitudinal diameter of an eye in the middle, from above roundly narrowed behind eyes, from side parallel to hind margin of an eye; occipital carina from above deeply but roundly impressed but not reach level of eyes and hind ocelli, sharp all round, meeting with hypostomal carina not reach of mandible base; hypostomal carina visible from lateral; length of abscissula 3,5 lesser than mandible base width; sulcus genalis broadly impressed; mandibles narrow, sickle-shaped, smoothly and evenly curved, upper tooth long, lower only just developed, but distinct, sharp, strongly moved inside in comparison with upper tooth, not visible from front; clypeus broad, convex at base and strongly impressed and flattened to apex, with slightly emarginate almost straight front margin and with raised and roundly oblique lateral margins, only just separated from face by broad impression, with slight longitudinal wrinkles along front margin; clypeal foveae deep, rounded, impression around them slightly expressed; labrum short, rounded, equal by breadth to front margin of clypeus; middle field of face only just elevated above lateral fields and not separated by distinct impressions, broad, 1,5 times broadly then lateral fields in middle; antennal cavities not great, very deep, laterally reach borders of eyes, but far not reach level of front ocellus, with slight lateral tubercles and expressed tooth between antennal fossae; front margins of antennal fossae slightly elevated above face surface; ocelli of normal size, diameter of lateral ocellus 1,8 times less than distance from ocellus to an eye; ocellar triangle distinctly elevated. Upper part of clypeus punctured by big connivent smoothed punctures, apical third smooth; surface of face roughly wrinklypunctured to cellular in upper third; antennal cavities with transversal wrinkling in upper half to entirely smooth; frons roughly wrinkly-punctured by irregular punctures; vertex and temples smoothly wrinkly-punctured; surface of head without microsculpture or with only just wisible microsculpture, more over shining.

T h o r a x: Collar of pronotum long with straight front margin, transverse furrow of pronotum broad, shallow, interrupted by low broad keel; pronotal base gradually uniformly curved almost with raight apical angle; epomiae strong, pronotal ridge not thickened; pronotum surface wrinkled in central part and densely punctured by punctures with expressed angles (reticulate-wrinkled) in upper one. Mesonotum moderately convex, only just longer than breadth; notauli only just marked in front third (middle field slightly elevated); surface very densely punctured by connivent punctures, dull as at all representatives of the tribe, interspaces between punctures not granulated; prepectus behind front coxae wrikly-punctured, slightly shining; prepectal carina apically not reach margin of pronotum, from below curved in area of sternauli; subalarum thin, high, sharp; surface of speculum roughly wrikly-punctured as other part of mesopleurae; mesopleural fovea small, slightly impressed, longutudinal; lower part of mesopleurae (mesosternum) not separated angularly with gradual bend; sternauli in a form of distinct impressions up to middle; mesopleurae including speculum (with exception of small shining stripe above mesopleural fovea) densely wrinkly-punctured; surface of mesopleurae matt; scutellum high elevated above postscutellum, bordered laterally up to apex by high carinae, horizontal part of scutellum convex, densely punctured by big punctures, matt, dull, vertical surface and postscutellum with sharp longitudinal wrinkles. Hind margin of metanotum with triangle projections opposite lateral longitudinal carinae. Propodeum from lateral of broken type, but with convex horizontal part, its horizontal part 1,4 times shorter than area posteromedia in middle; all carinae of propodeum sharp; basal area short, convex; area superomedia elongated, hexagonal, narrowed from costulae more strongly forward and slightly backwards, carina limited area superomedia in front roundly convex anteriad, costulae before middle of area superomedia; areae dentiparae at apices with short denticles, carina closing area dentipara at apex broadened; spiracles long, 3 times longer than width. Horizontal and vertical surface of propodeum densely wrinkly-punctured by big irregular punctures (to almost cellular on 2nd lateral field), with slight shine; metapleurae very roughly wrinkly-punctured, dull.

L e g s: Very long and slender; hind coxae of female without traces of scopa with dense puncturation; claws smooth with strongly curved apex.

W i n g s: Areolet pentagonal, symmetrical (wide of base varies considerably); stigma narrow, pale brown; radius slightly sinuous in a most part, curved at apex; nervulus intersticial, ramulus from distinct to only just marked; veins of both of wings light, apical veins strongly unsclerotized, apical veins of hind wings in a form of trace; membrane of wings hyaline. Length of front wing 1,5 times shorter than body length.

A b d o m e n: From above narrow, fusiform, with strongly projecting sheath of ovipositor, sharply oxypygous; tergite 2 from above 1,1 times shorter than width at apex; sheath of ovipositor from above far protrude behind apex of abdomen, more than length of tergites 6-7. First tergite from lateral with gradual bend of petiolus to postpetiolus; petiolus carinated only ventrally, lateral surface of petiolus sculpturated by fine transversal ribs, shining; petiolus narrow, from above gradually broadened to postpetiolus; middle field of postpetiolus distinctly elevated, not carinated, wider than lateral fields; middle field of postpetiolus usually smooth (at some species from Russian Far East to superficially irregularly-wrinkled), lateral fields with big smoothed punctures, surface of postpetiolus shining. Gastrocoeli deeply impressed, short and slightly oblique; thyridia distinct, 1,8 times wider than interval between them; lunulae distinct, of middle size, situated behind middle of 2nd tergite; surface of tergites 2-4 densely punctured by small superficial punctures, shining, other tergites smooth, shining. Sternites 2-4 completely unsclerotized, 5th in a most part. Hypopygium with fold.

C o l o r a t i o n: Head and thorax entirely black; tergites 1-3 and 4 up to middle red, tergites 5-7 black, 6-7 with big white spots; Legs predominately red; front and middle coxae predominately black to red, hind one predominately red, darkened in different degree, less often black; trochanteres 1 of front and middle legs black, hind femora and tibiae at apex and tarsi darkened; front tibiae yellow in front.

S i z e: Body length – 8,5-9,5; front wing – 5,2-6,0; flagellum – 5,2-6,2 mm.

V a r i a b i l i t y Tergite 4 sometimes entirely red, more often red up to middle. Coxae of all the legs more often red in a varying degree, less often entirely black.

M a l e

F l a g e l l u m: Slender, bristle-shaped, 31-32 segments, reddish-brown ventrally with traces of white annulus on segments 13-14(15) dorsally; flagellum sharply ribbed with transverse, bristle-topped carinae from 2nd segment; tyloides (Plate. 1,8) roundish, indistinct, on segments (9)10-13(14) to entirely their absence at some speciments.

H e a d: As at female.

T h o r a x: Similar by morphology with female. Sexual dimorphism is revealed in more rough sculpture of horizontal part of scutellum to roughly-wrinkled at some samples and considerably variation of hexagonal shape of area superomedia, from narrowed in front from costulae to broadened.

L e g s: More stout than at females, segments 3-4 of tarsi with white pattern dorsally.

W i n g s: Unlike females apical veins both of wings sclerotized normally.

A b d o m e n: First tergite with black base to black entirely, tergites 2-3, sometimes only tergite 2 (samples from Russian Far East) red, tergites 4-5 enterely black, less often base of tergite 4 red; only tergite 7 in a most part and paramerae white.