Degeeria multifasciata Tullberg, 1871 Entomobrya multifasciata , Brook 1884 Entomobrya nicoleti var. multifasciata , Börner 1901 Entomobrya aurantiaca Stach, 1922 Entomobrya multifasciata Biogeographical and ecological insights from Australasian faunas: the megadiverse collembolan genus, Entomobrya (Entomobryidae) Jordana, Rafael Greenslade, Penelope Zootaxa 2020 2020-05-06 4770 1 1 104 (Tullberg, 1871) Brook, 1884 , Brook 1884 Tullberg 1871 [151,832,826,853] Entognatha Entomobryidae Entomobrya Animalia Collembola 51 52 Arthropoda species multifasciata  ( Figs 2J, 5D, 27A–D)    Degeeria multifasciata Tullberg, 1871   Entomobrya multifasciata, Brook 1884   Entomobrya nicoletivar. multifasciata, Börner 1901   Entomobrya aurantiaca Stach, 1922   Entomobrya multifasciataf. nigra Uchida, 1954     Typelocality. Sweden  Material examined.  SAMA: male, ACT, Constitution Ave, Russell, Canberra, - 35.296°S, 149.150°E,  578masl, pitfallsite 5,  1–5.iv.1991, J. Dorrowleg.;  NSW, Armidale, Chiswickfarm, nr Armidale, - 32°S, 147°E,  980masl, pasture, grassland,  September 1973, KK leg.; 6 specimenmale;  NSW, Legume, - 28.40472°S, 152.30758°S,  583maslopen area in woodland,  20.ix.1974, JG leg.; 3 specimensfemale,  NSW, Armidale, - 32°S, 147°E,  980masl; female Juvenile,  NSW, Arawana Headland, - 30.065°S, 153.200°E,  5maslin sea weed on beach,  11.i.1974; female  NSW Richmond, - 33.600°S, 150.750°E,  19 masl, grass in garden,  31.xii.1975; 4 specimens,  NSW, Armidale, - 32°S, 147°E,  980masl; female,  QLD,, nr Leslie Dam, - 28.21°S, 151.97°E, sweeping,  13.05.1974, PG leg.; female  QLD, Brisbane, - 27.469°S, 153.025°E,  22masl, in irrigated pasture  8.ix.1933; female SA, Waite Institute, Urrbrae, - 34.967°S, 138.636°E,  118 masl, sub clover,  July 1930, 24,viii.31, HW leg.; female SA, Cambrai, - 34.663°S, 139.321°E,  305masl,, bracken, March, 1972, PG leg.; female SA, Andamooka, - 30.453°S, 137.164°E,  76masl, 2.iv.72; SA, Belair, 1.2  ii.1976, PG leg.; Belair, - 35.007°S, 138.633°E,  300masl; 2 femaleand male, juvenile, SA, Glen Osmond, - 34.960°S, 138.650°E,  300masl,  1.ix.1974; female, SA, Mitcham, - 34.987°S, 138.625°E,  62masl, garden soil, 21.xi.85,  R. V. Southcottleg.; 1male, 1female, SA, Hindmarsh Island, Lane’sproperty, - 35.533°S, 138.86°E,  25masl, pitfalls,  4.x.2013, PG GH leg.; male SA, Hindmarsh Island, Anne’sproperty, young,  4.x.2013, PG GH leg.; - 35.516°S, 138.866°E,  25masl,  5.x.2013, PG GH leg.; Juvenile. SA, Glen Osmond, stray in funnels in laboratory, - 34.960°S, 138.650°E,  300masl, and - 35.533°S, 138.86°E,  25masl; male SA, Hindmarsh Island, Hill’sproperty, - 35.516°S, 138.866°E,  25masl; female TAS, New Town, Hobart, - 42.859°S, 147.305°E,  49masl, 3.ix.32; juvenile, same location in Tasmania, 3.ix.32; juvenile, 1 male, 1female, WA, Mundijong, - 32.284°S, 115.973°E,  42masl, 25.vi.98, PG leg.; 3 specimens- 34.9600°S, 138.6502°E, 300masl PG leg.   Redescription based on Australian material.Size. Length up to 0.9 mmexcluding antennae (n=7). Colour. Pattern as in Fig 2J, some colour variations shown. Head. Antennal length 847–1020 μm, 2–3 times the length of the head; Ant IV with a bilobed apical vesicle. Relative lengths of Ant I/II/III/IV=1.0/2.1/1.9/2.6. Four spinulate labral papillae or with some projections, four ciliated prelabral chaetae and 5,5,4 labral smooth chaetae. Lateral process of papilla E reaching at the papilla tip. Labial posterior row with MREL 1L 2, R smaller than M; eight eyes, G and H similar in size to E and F. Thorax and abdomen. Ratio of lengths of Abd IV/III=4 (n= 7). Trochanteral organ near of 22 chaetae. Unguis with 4 teeth on internal edge: first pair at 50% distance from base of unguis, and 2 unpaired teeth, first one at 75% from the base and the most distal one minute. Dorsal tooth approximately at half distance between the base of unguis and internal pair of teeth. Unguiculus spike–like, with a smooth or external edge on leg III. Furcal length 636 μm (n=7). Manubrial plate with 4 chaetae and 2 psp. Mucro with 2 teeth, subapical tooth in size similar to the apical one. Mucronal spine present. Macrochaetotaxy. Simplified formula: 3,1,0,3,2/2,3/2,2/1,2,1/0,2,3,2,2. Head chaetotaxy with H1 area with An 2, An 3a1and An 3, and as in Fig. 27A, H 2with A 5, S’ 0Mc no present; H4 with S 1, S 3and S 4iMc, H5 with Ps2 and Ps5. Thoracic chaetotaxy Area T1 on Th II with 2 Mc (m 1and m 2ior m 2present) ( Fig. 27B). Area T2 on Th II with 3 Mc present (a 5, m 4and m4i). Abdominal chaetotaxy Area A1 on Abd II with 2 Mc as in Fig. 85 and area A2 on Abd II with 2 Mc. Abd III with 1 Mc on areas A3 and A5, 2 Mc on area A4 as in Fig. 27C. Abd IV with 18 Mc (A 3–A 6, B 4–B 6, E 1and C 2a) sometimes a variation on Area A7 on Abd IV 1–3 Mc. ( Fig 27D). Sensillary chaetotaxy S =21/02293 and Ms =10/101, similar position to  E. nigrocincta.   Remarks.The characteristic colour may fade and only remain on the anterior part of Th II and on a distal part of the body. The only difference from the European  E. multifasciata(cf. Jordana, 2012) is the presence of 3 Mc on H 4 inthe head, but some Australian specimens are asymmetric in this character.   FIGURE 27.  Entomobrya multifasciataLinedrawings A–D: A, head chaetotaxy; B, Th II chaetotaxy; C, Abd II–III chaetotaxy; D, Abd IV chaetotaxy. Bar=0.05 mm. Open circles mean unstable Mc. This introduced species is the most widespread and abundant species of the genus in Australia. It has been misidentified as  E. tenuicaudapreviously and with females of  E. nigrocinctabecause of a similar colour. It is rarely found in native vegetation and nearly always in improved exotic grasses in pasture and on other open disturbed land. 2610568471 [151,405,1112,1139] Sweden Sweden 51 52 1 holotype 2610568499 1991-04-01 1991-04-05 1991-04-01 SAMA J. Dorrow Constitution Ave 578 -35.296 Canberra 71 149.15 Russell 51 52 1 2610568341 1973-09 NSW Armidale 980 -32.0 Armidale 31797 147.0 Chiswick 51 52 6 2610568454 1974-09-20 NSW 583 -152.30759 Legume 51 52 3 2610568492 NSW Juvenile 980 -32.0 Armidale 31797 147.0 51 52 1 2610568468 1974-01-11 NSW 5 -30.065 Arawana Headland 73 153.2 51 52 1 2610568409 [213,1241,1328,1355] 1975-12-31 NSW 19 -33.6 Richmond 72 150.75 51 52 4 2610568544 NSW 980 -32.0 Armidale 31797 147.0 51 52 1 2610568555 [519,1436,1364,1391] 1974-05-13 QLD -28.21 Leslie Dam 740 151.97 51 52 1 2610568515 [151,1108,1400,1427] 1933-09-08 QLD 22 -27.469 Brisbane 74 153.025 51 52 1 2610568551 1930-07 SA 118 -34.967 Urrbrae 71 138.636 Waite Institute 51 52 1 2610568436 SA March 305 -34.663 Cambrai 71 139.321 51 52 1 2610568460 SA 76 -30.453 Andamooka 72 137.164 51 52 1 2610568532 [246,1204,1508,1535] 1976-02 SA 300 -35.007 Belair 71 138.633 Belair 51 52 2 2 2610568443 [151,949,1544,1571] 1974-09-01 SA 300 -34.96 Glen Osmond 71 138.65 51 52 1 2610568449 SA 62 -34.987 Mitcham 71 138.625 51 52 1 2610568416 [443,848,1581,1607] R, V Southcott 51 52 2 1 1 2610568452 2013-10-04 SA 25 -35.533 Lane's 455 138.86 Hindmarsh Island 51 52 1 2610568500 2013-10-04 2013-10-05 2013-10-04 SA Juvenile 25 -35.516 Anne's 71 138.866 Hindmarsh Island 51 52 1 2610568374 SA 300 -35.533 Glen Osmond 455 138.86 51 52 1 2610568312 SA Hindmarsh Island 25 -42.859 New Town 68 147.305 Hill's 51 52 2 1 1 Tasmania 2610568507 42 -32.284 Mundijong 72 115.973 51 52 3 Tasmania