Compsocommosis Heppner & Bae

Heppner, John B. & Bae, Yang-Seop, 2015, Compsocommosis new genus, with a new species in Vietnam, and Transfer of Mictocommosis to Archipini (Lepidoptera: Tortricidae: Tortricinae: Archipini), Zootaxa 3999 (1), pp. 144-150 : 145-147

Compsocommosis Heppner & Bae , new genus

Figs. 1–5

Type-species: COmpSOcOmmOSiS vietnamenSiS Heppner & Bae , new species

Diagnosis. This new genus is related to the Asian Mictocommosis but differs significantly in genital characters, with the female having the sterigma split posteriorly into two plates, unique sclerotized and dentate plates at the ductus bursae-bursal border, and a single small thorn-like signum. The genitalia characters are very different from those of Mictocommosis : females have distinctive sterigma-ostium and ductus plates, both of which are simplified in Mictocommosis , and ductus plates are lacking in the latter genus. The forewing has only 2 apical silver lines (three in Nexosa ). The male is unknown.

Description. Adults ( Fig. 1 ) small (ca. 14 mm). Head ( Figs. 2–3 ): Normal tortricoid form; ocellus moderate; labial palpus small (slightly larger than eye), somewhat upturned; antenna filiform, unmodified. Thorax: Unmodified; legs unmodified. Forewing ( Fig. 4 ) elongated, with sharp apex and sinuate termen and rounded tornus, R1–R4 to costa, R5 to termen, M1–M3 equally spaced and M3 approximate with CuA1 at end of cell, CuA2 parallel to CuA1, CuP strong to base, and A1+2 with long basal fork and long to near tornus; hindwing subtriangular with sharply rounded apex and convex costal margin, with tornus merged to rounded anal margin, Sc+R1 to costa at apical quarter, Rs to just above apex, M1–M3 nearly subequal (M1 more distant from M2 than M3 is from M2), M3 and CuA1 approximate at end of cell, CuA2 to tornus and distant from CuA1, CuP strong to midwing, A1+2 with short basal fork, and A3 evident near anal margin. Abdomen. Unmodified. Female genitalia ( Fig. 6 ) elongate, with short ovipositor, papilla anales tortricoid, setous and flattened; lamella postvaginalis split with two plates; sterigma a curved cup-shape; ostium bursae ovate, simple, with bulbous anterior end (before a constriction entering ductus bursae); ductus bursae simple, merging into bursae with two spined plates at the border; ductus seminalis large, emergent dorsally from ductus bursae near subostium; corpus bursae simple, elongated tear-drop shape, gradually merging with ductus bursae without clear demarcation other than the ductal plates; signum a small thorn-like process. Male genitalia unknown.

Etymology. The generic name means "elegant" (compso) "ornamentation" (commosis) in Greek.

Discussion. The genus appears most related to Mictocommosis , although the female genitalia are unique. Wing venation is similar to that of Mictocommosis , except R4–R5 are closer together at the end of cell in the forewing (more separate in Mictocommosis ), and in the hindwing Rs extends to just above the wing apex (at the apex in Mictocommosis ). Compsocommosis and Mictocommosis clearly belong in Archipini based on characters of the tribe ( Horak 1984, 1998, Razowski 1987), most notably the signum of the females with the long-curved, spine-like shape (although short in Compsocommosis ) and the absence of hami in the male genitalia, all atypical for Hilarographini , although we do not yet know the male for Compsocommosis . Likewise, the labial palpi are only somewhat upturned, like other Archipini , while in Hilarographini they are much recurved. The main confusion with Mictocommosis has likely been the male genitalia, with the hilarographine-like socius formation. While somewhat atypical for Archipini , it is within the character range for the tribe, but much less developed than found for genera like Hilarographa and Thaumatographa of the true Hilarographini , which also have hami. Wing venation in these genera also clarifies their placement: for Compsocommosis and Mictocommosis the venation does not have the hindwing radius stalked with M1 (likewise in the Neotropical genus Mictopsichia ), while in true hilarographines (viz., Hilarographa and Thaumatographa ) these veins are stalked.

Figures

FIGURE 1. COmpSOcOmmOSiS vietnamenSiS n. sp., female holotype (14.5 mm wingspan).

FIGURES 2 – 3. COmpSOcOmmOSiS vietnamenSiS n. sp., head close-ups (lateral and dorsal views), female.

FIGURES 4 – 5. Wing venations: 4) COmpSOcOmmOSiS n. gen. (C. vietnamenSiS n. sp., female); 5) MictOcOmmOSiS Diakonoff (M. nigrOmaculata (Issiki), male, redrawn from Diakonoff, 1977).

FIGURE 6. COmpSOcOmmOSiS vietnamenSiS n. sp., female genitalia, with details a) sterigma and ostium, with anterior split colliculum below subostium; b) ductus bursae cestum-like dentate plates; and c) signum (slide JBH 2859).