Taxonomic revision of the Oriental species of Notoxus (Coleoptera: Anthicidae) Kejval, Zbyněk Acta Entomologica Musei Nationalis Pragae 2011 2011-12-09 51 2 627 673 Kejval, 2011 Kejval 2011 [487,699,442,467] Insecta Anthicidae Notoxus Animalia Coleoptera 22 649 Arthropoda species impavidus sp. nov.  ( Figs. 22, 53, 73)    Typelocality. Laos, Vientiane Province, Phou Khao Khouay National Park, Tad Leuk, 18°23′N, 103°04′E, 150–200 ma.s.l.  Type material.  HOLOTYPE: J, ‘ LAOScentral, Saisombunzone PHOU KHAO KHOUAYNat. Park TAD LEUK 18°23′N, 103°04′E  15.–21.V.2001, alt.  150–200 m E. Jendek& O. Šaušaleg.’ ( NMPC).  PARATYPES: 16 JJ 3 ♀♀, same data as holotype ( ZKDC, 1 spec.each in BMNH, DCDC, MHNG, MNHN, NHMB, ZMHB); 1 J, ‘ LAOScentr., Khammouanpr., BAN KHOUN NGEUNvill. env.,  17.v.–6.vi.2007, ~  300 m, M. Štrbaleg.’ ( ZKDC); 8 JJ, ‘C-LAOS: Prov. Viangchan Phou Khao Khouay NP Nam Leukenv. Tad Leuk Waterf. // 1.-  8.6.1996,  200mat light leg. Schillhammer(15a) // Notoxus inbasalis Picdet. G. Uhmann1998’ ( NHMW, ZSMC).   Description. Male( holotype). Body length 3.5 mm. Head and pronotum reddish brown. Elytra brown, with reddish tinge basally, with indication of two vaguely outlined, transverse, pale reddish bands. Legs and antennae reddish. Body only moderately glossy, rather finely, densely and evenly punctured; pronotum with some scattered coarser punctures; punctures of elytra rather fine but distinct ( Fig. 73), slightly coarser than those on pronotum, simple, with smooth margins. Body setation mostly subdecumbent; pronotum with numerous conspicuously long, erect setae, especially anterolaterally; elytra with numerous, moderately longer suberect to erect setae. Antebasal setose band of pronotum conspicuous, widely interrupted medially. Head with rather large eyes. Antennae long, only moderately enlarged in terminal third. Pronotum 1.1 times as wide as head across eyes, globose in dorsal view. Pronotal horn moderately long, slender, largely subparallel, with continuously fringed apical margins and 3 lateral lobules on each side, becoming distinctly smaller towards horn base; horn crest only moderately raised, evenly lowering towards horn apex, its lateral margins feebly marked by series of somewhat coarser, distinctly separate lobules / rugules; dorsal surface rather evenly and densely covered with distinct rounded rugules. Elytra 1.9 times as long as wide; humeri distinctly protruding; omoplates and postbasal moderately indicated; elytral apices distinctly modified, subtruncate, with rounded gland opening near distinctly produced lateral angle. Legs simple. Abdominal sternum VII moderately impressed postero-medially, its posterior margin somewhat unevenly rounded. Tergum VIII forming simple, evenly arched sclerite. Aedeagus as figured ( Fig. 22).   Female.Externally differing from male by simply rounded elytral apices.  Variation.Body length (J ♀) 3.2–3.6 mm. Moderately variable in width of pronotal horn; its lateral margins with 2–4 distinct lobules on each side.   Differential diagnosis.  Notoxus impavidus sp. nov.shares long bifurcate dorsal plate of the tegmen of the aedeagus with  N. alesand  N. garuda sp. nov., but differs by somewhat paler body colouration (dark markings of elytra less distinct, vaguely outlined), rather fine and dense punctation of the elytra ( Fig. 73), and by the morphology of the aedeagus.   Etymology.From the Latin,  impavidus(fearless, intrepid); named in reference to the location of the typelocality in the former Saisomboun special zone, administered by military forces.   Distribution. Laos. 2001-05-15 2001-05-21 2001-05-15 PHOU, KHAO, KHOUAY, LEUK, NMPC E. Jendek & O. Sausa Laos 175 18.383333 Saisombun 1275 103.066666 22 649 1 holotype 2001-05-15 2001-05-21 2001-05-15 ZKDC, BMNH, DCDC, MHNG, MNHN, NHMB, ZMHB E. Jendek & O. Sausa Laos 175 18.383333 Saisombun 1275 103.066666 22 649 4 3 paratype 2007-05-17 2007-06-06 2007-05-17 KHOUN, NGEUN, ZKDC M. Strba Laos 300 Khammouan 22 649 1 Khammouan paratype 1996-06-08 NHMW, ZSMC Phou Khao Khouay NP Nam Leuk & Schillhammer Laos G. Uhmann 200 Viangchan 22 649 1 Viangchan paratype