Nomenclatural Status Of The Palaeobotanical “ Artificial Taxa ” Established In Brongniart’S 1822 “ Classification ” Paper Cleal, Christopher J. Thomas, Barry A. Fossil Imprint 2018 2018-08-31 74 1 - 2 9 28 BRONGNIART BRONGNIART [937,1342,1472,1496] Filicites GBIF Plantae 9 18 Tracheophyta section Pecopteris   Text-fig. 1g–i 1822a  Filicites( Pecopteris) BRONGNIART, p. 233. 1825  Pecopteris (BRONGNIART) STERNBERG, Tentamenp. xvii. Ty p e.  Filicites(sect. Pecopteris) penniformis BRONGNIART, 1822a, p. 233, pl. 2, fig. 3; Loc. Middle Pennsylvanian Series, Dudweiler, near Saarbrücken, Germany(vide Brongniart 1836: 345); ≡  Pecopteris penniformis(BRONGNIART) BRONGNIART, 1828a. D i a g n o s i s. “la fronde est pinnatifide à pinnules adherents par leur base au rachis, transversée par une nervure médiane et à nervures secondaires pennées.” D i s c u s s i o n. When Sternberg (1825)raised this species in rank to genus, he illegitimately renamed the typespecies  Pecopteris pennata STERNBERG, 1825, but this name must be supressed as a later homotypic synonym. In the original definition of Brongniart (1822a)this name was used for both fern and seed-plant fronds. However, mainly through the separation off of species with larger pinnules such as  Alethopteris STERNBERG, 1825, and Callipteridium(E. WEISS) GRAND’ EURY, 1877 (≡  Odontopteris( Callipteridium) E. WEISS, 1870) the name has become almost exclusively used for Palaeozoic fern foliage. Only fronds of the seedplant family Callistophytaceae STIDD et HALL, 1970, have occasionally still been referred to  Pecopteris, but even this is nowadays unusual; such fronds are normally now referred to  Dicksonites STERZEL, 1881. Most Palaeozoic pecopteroid foliage belong to two families: the Psaroniaceae UNGER ex ENDLICHER, 1842, ( Marattiales) and Tedeleaceae EGGERT et TAYLOR, 1966, (Botryopteridales). The typeof  Pecopterisunequivocally belongs to the family today usually referred to as the Tedeleaceaeand so the name should be restricted to species belonging to that family, most notably  P. penniformis,  Pecopteris plumosa(ARTIS) BRONGNIART, 1836(≡  Filicites plumosus ARTIS, 1825) and  Pecopteris volkmannii SAUVEUR, 1848. These species have in the past been placed in a separate fossil-genus based on the distinctive soral structures, variously named  Senftenbergia CORDA, 1845and  Dactylotheca ZEILLER, 1883, but (as pointed out by Cleal 2015) these names should be supressed as later taxonomic synonyms of  Pecopteris. On this basis,  Pecopterisshould not be used for species that are demonstrably attributable to the Psaroniaceae. Most of the well-established Psaroniaceaefossil-species can be assigned to one or other of the fossil-genera that are based on a combination of pinnule morphology and venation, and sporangial form (as summarised in Cleal 2015).