Porcellanidae (Crustacea: Decapoda: Anomura) from New Caledonia and the Loyalty Islands Osawa, Masayuki Zootaxa 2007 1548 1 49 a54a6f46-eeec-47d4-a8a5-7c28b361cfd0 [640,811,301,327] Malacostraca Porcellanidae Animalia Decapoda 3 4 Arthropoda family    1. Antennal peduncle with movable (second to fourth) articles freely accessible to orbit .............................. 2  - Antennal peduncle with movable articles excluded from orbit by projection of immovable (first) article meeting anterior margin of carapace ........................................................................................................... 6   2. Chelipeds flattened. Pterygostomian flap entire .......................................................................................... 3  - Chelipeds robust. Posterior part of pterygostomian flap composed of 1 or more pieces separated by mem- branous interspaces from anterior part ............................................................  Pachycheles Stimpson, 1858   3. Rostrum narrow, triangular, with row of spines on each lateral margin. Telson composed of 5 plates ........ ................................................................................................................................  Petrocheles Miers, 1876  - Rostrum broad, unarmed except for sometimes with supraocular spine on each lateral margin. Telson composed of 7 plates ................................................................................................................................... 4   4. Carapace with gastric region strongly elevated. Ambulatory legs subcylindrical......................................... .....................................................................................................................  Neopetrolisthes Miyake, 1937a  - Carapace flattish or somewhat convex on dorsal surface. Ambulatory legs flattish ................................... 5   5. Carapace uneven on dorsal surface. Median lobe of rostrum with pair of lateral elevations........................ ...........................................................................................................................  Novorostrum Osawa, 1998a  - Carapace genarally even on dorsal surface. Median lobe of rostrum without lateral elevations .................. ..........................................................................................................................  Petrolisthes Stimpson, 1858   6. Carapace broader than long ......................................................................................................................... 7  - Carapace as long as or longer than broad .................................................................................................. 11   7. Ambulatory legs with dactyli each terminating in simple claw (no corneous spines present on flexor mar- gin) .................................................................................................................... Raphidopus Stimpson, 1858  - Ambulatory legs with dactyli each with biunguiculate claw....................................................................... 8   8. Branchial margins of carapace armed with spines, outer orbital angle terminating in small spine............... ................................................................................................... Aliaporcellana Nakasone & Miyake, 1969  - Branchial margins of carapace without spines, outer orbital angle unarmed .............................................. 9   9. Carapace with pair of transverse ridges along cervical regions. Telson composed of 5 plates ..................... ........................................................................................................................  Heteropolyonyx Osawa, 2001  - Carapace with weak or distinct, cervical grooves. Telson composed of 7 plates ...................................... 10   10. Carapace with dorsal regions well defined by grooves; cervical and gastric grooves formed by thinning and gradual decalcification of cuticle ........................................................Eulenaios Ng & Nakasone, 1993  - Carapace with dorsal regions not distinctly defined by grooves ..........................  Polyonyx Stimpson, 1858   11. Carapace distinctly elongate. Dactyli of ambulatory legs triunguiculate or quadriunguiculate................ 12  - Carapace not elongate. Dactyli of ambulatory legs each terminating in a single claw ............................. 13   12. Carpi of chelipeds longer than broad. Dactyli of ambulatory legs quadriunguiculate .................................. .............................................................................................................................  Porcellanella White, 1852  - Carpi of chelipeds broader than long. Dactyli of ambulatory legs triunguiculate......................................... .................................................................................................... Pseudoporcellanella Sankarankutty, 1961   13. Carapace strongly areolate on dorsal surface ............................................................................................ 14  - Carapace nearly smooth or striate on dorsal surface ................................................................................. 15   14. Branchial margins of carapace with spines. Chelae flattened ...........................  Enosteoides Johnson, 1970  - Branchial margins of carapace without spines. Chelae inflated .....................  Capilliporcellana Haig, 1978   15. Chelipeds subequal, dactyulus of smaller cheliped not twisted ..........................  Porcellana Lamarck, 1801  - Chelipeds unequal, dactylus of smaller cheliped usually twisted.............................................................. 16   16. Carapace with dorsal regions clearly defined. Rostrum broadly triangular or rounded in dorsal view, median lobe strongly deflexed ............................................................................... Ancylocheles Haig, 1978  - Carapace with dorsal regions weakly defined. Rostrum horizontal, tridentate or trilobate in dorsal view [except for  Pisidia dispar( Stimpson, 1858)with rostrum strongly bent ventrally, broadly rounded in dor- sal view]..................................................................................................................................................... 17   17. Chelae rather flattish, showing pronounced sexual dimorphism in spinulation as well as in degree of dis- tortion of fingers ............................................................................................................  Pisidia Leach, 1820  - Chelae subcylindrical, generally similar in both sexes ......................................  Lissoporcellana Haig, 1978