Bromopsis dolichocarpa (Wagnon) Holub (1973: 167)
A taxonomic revision of Bromus (Poaceae: Pooideae: Bromeae) in México and Central America
Saarela, Jeffery M.
Peterson, Paul M.
Valdés-Reyna, Jesus
Phytotaxa
2014
2014-11-14
185
1
1
147
9. Bromus dolichocarpus Wagnon (1950: 65). Figs. 32, 33.
Bromopsis dolichocarpa(Wagnon) Holub (1973: 167). Type:— MÉXICO. Michoacán: in fir forest, ca. 10 miNW of Ciudad Hidalgo, 18 March 1949, R. McVaugh 9887( holotype MICH-1108613!, isotype US-2012739!). Plants perennial, not rhizomatous. Culms up to 192 cmtall, 2–4 mmwide at base, erect, the bases ± decumbent, pubescent below inflorescences; nodes 2–6, glabrous, puberulent or pubescent. Leaf sheaths sparsely to moderately pilose, hairs up to 1.5 mmlong; auricles usually present; ligules 0.6–1.5(–3) mm long, glabrous; blades up to 39 cm× 3–16 mm, flat, adaxial surfaces pilose, hairs up to 1.5 mmlong, abaxial surfaces glabrous, sometimes with scattered hairs adjacent to the midnerve, or sparsely pubescent, margins serrulate. Panicles up to 33 cm× 3–11 cm, open, nodding, branches ascending, 0.7–12 cmlong, shorter or longer than spikelets, scabrous to densely pubescent, hairs up to 0.6 mmlong, 1–2(–3) spikelets per branch. Spikelets 2.5–3.6(–4) cm long, 5–8-flowered, elliptic to lanceolate, terete to moderately laterally compressed, rachillas sometimes visible at maturity; glumes glabrous, sometimes a few hairs proximally and along the margins, or pubescent proximally, margins hyaline, midnerves scabrous, sometimes pubescent proximally, hairs up to 0.6 mmlong; lower glumes 10–13(–14) mm × 0.3–0.7(–1) mm, narrowly lanceolate, 1(–3)-nerved, green along the nerve(s), apices acute, rarely acuminate; upper glumes 12–19 mm× 0.7–1.3 mm, lanceolate, 3-nerved, green along the nerves, green to light green–translucent between the nerves, apices acute, rarely acuminate; lemmas (13–) 14–20 mm× 0.9–1.5 mm, linear-lanceolate, rounded over the backs, apices acute, 3(–5)-nerved, the nerves strong, backs puberulent proximally, glabrous distally, pubescent to pilose between the lateral nerves and the margins, hairs up to 0.5 mmlong, light greentranslucent between the nerves, margins hyaline, nerves scabrous or pilose, hairs up to 0.6 mmlong; awns 7–13 mmlong, attached up to 0.5 mmbelow lemma apex, straight; paleas shorter and narrower than the lemmas, backs
ATAXONOMIC REVISION OF BROMUS(POACEAE) Phytotaxa185 (1) © 2014 Magnolia Press 77 scabridulous to pubescent, keels ciliate, cilia up to 0.1 mmlong; anthers 1.1–2.5 mmlong; caryopses 10−15 mmlong, light brown. 2 n= 28 ( Wagnon 1950). FIGURE 32.Illustration of Bromus dolichocarpus. A. Spikelet. B. Lemma C. Inflorescence. D. Ligule. Based on Breedlove 55564(GH). Illustration by Paulette Dennis © Paulette Dennis. 78 Phytotaxa185 (1) © 2014 Magnolia Press SAARELA ET AL.
Distribution:―Native. Bromus dolichocarpusis distributed from central México( Chiapas, Distrito Federal, Durango, Guerrero, Hidalgo, Jalisco, Estado de México, Michoácan, Morelos, Oaxaca, Puebla, Veracruz) to Guatemala( El Progreso, Quezaltenango, San Marcos) ( Fig. 34). It was reported from Colima and Jalisco by Beetle (1977), but we have not seen specimens from these states. This is the first report of B. dolichocarpusfrom Durango, as the species was not reported for the state by Herrera Arrieta (2001, 2014).
Ecology:— Bromus dolichocarpusis found on gentle slopes and flats that are often steep and rocky and along small draws; associated with Pinusspp., Pinus hartwegii Lindley (1839: 62), Abies religiosa, Quercusspp., Juniperus deppeana, Arbutus xalapensis, Gaultheria Linnaeus (1753: 395), PteridiumGled. ex Scopoli (1760: 169), Tagetes Linnaeus (1753: 887), Symphoricarpos Duhamel du Monceau (1755: 295), Arctostaphylos pungens, Alnus, Ribes Linnaeus (1753: 200), Prunus serotina Ehrhart (1783: 285), Rhamnus Linnaeus (1753: 193), Crataegus Linnaeus (1753: 475), Toxicodendron Miller (1754a), Geranium( Linnaeus 1753: 676)and Baccharis. Elevation: 2500–3800 m( Guatemala), (1900–) 2200–3920 m( México). Common Names:―Unknown.
Comments:― Bromus dolichocarpusis one of three Mexican species with 3-nerved lemmas, which are newly recognized here in sect. Mexibromus. It is easily distinguished from B. attenuatusby its pubescent lemmas, longer awns and leaf blades that do not distinctly narrow towards their bases, and from B. densusby its longer awns, shorter anthers and wider leaf blades (see taxonomic key). Specimens Examined:― GUATEMALA. El Progreso: Sierra de Las Minas, between Finca Piamonteand top of Montaña Piamonte, along Joya Pacayal, [ 15.13°N, 89.75°W], 2500–3000 m, 7 February 1942, J. A . Steyermark43619(F-1202053, US-2436906). Quezaltenango: Cerro Quemado, [ 14.8°N, 91.52°W], 2600 m, September 1954, M . de Koninck210( US-2153572); mountains SE of Palestina, on old road to San Juan Ostuncalco, [ 14.87°N, 91.62°W], 2550–2850 m, 21 January 1941, P. C . Standley 84221(F-1202285). San Marcos:along road between SanSebastián at km 21 and km 8, 8– 18 mi NWof San Marcos, [ 15.06°N, 91.83°W], 2700–3800 m, 15 February 1940, J. A . Steyermark35769(F-1052081) & 35771(F-1052024); Sierra Madre Mountains, ca. 6 km(airline) Nof San Marcos, [ 14.96°N, 91.79°W], 2700 m, 13 December 1963, L. O Williams, A. MolinaR. & T.P. Williams25876(F-1656824, US-2537602) & 25885( NY, mixed sheet with B. carinatusvar. marginatus). MÉXICO. Chiapas: Mpio. Zinacantán, near the summit of Muk'tavits, [ 16.3333°N, 92.75°W], 2745 m, 16 November 1981, D. E . Breedlove& B. Bartholomew55564( CAS, GH, MO). Distrito Federal:cerca de Eslava, 19.42°N, 99.17°W, 19 October 1952, J . Rzedowski2005( MEXU-100903, MICH-1119221); Cumbre de Estepa, 18.05°N, 97.7°W, April 1842, Liebmann487( US-2473643); in imperio Mexicano, from Zuccarini, Karwinskys.n.( US-1009477); San Rafael, 19.25°N, 98.79°W, 2600 m, 18 September 1932, M . St. Pierre938( MICH-1119222, P-03629787, US); SanRoman, Liebmann501( MO, US-2473641); 10 mi EofAmecamecaon road towards Paso de Cortés, in Parque Nacional Izta-Popo, 19.0764°N, 98.71°W, 2840 m, 12 October 2001, P. M . Peterson& Y. Herrera-Arrieta16160( CAN, MO, US); 1er Dínamo, Delegacíon Magdalena Contreras, [ 19.42°N, 99.13°W], 2580 m, 17 October 1985, A . Miranda& P. Guerrero141( MEXU-1110774); Desierto de Los Leones, 19.2531°N, 99.3286°W, 19 October 1960, T . Tateoka389( MEXU-151424) & 1151( US-2380596); Desierto de Los Leones, Delegación de Guajimalpa, [ 19.2533°N, 99.3308°W], 25 January 1976, M . González E. 82( ANSM); Mpio. Cuajimalpa, Desierto de Los Leones, [ 19.2533°N, 99.3308°W], 2 November 1979, 2700 m, F . Santoyo178( ANSM); Río Grande, July 1928, E . Lyonnet266( MEXU-258403); Vallede México, [ 19.42°N, 99.13°W], 3500 m, 9 December 1951, E . Matuda25754( MEXU-499579, MEXU-499578); Vallede México, Lago Zempoala, Hdo.de Morelos, 19.0494°N, 99.3175°W, 3000 m, 7 October 1951, E . Matuda25587( MEXU, US-2079183); Vallede México, San Rafael, 19.2333°N, 98.75°W, 2600 m, 19 November 1950, E . Matuda18722( MEXU, US-2040890). Durango:Mpio. Súchil, Reserva de la Biosfera La Michilia, Mesa El Burro, 23.4108°N, 104.3108°W, 2650 m, 16 September 1982, Y. Herrera261( ANSM). Guerrero: Mpio. Chilpancingode los Bravos, Omiltemi, [ 17.55°N, 99.51°W], 1900 m, 15 November 1991, V. Nandi134( MEXU-1089776); Mpio. Chilpancingode los Bravos, al NWde Omiltemien direccion a la toma de Aguade la Cañada de la Perra, [ 17.55°N, 99.5°W], 2120 m, 10 October 1985, R. Antonio Ocampo405( MEXU-502476). Hidalgo: El Chico, 20.2167°N, 98.7333°W, July 1929, P. Lyonnet262( US-1034151); Las Ventanas, Mineral del Chico, [ 20.17°N, 98.73°W], 2970 m, 19 May 1993, J. Práxedes Pérez23( MEXU); Pachuca Dist., below Pueblo Nuevoon road from Real del Monteto El Chico, 20.1667°N, 98.7333°W, 2850 m, 17 October 1946, H. E. Moore, Jr.1531( GH, US-1963095). Jalisco:on the divide above headwaters of Río Mascota( 25–30 km, airline, SE of Talpa de Allende), steep mountainsides and barrancas 1–2 mi Nofthe sawmill " LaCumbre", and 11–12 road mi SofEl Rincón, 20.0403°N, 103.8753°W, 2000–3000 m, ATAXONOMIC REVISION OF BROMUS(POACEAE) Phytotaxa 185 (1) © 2014 Magnolia Press 79 29 November 1960, R . McVaugh21533( NY); steep mountainsides above Amacueca, near the summit of the plateau, road to Tapalpa, [ 19.1556°N, 102.9889°W], 2100–2250 m, 2 November 1960, R . McVaugh20654( NY). Estado de México: 8 kmde la desviación al Nevado de Toluca, [ 19.1083°N, 99.7583°W], 3260 m, 29 November 1983, Manrique, Jaramilo& Núñez558( MEXU-1089780); a 6 kmsobre la desviación a Tequesquiapan, carretera Toluca-Temascaltepec, 19.31°N, 98.97°W, 2580 m, 25 October 1983, Manrique, Jaramillo& Nuñez445( MEXU- 1098071, MEXU-1033954); al SE de Zinacantepec, [ 19.14°N, 99.71°W], 3920 m, 8 April 1992, A . Vega, A. Ramírez& R. Mendoza2( MEXU-1089771); camino de terracería Malinalco-San José Tenería, [ 18.93°N, 99.49°W], 27 October 1988, J. G . Aldapeet al. 34( MEXU); carretera Toluca-Temascaltepec, [ 19.12°N, 99.88°W], 2530 m, Manrique, Guerrero, Guzmán& Jaramillo140( MEXU); Cieneguillas de Cabra, entre la carretera Sultepec-La Puerta, al Sdel Nevado de Toluca, [ 19.1083°N, 99.7583°W], 2600 m, 1 August 1981, R. Guzmán4021( MEXU); entre Texcaltitlán y Las Juntas, al Sdel Nevado de Toluca, [ 18.93°N, 99.95°W], 2660 m, 1 August 1981, R. Guzmán4015( MEXU-1110805); Estaciónexperimental SanCayetano, 2500 m, 2 August 1978, F. M . Fuchs97( MEXU-231170); Mpio. Amecameca, 1 kmal noroeste de San Antonio, 2550 m, 11 December 1977, N . Valentín M. 74( ANSM); Mpio. Amecameca, km 15 carretera Amecameca-Tlamacas, [ 19.08°N, 98.67°W], 2 October 1992, A . Miranda& G. Villegas636( MEXU); Mpio. Atlautlade Victoria, Volcán Popocatépetl, ladera W por Atlautla, [ 19.05°N, 98.64°W], 2725 m, 24 October 1987, J . Hernández230( MEXU-565906); Mpio. México, 3 kmal EdeSan Rafael, [ 19.21°N, 98.74°W], 2700 m, 19 September 1976, C . Flores G. s.n.( MEXU-292304); Mpio. Morelos, Valledel Tepeite, [ 19.27°N, 99.18°W], 16 October 1937, E . Lyonnet18112( MEXU-278684); Mpio. Ocuilan de Arteaga, 2 kmal NWde Sta. Mónica de Ocuilan, 18.9833°N, 99.4417°W, 2350 m, 4 February 1990, González& O . Miranda J. 328( MEXU-1031709); Mpio. Tepotzotlán, 2400 m, 26 October 1975, J. R . Torres V.( ANSM); Mpio. Texcoco, 2 kmal SE de San Pablo Ixayoc, sobre el camino hacia el Aguaje, 12.5 kmal SE de Texcoco, [ 19.41°N, 98.86°W], 2670 m, 21 October 1976, S . D. Koch76210( MEXU-265511, MO); Mpio. Tlalmanalco, 3 kmal EdeSan Rafael, [ 19.21°N, 98.76°W], 2700 m, 27 May 1965, J . Rzedowski19863( MEXU- 187911); Mpio. Tlalmanalco, a 3 kmal EdeSan Rafael, [ 19.21°N, 98.76°W], 2700 m, L. M . GarcíaR. 73( ANSM, MEXU-220002); Mpio. Villa de Allende, SanCayetano (estación experimental) al NdeAgua Escondiday a 26 kmal NEde Valle de Bravo, [ 19.37°N, 100.15°W], 2490 m, August–December 1974, Ma. E . Maury Hdez., V. Serrano Cardenas& S. Gallina Tessaro5( MEXU-200087); Mpio. Villa de Allende, SanCayetano ( Estaciónexperimental), al NdeAgua Escondiday a 26 km NEde ValledeBravo, [ 19.33°N, 100°W], 2490 m, August–December 1974, S . Gallina Tessaros.n.( MEXU); Mpio. Villa Guerrero, carretera de cuota de Tenango de Aristaa Ixtapan de la Sal, [ 18.84°N, 99.68°W], 2200 m, 30 September 1995, H . Vibrans5564( MEXU-895630); Mpio. Zimatlán, Cañada El Frijolón, 8 kmal NWde La Cofradía, comunidad de San Pedro El Alto, [ 19.61°N, 99.9°W], 2790 m, 8 October 1998, A. G . Miranda M. & O. L. Hernández Martínez248( MEXU-907764); 1 km WofAmecamecaalong road to Ayapango, 19.15°N, 98.7833°W, ca. 2500 m, 29 September 1978, H. H . Iltis, J. Doebley& A. Lasseigne764(F-1919412, MO, MSC-277227, RSA-POM-320576, US); 1.4 mi NofTequexquipan, 2490 m, 19.071°N, 99.9388°W, 10 October 2001, P. M . Peterson& Y. Herrera-Arrieta16139( CAN, MO, P-03216909, US); 1.9 mi WofChiltepec on Hwy.12, 18.9268°N, 99.8442°W, 2596 m, 10 October 2007, P. M . Peterson, J. M. Saarela& M. J. Flores Villegas21364( CAN, MO, P-03631106, US); 12.1 mi NEof San Miguel, SW of Volcán Toluca, 19.0317°N, 99.9102°W, 2826 m, 10 October 2001, P. M . Peterson& Y. Herrera-Arrieta16143( CAN, MO, P- 03216911, US); 5 kmal NWde Amecameca, laderas vecinas al Cerro Tenayo, [ 19.12°N, 98.76°W], 2550 m, 17 November 1968, A . PinedaR. 603( MICH-1119227, MSC-223104); 6.7 mi EofAmecamecatowards Pasode Cortés, 19.0706°N, 98.6923°W, 2488 m, 11 October 2007, P. M . Peterson, J. M. Saarela& M. J. Flores Villegas21382( CAN, MO, US); 8.7 mi NEof La Comumidad on Hwy.134 toward Toluca, 19.1858°N, 99.86°W, 3190 m, 9 October 2001, P. M . Peterson& Y. Herrera-Arrieta16132( US); at La Comunidad, along stream just below puebla ( LaComunidad), 19.1518°N, 99.9118°W, 2607 m, 9 October 2001, P. M . Peterson& Y. Herrera-Arrieta16136( CAN, MO, P-03216910, US); Coulter1651( GH); Mpio. Amecameca, 1 kmal NEde San Antonio, 19.12°N, 98.76°W, 2550 m, 21 October 1968, J. J . Rzedowski26431( MICH-1119223, MSC-233266); Mpio. Amecameca, cerca de San Pedro Nexapa, 19.08°N, 98.74°W, 9 October 1966, 2550 m, J. J . Rzedowski23285( MSC-217972); Mpio. Amecameca, Zumpango, carretera al Pasode Cortés, 16 kmal Edelentronque con la carretera Amecameca-Cuautla, Mor., 19.06°N, 98.7°W, 3300 m, 20 November 1976, S. D. Koch76275 B( US- 2824581); Mpio. Huixquilucan, 5 kmal Ede México, carretera de la Marquesaa Naucalpan, 19.3833°N, 99.3333°W, 2600 m, 29 July 1976, S . D. Koch76107( MEXU-270647, CHAPA, MO, US); Mpio. Texcoco, 2 kmal SE de San Pablo Ixayoc, 12.5 kmal SE de Texcoco, 19.42°N, 98.83°W, 2670 m, 21 October 1976, S .D. Koch 80 Phytotaxa185 (1) © 2014 Magnolia Press SAARELA ET AL.ATAXONOMIC REVISION OF BROMUS(POACEAE) FIGURE 33. Bromus dolichocarpus. PinedaR. 603(MICH-1119227). Phytotaxa185 (1) © 2014 Magnolia Press 81 76210(US-2824630); Sslopes of Nevado de Toluca, 35 km(road) SW of Toluca on Hwy. 130, 19.0833°N, 99.8333°W, ca. 3000 m, 29 August 1965, K. Roe, E. Roe& S. Mori1491 ( MSC-229117); Temascaltepec Dist., Comunidad, 19.1292°N, 99.9347°W, 2600 m, 25 November 1932, G. B. Hinton2459( NY, US-1867841); Temascaltepec, Crucero, 19.04°N, 100.23°W, 28 October 1935, G. B. Hinton8395( MO, NY, 2 sheets, RSA-POM- 364158); Tlalmanalco, 3.0 km al EdeSan Rafael, [ 19.21°N, 98.74°W], 2700 m, 19 September 1976, C. Flores G. s.n.( MO, SD-110652); Volcán de Toluca, 3760 m, 19.1014°N, 99.7694°W, 18 October 1953, E. R. Sohns& E. Matuda1004( US-2118910); Wslopesof Nevado de Toluca, 35 km(road) SW of Toluca on Hwy.130, 19.0833°N, 99.8333°W, 3000 m, 29 August 1965, K. Roe, E. Roe& S. Mori1491( US-3115415); Wslopesof Nevado de Toluca, 35 km(road) SW of Toluca on Hwy.130, [ 19.11°N, 99.79°W], ca. 3000 m, 29 August 1965, K. Roe, E. Roe& S. Mori( GH). Michoácan: Cerro Tancítaro, 27 kmal WdeUruapanen linea recta, 1 kmal oeste de El Tepetate, [ 19.41°N, 102.3°W], 2850 m, 29 August 1996, I. Garcia Ruíz, J.A. Machuca & M. Cházaro4153( MICH- 1119229); 11 kmal NdeIndaparapeo, sobre el camino a Las Peras, [ 19.78°N, 100.97°W], 2200 m, 8 November 1987, J. Rzedowski45692( MEXU-809934); 37 mi EofMoreliaon MEX15, 19.67°N, 100.73°W, 8000 ft[ 2438 m], 20 October 1972, L. H. Harvey& J. T. Witherspoon9419( MICH-1119244); 5.7 mi NWof Huajumbaroon road towards Santa Clara, 19.7364°N, 100.78°W, 2430 m, 8 October 2001, P. M. Peterson16129( CAN, MO, US); a 12 kmal NdeUruapan, carretera a Carapan, [ 19.41°N, 102.06°W], 2075 m, 22 August 1980, J. C. Soto Núñez2393( MEXU); above Villa Escalanteon road to Ariode Rosales, 19.4°N, 101.65°W, 2340 m, 11 November 1949, H. E. Moore, Jr., E. Hernández-X. & H. Porras-H. 5634( US-1983697); ca. 10 mi NWof C. Hidalgo, 19.66°N, 100.71°W, 4 July 1950, H. K. Wagnon1615( MEXU-772510, US-2154832); ca. 18 mi SofPátzcuaro, 19.31°N, 101.66°W, 2713–2743 m, 20–25 November 1961, R. M. King& T. R. Soderstrom5178( MICH-1119226, MEXU- 55441, NY, mixed sheet with B. carinatusvar. carinatus), US-2378973); canyon just NWof Congoon on Hwy.120 towards Pátzcuaro, 19.3806°N, 101.65°W, 2350 m, 8 October 2001, P. M. Peterson16128( CAN, MO, P- 03216912, US); Cerro Azul, vicinity of Morelia, 19.7167°N, 101.1833°W, 2300 m, 4 November 1909, G. Arséne3273( MO, US-1002650); grown at the Botanical Gardens, Universityof Michigan, Ann Arbor, from seed collected with the typein fir forests ca. 10 mi NWof Ciudad Hidalgo, 4 July 1950, H. K. Wagnon1615( MICH- 1119225[chromosome number voucher specimen, Wagnon 1950], MO); Mesa Karitshoo del Padre, SE de San Francisco Pichataro, [ 19.57°N, 101.81°W], 2625 m, 9 November 1978, J. Coballero& C. Mapes644( MEXU); Mpio. Charo, El Salto de Agua, 19.65°N, 100.9444°W, 1 February 1994, J. A. Torres Estrada35( MEXU-1009115, MEXU-1089785); Mpio. Morelia, 3 kmal WdeSan Miguel del Monte, [ 19.7°N, 101.18°W], 2300 m, 25 October 1986, J. Rzedowski41312( MEXU-658295); Mpio. Pátzcuaro, vertiente Wdel Cerro del Burro, [ 19.415°N, 101.5006°W], 2900 m, 23 November 1986, J. Rzedowski41962( ANSM); Mpio. Queréndaro, 2 kmal NdeReal de Otzumatlán, 2350 m, 20 November 1986, J. Rzedowski41855( ANSM); Mpio. Senguio, Cerro Ross Azul, al EdeChincus, 19.73°N, 100.35°W, 2800 m, 23 November 1986, J. Santos Martínez1969( MICH-1119219); Mpio. Zacapu, ± 5 km de Zacapu, por la carretera a Zamora, 2000 m, 9 September 1988, A. Grimaldo Núñez406( ANSM); Mpio. Zinapécuaro, 5 kmal SSE de Jeráhuaro, 2550 m, 7 December 1987, J. Rzedowski46150( ANSM, MEXU-915763); slopes of lava flow Eof San Juan Nuevo, ca. 8 km SofUruapan, 19.41°N, 102.06°W, 11–15 October 1961, R. M. King& T. R. Soderstrom4760( MEXU-55372); vicinity of Morelia, Cerro Agul, [ 19.69°N, 101.16°W], 4 November 1909, G. Arséne3293[?] ( GH); Campo Experimental INIF, Uruapán, 2100 m, 9 October 1976, C. L. Días Luna7504( ANSM); carretera Pátzcuaro-Uruapankm 25, [ 19.44°N, 101.84°W], 4 December 1986, J. J. Ortíz1195( MEXU-638476); Mpio. Ocampo, 2 kmal EdeOcampo, camino al Rosario, [ 19.58°N, 100.34°W], 2400 m, 2 November 1989, R. Torres& M. Ramírez13553( MEXU-904824); Mpio. Pátzcuaro, 6 kmal SdePátzcuaro, carretera a Opopeo, [ 19.5158°N, 101.6094°W], 2400 m, 10 November 1985, J. Espinosa Garduño 1990( MEXU-993903); Mpio. Pátzcuaro, lado SW del Cerro El Fríjol, [ 19.5158°N, 101.6094°W], 2700 m, 31 October 1985, H. Díaz Barriga1652( MEXU-662626). Morelos:km 62 de la carretera Curenava-México, [ 19.27°N, 99.17°W], 2460 m, 8 February 1984, Manrique, Beetleet al. 733( MEXU-1110847); Valle del Tepeite, [ 19.27°N, 99.18°W], 15 October 1937, E. Lyonnet1812( MEXU-278689); Valle del Tepeite, 19.27°N, 99.25°W, 16 October 1937, E. Lyonnet& J. Elcoro1811( MEXU, US-1746382). Oaxaca: 1.6 mi NofSan Miguel Suchixtepecalong Hwy.175, 16.1157°N, 96.4745°W, 2722 m, 23 September 2008, P. M. Peterson& J. M. Saarela22378( CAN, US); 9.5 mi NofSan Cualimojoyason road towards Santa María Yavesia, 17.1966°N, 96.4512°W, 2727 m, 20 September 2008, P. M. Peterson& J. M. Saarela22307( CAN, US); Campamento Río de Molino, 4 kmal SW de San Miguel Suchistepec, 16.09°N, 96.47°W, 2250 m, 21 September 1965, J. Rzedowski20999( MICH-1119224), 21039( MICH-1119220) & 21082( MSC-210582); Cerro de San Felipe, Distrito del Centro, 17.1°N, 96.85°W, 82 Phytotaxa185 (1) © 2014 Magnolia Press SAARELA ET AL. 2300 m, 12 October 1920, C. Conzatti 4069( MEXU-6111, US-1014111); Mpio. Zimatlán, Comunidad El Tlacuache, San Pedro El Alto, 16.04°N, 96.47°W, 2420 m, 9 October 1998, A. G. Miranda& O. Hernández M. 306( MEXU-901092); 1.2 mi EofSan José del Pacificoon road towards SanSebastián, 16.1698°N, 96.4898°W, 2641 m, 22 September 2008, P. M. Peterson& J. M. Saarela22353( CAN, US). Puebla: 23.4 Eof Amecamecaon road towards Puebla in Parque Nacional Izta-Popo, 19.0897°N, 98.6064°W, 3330 m, 12 October 2001, P. M. Peterson& Y. Herrera-Arrieta16170( US); 24.3 mi EofAmecamecaon road towards Puebla, in Parque Nacional Izta-Popo, 19.085°N, 98.6°W, 3316 m, 13 October 2001, P. M. Peterson& Y. Herrera-Arrieta16175( CAN, US). Veracruz: Orizaba, 19.0167°N, 97.22°W, s.d., Liebmann 496( US-2473642).
[517,712,1330,1352]
1949-03-18
NW, R
Mexico
Ciudad
Type
76
77
1
Michoacan de Ocampo
[488,1163,1361,1383]
MICH, US
Mexico
76
77
MICH-1108613, US-2012739
1
Michoacan de Ocampo
holotype
1942-02-07
J, A
Guatemala
Sierra de Las Minas
2750
15.13
Montana Piamonte
772
-89.75
Finca Piamonte
78
79
1
El Progreso
1954-09
M
Guatemala
Steyermark
2600
14.8
Cerro Quemado
5581
-91.52
Quezaltenango
78
79
US-2436906
1
El Progreso
1941-01-21
P, C
de Koninck
Guatemala
2700
14.87
San Juan Ostuncalco
772
-91.62
78
79
US-2153572
1
El Progreso
1940-02-15
NW, J, A
Guatemala
3250
15.06
San
772
-91.83
78
79
1
San Marcos
1963-12-13
N, L, O
Guatemala
2700
14.96
Sierra Madre Mountains
772
-91.79
Steyermark
78
79
1
San Marcos
[1300,1432,1015,1040]
A
Guatemala
Williams
78
79
1
San Marcos
NY
Guatemala
Williams
78
79
US-2537602
1
San Marcos
1981-11-16
E
Mexico
2745
16.3333
Muk'tavits
533
-92.75
Mpio. Zinacantan
78
79
1
Chiapas
[488,662,1124,1149]
B
Mexico
Breedlove
78
79
1
Chiapas
[674,1113,1124,1149]
CAS, GH, MO
Mexico
Bartholomew
78
79
1
Chiapas
1952-10-19
J
de Eslava
Mexico
19.42
Distrito Federal
763
-99.17
78
79
1
Distrito Federal
1842-04
1932-09-18
1842-04
M
Mexico
Rzedowski
2600
19.25
Liebmann
763
-98.79
Cumbre de Estepa
78
79
MEXU-100903, MICH-1119221, US-2473643, US-1009477
1
Distrito Federal
2001-10-12
US, MO, E, P, M
Parque Nacional Izta-Popo
Mexico
St. Pierre
2840
19.0764
Liebmann
525
-98.71
San
78
79
MICH-1119222, P-03629787, US-2473641
1
Distrito Federal
[151,305,1340,1364]
Y
Mexico
Peterson
78
79
1
Distrito Federal
1985-10-17
CAN, MO, US, A
Dinamo & Delegacion Magdalena Contreras
Mexico
2580
19.42
Herrera-Arrieta
763
-99.13
78
79
1
Distrito Federal
[613,766,1375,1400]
P
Mexico
Miranda
78
79
1
Distrito Federal
1960-10-19
T
Mexico
19.2531
Desierto de Los Leones
7
-99.3286
78
79
MEXU-1110774
1
Guerrero
1976-01-25
M
Mexico
Tateoka
19.2533
Delegacion de Guajimalpa
7
-99.3308
Desierto de Los Leones
78
79
MEXU-151424, US-2380596
1
Distrito Federal
[1139,1276,1447,1472]
E
Mexico
Gonzalez
78
79
1
Distrito Federal
1979-11-02
ANSM, F
Mpio. Cuajimalpa
Mexico
2700
19.2533
Desierto de Los Leones
7
-99.3308
78
79
1
Distrito Federal
1928-07
ANSM, E
Mexico
Rio Grande
Santoyo
78
79
1
Distrito Federal
1951-12-09
E
Mexico
3500
19.42
Valle
763
-99.13
Lyonnet
78
79
MEXU-258403
1
Mexico
1951-10-07
E
Mexico
Matuda
3000
19.0494
Lago Zempoala
7
-99.3175
Valle
78
79
MEXU-499579, MEXU-499578
1
Mexico
1950-11-19
MEXU, E
Valle
Mexico
2600
19.2333
San Rafael
524
-98.75
Matuda
78
79
US-2079183
1
Mexico
[999,1433,1628,1653]
MEXU
Mexico
Matuda
78
79
US-2040890
1
Distrito Federal
1982-09-16
Y, ANSM
Herrera & Guerrero & Mpio. Chilpancingo & Bravos
1900
17.55
Omiltemi
767
-99.51
78
79
1
1991-11-15
V, NW
Mpio. Chilpancingo & Bravos & de Agua & de la Perra
Canada
2120
17.55
Nandi
5325
-99.5
78
79
MEXU-1089776
1
1985-10-10
R
Hidalgo & El Chico
Canada
20.2167
Ocampo
7
-98.7333
78
79
MEXU-502476
1
1929-07
P
Las Ventanas
Canada
2970
20.17
Lyonnet
761
-98.73
Mineral del Chico
78
79
US-1034151
1
1993-05-19
J, MEXU
Praxedes Perez & Pueblo Nuevo & El Chico
Canada
2850
20.1667
Pachuca Dist.
7
-98.7333
Real del Monte
78
79
1
1946-10-17
H, E, GH
Jalisco
Canada
Rio Mascota
Moore
Jr.
78
79
US-1963095
1
[236,1431,1988,2013]
N, S
Canada
2500
20.0403
La
7
-103.8753
78
79
1
1960-11-29
R
Magnolia Press
79
80
78
79
1
1960-11-02
NY, R
McVaugh
2175
19.1556
Tapalpa
7
-102.9889
Amacueca
79
80
1
1983-10-25
1992-04-08
1983-10-25
NY, A
de Toluca & Jaramilo & Nunez & Jaramillo
3260
19.14
McVaugh
763
-99.71
79
80
MEXU-1089780, MEXU- 1098071, MEXU-1033954
1
Mexico
[1338,1432,332,356]
A
Vega
79
80
1
Mexico
[151,311,368,392]
R
Ramirez
79
80
1
Mexico
1988-10-27
J, G
Malinalco-San Jose Teneria
18.93
Mendoza
764
-99.49
79
80
MEXU-1089771
1
Mendoza
MEXU, S
Guzman & Jaramillo & de Cabra
Aldape
2530
19.12
Cieneguillas de Cabra
763
-99.88
Toluca-Temascaltepec
79
80
1
Guerrero
[417,476,476,501]
1981-08-01
R
de Toluca
2600
19.1083
Nevado
7
-99.7583
79
80
1
Nevada
MEXU, S
Guzman
entre Texcaltitlan y Las Juntas
79
80
1
Nevada
[712,769,511,536]
1981-08-01
R
de Toluca
2660
18.93
Nevado
764
-99.95
79
80
1
Nevada
[257,1436,548,573]
1978-08-02
F, M
Guzman & Estacion
2500
San
79
80
MEXU-1110805
1
Nevada
[151,1432,583,608]
1977-12-11
N
Fuchs & Mpio. Amecameca & de San Antonio
2550
79
80
MEXU-231170
1
Nevada
[151,283,620,645]
M
Valentin
79
80
1
Nevada
1992-10-02
ANSM, A
19.08
Amecameca-Tlamacas
764
-98.67
Mpio. Amecameca
79
80
1
Nevada
[350,505,655,680]
G
Miranda
79
80
1
Nevada
[516,1436,655,680]
MEXU, W
Villegas & Volcan Popocatepetl
Mpio. Atlautla
79
80
1
Victoria
[151,853,692,717]
1987-10-24
J
Atlautla
2725
19.05
764
-98.64
79
80
1
Victoria
1976-09-19
E, C
Hernandez & Mpio.
2700
19.21
E de San Rafael
763
-98.74
79
80
MEXU-565906
1
Mexico
[1063,1167,727,752]
G
Flores
79
80
1
Mexico
1937-10-16
E
Mpio. & Tepeite
19.27
Morelos, Valle
763
-99.18
79
80
MEXU-292304
1
Morelos, Valle
1990-02-04
NW, O
Lyonnet & Gonzalez
2350
18.9833
Mpio. Ocuilan de Arteaga
7
-99.4417
79
80
MEXU-278684
1
Morelos, Valle
[394,514,836,861]
J
Miranda
79
80
1
Morelos, Valle
[524,1355,836,861]
1975-10-26
J, R
Mpio. Tepotzotlan
2400
79
80
MEXU-1031709
1
Morelos, Valle
1976-10-21
V, ANSM, S
Torres & el Aguaje
2670
19.41
Texcoco
763
-98.86
Texcoco
79
80
1
Morelos, Valle
1965-05-27
MO, E, J
D. Koch
2700
19.21
Mpio. Tlalmanalco
763
-98.76
79
80
MEXU-265511
1
Morelos, Valle
E, L, M
Rzedowski & Mpio. Tlalmanalco
2700
19.21
E de San Rafael
763
-98.76
79
80
MEXU- 187911
1
Morelos, Valle
1974-08
1974-12-31
1974-08
ANSM, N, NE, E
de Bravo & Ma.
2490
19.37
San
763
-100.15
Mpio. Villa de Allende
79
80
MEXU-220002
1
Valle
[1158,1335,1052,1077]
V
Maury Hdez.
79
80
1
Valle
S
Serrano Cardenas
79
80
1
Valle
1974-08
1974-12-31
1974-08
N, NE, S
Estacion & de Bravo
2490
19.33
San
0
-100.0
Mpio. Villa de Allende
79
80
MEXU-200087
1
Valle
1995-09-30
MEXU, H
Mpio. Villa
2200
18.84
Ixtapan de la Sal
764
-99.68
Tenango de Arista
79
80
1
Guerrero
1998-10-08
NW, A, G
Vibrans & Mpio. Zimatlan & Canada El Frijolon & de San Pedro El Alto
2790
19.61
NW de La Cofradia
5262
-99.9
79
80
MEXU-895630
1
Guerrero
[598,814,1268,1293]
M, O, L
Miranda
79
80
1
Guerrero
1978-09-29
W, H
Hernandez Martinez & Ayapango
2500
19.15
1 km W of Amecameca
555
-98.7833
79
80
MEXU-907764
1
Guerrero
[1258,1332,1303,1328]
J
Iltis
79
80
1
Guerrero
A
Doebley
79
80
1
Guerrero
2001-10-10
MO, US, N, P, M
Lasseigne
2490
19.071
1.4 mi N of Tequexquipan
55
-99.9388
79
80
MSC-277227, RSA-POM-320576
1
Guerrero
[669,820,1376,1400]
Y
Peterson
79
80
1
Guerrero
2007-10-10
CAN, MO, US, W, P, M
Herrera-Arrieta
2596
18.9268
1.9 mi W of Chiltepec on Hwy.
7
-99.8442
79
80
P-03216909
1
Guerrero
[1180,1339,1413,1436]
J, M
Peterson
79
80
1
Guerrero
M, J
Saarela
79
80
1
Guerrero
2001-10-10
CAN, MO, US, NE, P, M
Flores Villegas
2826
19.0317
SW of Volcan Toluca
7
-99.9102
79
80
P-03631106
1
San Miguel
[816,974,1484,1508]
Y
Peterson
79
80
1
San Miguel
1968-11-17
CAN, MO, US, NW, A
Herrera-Arrieta
2550
19.12
Cerro Tenayo
763
-98.76
79
80
P- 03216911
1
San Miguel
2007-10-11
E, P, M
Paso
2488
19.0706
6.7 mi E of Amecameca
7
-98.6923
79
80
MICH-1119227, MSC-223104
1
Cortes
[899,1064,1593,1616]
J, M
Peterson
79
80
1
Cortes
[1076,1250,1591,1616]
M, J
Saarela
79
80
1
Cortes
2001-10-09
CAN, MO, US, NE, P, M
Flores Villegas
3190
19.1858
Toluca
524
-99.86
79
80
1
Cortes
[411,571,1664,1688]
Y
Peterson
79
80
1
Cortes
2001-10-09
US, P, M
Herrera-Arrieta & La Comunidad
2607
19.1518
La
7
-99.9118
79
80
1
Cortes
[1082,1239,1700,1724]
Y
Peterson
79
80
1
Cortes
1968-10-21
CAN, MO, US, GH, NE, J
Herrera-Arrieta
2550
19.12
Amecameca
763
-98.76
Coulter
79
80
P-03216910
1
Cortes
1966-10-09
J
Rzedowski & de San Pedro Nexapa
19.08
Amecameca
764
-98.74
79
80
MICH-1119223, MSC-233266
1
Cortes
E
Rzedowski & Mpio. Amecameca & Zumpango & Paso
Cortes
79
80
MSC-217972
1
Cortes
[1170,1230,1844,1869]
1976-11-20
S
Amecameca-Cuautla & Mor.
3300
19.06
5279
-98.7
79
80
1
Cortes
1976-07-29
B, E, S
D. Koch & de la Marquesa
2600
19.3833
Naucalpan
7
-99.3333
Mpio. Huixquilucan
79
80
US- 2824581
1
Mexico
1976-10-21
CHAPA, MO, US, S
D. Koch
2670
19.42
Texcoco
763
-98.83
Texcoco
79
80
MEXU-270647
1
Mexico
1965-08-29
K, E, S
Roe & Mori & Comunidad
3000
19.1292
Temascaltepec Dist.
7
-99.9347
81
82
MSC-229117
1
1932-11-25
G, B, NY
Hinton & Temascaltepec & Crucero
2600
19.04
764
-100.23
81
82
US-1867841
1
1935-10-28
G, B, MO, NY, E
Tlalmanalco
19.21
Hinton
763
-98.74
81
82
RSA-POM- 364158
1
1976-09-19
C, G, MO
Indonesia
2700
Volcan de Toluca
81
82
SD-110652
1
1953-10-18
E, R, W
Indonesia
Sohns
19.0833
Nevado de Toluca
7
-99.8333
Matuda
81
82
US-2118910
1
1965-08-29
K, E, S, W
Roe & Mori
Indonesia
3000
19.11
Nevado de Toluca
763
-99.79
81
82
US-3115415
1
1965-08-29
K, E, S, GH, W
Roe & Mori & Cerro Tancitaro & de El Tepetate
Indonesia
3000
19.41
W de Uruapan
5268
-102.3
81
82
1
Michoacan de Ocampo
1996-08-29
I, N
Las Peras
Indonesia
2850
19.78
N de Indaparapeo
762
-100.97
81
82
MICH- 1119229
1
Michoacan de Ocampo
1987-11-08
J, E
Rzedowski
Mexico
2200
19.67
37 mi E of Morelia
763
-100.73
81
82
MEXU-809934
1
1972-10-20
L, H, J, T, NW
Harvey & Witherspoon & Santa Clara
Mexico
2438
19.7364
5.7 mi NW of Huajumbaro
522
-100.78
81
82
MICH-1119244
1
2001-10-08
P, M, CAN, MO, N
Peterson & Carapan
United States of America
2430
19.41
N de Uruapan
763
-102.06
81
82
1
1980-08-22
J, C, MEXU
Soto Nunez & Villa Escalante & Ario
United States of America
2075
19.4
5580
-101.65
81
82
1
1949-11-11
H, E, NW, C
Moore & Jr. & Hidalgo
United States of America
2340
19.66
763
-100.71
81
82
US-1983697
1
1950-07-04
H, K, S
Wagnon
United States of America
19.31
18 mi S of Patzcuaro
763
-101.66
81
82
MEXU-772510, US-2154832
1
1961-11-20
1961-11-25
1961-11-20
R, M, T, NY, NW
King & Soderstrom
United States of America
2728
19.3806
Patzcuaro
524
-101.65
81
82
MICH-1119226, MEXU- 55441, US-2378973
1
2001-10-08
P, M, CAN, MO
Peterson & Cerro Azul
United States of America
2350
19.7167
Morelia
7
-101.1833
81
82
P- 03216912
1
1909-11-04
1950-07-04
1909-11-04
G, MO, NW, H, K
Botanical Gardens & University & Ann Arbor & Wagnon & Mesa Karitsho & Padre
United States of America
2300
19.57
Ciudad Hidalgo
763
-101.81
Arsene
81
82
US-1002650, MICH- 1119225
1
Michigan
1978-11-09
J, C, MEXU
Coballero & Mapes & Mpio. Charo
United States of America
2625
19.65
El Salto de Agua
555
-100.9444
81
82
1
Michigan
1994-02-01
J, A, W
Torres Estrada
United States of America
19.7
Morelia
5580
-101.18
81
82
MEXU-1009115, MEXU-1089785
1
Michigan
1986-10-25
J, W
Rzedowski & Mpio. Patzcuaro
United States of America
2300
19.415
Cerro del Burro
55
-101.5006
81
82
MEXU-658295
1
Michigan
1986-11-23
J, ANSM, N
Rzedowski & Mpio. Querendaro
United States of America
2900
N de Real de Otzumatlan
81
82
1
Michigan
1986-11-20
J, ANSM, E
Rzedowski & Mpio. Senguio & Cerro Ross Azul
United States of America
2350
19.73
E de Chincus
762
-100.35
81
82
1
Michigan
1986-11-23
J
Santos Martinez & Mpio. Zacapu & de Zacapu & Zamora
United States of America
2800
81
82
MICH-1119219
1
Michigan
1988-09-09
A, ANSM
Grimaldo Nunez & Mpio. Zinapecuaro
United States of America
2000
SSE de Jerahuaro
81
82
1
Michigan
1987-12-07
J, ANSM, E, S
Rzedowski
United States of America
2550
19.41
San Juan Nuevo
763
-102.06
81
82
MEXU-915763
1
Michigan
1909-11-04
1961-10-15
1909-11-04
R, M, T, G, GH, INIF
Arsene & Campo Experimental
United States of America
King
19.69
Morelia
763
-101.16
Soderstrom
81
82
MEXU-55372
1
Michigan
1976-10-09
1986-12-04
1976-10-09
C, L, ANSM, J, E
Ortiz & Rosario
United States of America
Dias Luna
2100
19.58
Ocampo
763
-100.34
Patzcuaro-Uruapan
81
82
MEXU-638476
1
Michigan
1989-11-02
R, M, S
Torres & Ramirez & Mpio. Patzcuaro & Opopeo
United States of America
2400
19.5158
S de Patzcuaro
7
-101.6094
81
82
MEXU-904824
1
Michigan
1985-11-10
J
Mpio. Patzcuaro & Cerro El Frijol
United States of America
2400
19.5158
7
-101.6094
81
82
MEXU-993903
1
Michigan
1985-10-31
H
Diaz Barriga & Morelos & Curenava-Mexico
United States of America
2700
19.27
763
-99.17
81
82
MEXU-662626
1
Michigan
1937-10-15
1984-02-08
1937-10-15
E
Lyonnet
United States of America
Manrique
2460
19.27
Valle del Tepeite
763
-99.25
Beetle
81
82
MEXU-1110847, MEXU-278689
1
Michigan
1937-10-16
E, J, MEXU, N
Oaxaca & Hwy.
United States of America
16.1157
Elcoro
7
-96.4745
Lyonnet
81
82
US-1746382
1
Michigan
2008-09-23
P, M, J, CAN, N
Peterson & Saarela & Santa Maria Yavesia
United States of America
2722
17.1966
9.5 mi N of San Cualimojoyas
7
-96.4512
81
82
1
2008-09-20
P, M, J, CAN
Peterson & Saarela
United States of America
2727
16.09
Campamento Rio de Molino
769
-96.47
81
82
1
1965-09-21
J
Rzedowski
United States of America
2250
17.1
Distrito del Centro
5580
-96.85
Cerro de San Felipe
81
82
MICH-1119224, MICH-1119220, MSC-210582
1
1920-10-12
C
Mpio. Zimatlan & Comunidad El Tlacuache & San Pedro El Alto
2300
16.04
769
-96.47
82
83
MEXU-6111, US-1014111
1
1998-10-09
A, G, O, M, E
San
2420
16.1698
Miranda
7
-96.4898
Hernandez
82
83
MEXU-901092
1
2008-09-22
P, M, J, CAN
Peterson & Saarela & Puebla
United States of America
2641
US
82
83
1
United States of America
19.0897
Puebla in Parque Nacional Izta-Popo
7
-98.6064
82
83
1
2001-10-12
P, M, Y, E
Peterson & Herrera-Arrieta & Puebla & Parque Nacional Izta-Popo
United States of America
3330
19.085
24.3 mi E of Amecameca
5250
-98.6
82
83
1
2001-10-13
P, M, Y, CAN
Peterson & Herrera-Arrieta & Veracruz & Orizaba
United States of America
3316
19.0167
US
525
-97.22
82
83
1
[502,888,404,429]
US
United States of America
82
83
US-2473642
1