Sogana Matsumura, 1914

Constant, Jérôme & Pham, Thai Hong, 2020, Sogana cysana sp. nov., a new tropiduchid planthopper from Chu Yang Sin National Park in Vietnam and key to Vietnamese species of the genus (Hemiptera: Fulgoromorpha: Tropiduchidae), Belgian Journal of Entomology 95, pp. 1-13 : 1-13

Genus Sogana Matsumura, 1914

Sogana MATSUMURA, 1914: 268 .

Type species: Sogana hopponis Matsumura, 1914 , by monotypy.

Sogana – MELICHAR, 1914: 194; DISTANT, 1916: 54; METCALF, 1954: 130; TSAUR, 1990: 245; LIANG & WANG, 2008: 30; CONSTANT, 2010: 64; CONSTANT & PHAM, 2013: 72; CONSTANT, 2019: 4; CONSTANT & PHAM, 2019: 4.

FLOW, 2020:

https://www.hemiptera-bases.org/flow/?page=explorer&db=flow&lang=en&card=taxon&rank=genus&id=8083

SPECIES INCLUDED (TYPE LOCALITY):

Sogana bachmana Constant & Pham, 2019 ( Vietnam);

Sogana baviana Constant & Pham, 2019 ( Vietnam);

Sogana chartieri Constant, 2019 ( Cambodia);

Sogana clara Liang & Wang, 2008 ( Laos);

Sogana condaoana Constant & Pham, 2013 ( Vietnam);

Sogana cucphuongana Constant & Pham, 2013 ( Vietnam);

Sogana cysana sp. nov. ( Vietnam);

Sogana extrema Melichar, 1914 ( Myanmar);

Sogana floreni Constant, 2010 (Borneo);

Sogana hopponis Matsumura, 1914 (Taiwan);

Sogana longiceps Fennah, 1978 ( Vietnam);

Sogana pseudohopponis Liang & Wang, 2008 ( Laos);

Sogana robustocarina Liang & Wang, 2008 (Borneo);

Sogana stimulata Melichar, 1914 (Mentawai Islands).

Identification key to the species of Sogana from Vietnam

1. Vertex more or less subquadrate ( Fig. 1 B, D ), ratio LV/BV <0.85; apical margin of anal tube roundly notched apically in dorsal view ( Fig. 1 N, O ); aedeagus with strong, deeply furcate or strongly dentate process on left side ( Fig. 1 S–T , 3 F )....................................... 2

– Vertex elongate ( Fig. 1 A, C, E ), ratio LV/BV> 0.85; apical margin of anal tube rounded or oblique apically in dorsal view, not notched ( Fig. 1 K–M ); aedeagus without strong furcate process ( Fig. 1 P–R )................................................................................................ 4

2. Tegmina more elongate, LTg/BTg = 3.8, with 18–19 apical cells ( Fig. 1 I ); vertex more elongate, LV/BV = 0.8 ( Fig. 1 B ); ventral process of periandrium ended in a hook ( Fig. 1 X )................................................................................ S. baviana Constant & Pham, 2019

– Tegmina less elongate, LTg/BTg = 3.3–3.5, with 13 ( Fig. 2 E ) or 19–21 apical cells ( Fig. 1 J ); vertex shorter, LV/BV = 0.63–0.7 ( Figs. 1 D , 2 G ); ventral process of periandrium not ended in a hook ( Fig. 1 Y ) .................................................................................................. 3

3. Size smaller, ♂ LT = 8.8 mm; tegmina more elongate, LTg/BTg = 3.5, with 13 apical cells ( Fig. 2 E ); vertex slightly more elongate, LV/BV = 0.7 ( Fig. 2 G ); right dorsal process of periandrium not furcate ( Fig. 3 E, H ); apical projections of anal tube pointed ( Fig. 3 A–C ) .................................................................................................................. S. cysana sp. nov.

– Size larger, ♂ LT = 13 mm; tegmina less elongate, LTg/BTg = 3.3, with 19–21 apical cells ( Fig. 1 J ); vertex shorter, LV/BV = 0.63 ( Fig. 1 D ); right dorsal process of periandrium furcate ( Fig. 1 T ); apical projections of anal tube not pointed ( Fig. 1 O ) ............................. ......................................................................... S. cucphuongana Constant & Pham, 2013

4. Vertex more elongate, ratio LV/BV = 1.15 ( Fig. 1 E ); aedeagus with two elongate dorsal processes ending in a single point ( Fig. 1 R, W ) ....................... S. longiceps Fennah, 1978

– Vertex less elongate, ratio LV/BV = 0.9 ( Fig. 1 A, C ); aedeagus with dorsal processes with multiple teeth or crenulations apically ( Fig. 1 P–Q ) ........................................................... 5

5. Tegmina more elongate, LTg/BTg = 3.6 ( Fig. 1 F ); tegmina with 14 apical cells ( Fig. 1 F ); clypeus mostly pale yellow-brown; dorsal processes of aedeagus with left one strongly sinuate and directed anteroventrally and right one spatulate apically and directed mostly dorsally ( Fig. 1 P )................................................... S. bachmana Constant & Pham, 2019

– Tegmina less elongate, LTg/BTg = 3.0 ( Fig. 1 G ); tegmina with 18 apical cells ( Fig. 1 G ); clypeus mostly pale brown; dorsal processes of aedeagus both nearly straight and directed anterodorsally ( Fig. 1 Q ) ....................................... S. condaoana Constant & Pham, 2013

Figures

Fig. 1. Sogana spp., diagnostic characters. A–E, head and thorax, dorsal view. F–J, right tegmen. K–O, anal tube, dorsal view. P–T, aedeagus, left lateral view. U–Y, aedeagus, ventral view. S. bachmana: A, F, K, P, U; S. baviana: B, I, N, S, X; S. condaoana: C, G, L, Q, V; S. cucphuongana: D, J, O, T, Y; S. longiceps: E, H, M, R, W.

Fig. 3. Sogana cysana sp. nov., holotype ♂. A, pygofer, anal tube and gonostyli, left lateral view. B, pygofer, anal tube and gonostyli, right lateral view. C, pygofer, anal tube and gonostyli, dorsal view. D, left gonostylus, dorsal view. E, aedeagus, right lateral view. F, aedeagus, left lateral view. G, aedeagus, left laterodorsal view. H, aedeagus, dorsal view. I, aedeagus, left laterodorsal view. J, aedeagus, ventral view. An: anal tube – apa: apical process of aedeagus– cn: connective – dpa: dorsal process of aedeagus – esp: ear-shaped process of aedeagus – G: gonostylus – ldp: left dorsal process of aedeagus – pe: periandrium – Py: pygofer – vpp: ventral process of periandrium.

Fig. 2. Sogana cysana sp. nov., holotype ♂ (RBINS). A, habitus dorsal view. B, habitus, ventral view. C, habitus, perpendicular view of frons. D, habitus, lateral view. E, right tegmen. F, right posterior wing. G, head and thorax, dorsal view. H, head and thorax, laterodorsal view. I, head and thorax, lateral view. J, frons, perpendicular view. K, distal portion of left posterior leg, ventral view.