Studies of the genus Anthelephila Hope (Coleoptera: Anthicidae) 14. Twenty-four new species from Asia and new records of A. fossicollis Kejval Kejval, Zbyněk Zootaxa 2017 4306 1 1 52 Kejval, 2017 Kejval 2017 [151,400,151,177] Insecta Anthicidae Anthelephila Animalia Coleoptera 10 11 Arthropoda species arcana sp. nov.   Description.Male ( holotype). Body length 4.1 mm. Head and pronotum reddish; elytra brownish black, with slight bluish reflection; legs reddish-brown, tibiae and distal part of femora darker, antennae dark reddish, distal antennomeres nearly brownish black. Head 1.1 times as long as wide, differentiated from short neck; tempora strongly narrowing posteriad, posterior angles absent. Eyes medium-sized, rather convex. Dorsal surface moderately glossy, distinctly punctate and partly corrugated (especially medially to eyes), with indication of smooth, impunctate median strip posteriorly; punctures distinctly separated. Setation short, subdecumbent; few tactile setae. Antennae moderately enlarged in terminal third; antennomere X 1.6 times, XI 2.4 times as long as wide. Pronotum 1.5 times as long as wide, distinctly narrower than head including eyes, nearly evenly rounded anteriorly, strongly narrowed and moderately impressed (constricted) postero-laterally in dorsal view; pronotal disc evenly convex in lateral view. Surface rather glossy; antero-lateral sides minutely and sparsely punctate, nearly impunctate near procoxal cavities; disc more distinctly and densely punctate posteriorly, with several transverse wrinkles before smooth antebasal area; postero-lateral impressions shortly and rather finely, longitudinally wrinkled, adjacent dorso-lateral surface rugose. Setation as on head. Mesosternum with slight indication of median longitudinal bulge. Metasternum simple. Elytra 1.8 times as long as wide, conjointly rounded apically; humeri distinct; postscutellar impression absent. Surface glossy, distinctly punctate; punctation indistinctly double, setiferous punctures rather widely separated. Setation nearly evenly developed, sparser near base, generally longer and more raised than on head, decumbent; numerous tactile setae. Metathoracic wings fully developed. Fore legs modified ( Fig. 20); profemora with short tooth-like process, rounded apically, with short fringe of stiff setae subapically; protibiae nearly simple, moderately swollen and with small protrusion on inner side at about mid-length; penultimate tarsomere widened/flattened distally, with terminal tarsomere articulated dorsally near base in all tarsi. Setation normally developed; pro- and mesotibiae with two, metatibiae with single distinct terminal spur. Abdominal sternum III with paired submedian, longitudinal protrusions at base. Sternum VII shallowly emarginate and long, densely setose ( Fig. 21). Tergum VII simple. Sternite VIII ( Fig. 22); paired prongs moderately lobed and with long slender projection medially. Tergite VIII with paired sclerites somewhat unevenly rounded apically, narrowly connected medially. Terminal sclerite of segment IX (spiculum) membranous, widely rounded and setose apically. Aedeagus ( Fig. 23); apical portion of tegmen 0.4 times as long as basal-piece, trilobed apically. Female. Externally differing from male as follows: fore legs and abdominal sterna III, VII simple; both sternum and tergum VII rather strongly produced and narrowly rounded apically.  Variation.Body length (♂♀) 3.6–4.5 mm.     Typematerial. Holotype, ♂: LAOSnorth,  5–11.V.1997, 20 km NW Louang Namtha, N 21°09.2, E 101°18.7, alt. 900±  100 m, E. Jendek& O. Šausaleg. [p] ( NMPC).   Paratypes: 3 ♂♂, same data as holotype ( ZKDC);  2 ♂♂, same data, except: M. Štrba& R. Hergovitsleg. ( ZKDC);  2 ♂♂, LAOSnorth,  13–24.v.1997, 15 km NW Louang Namtha, N21°07.5, E 101°21.0, alt. 750±  100 m, E. Jendek& O. Šausaleg. [p] ( ZKDC);  2 ♂♂, north LAOS, 20 kmNW of Louang Namtha, 21°09.2N/ 101°18.7E(GPS),  900–1100 m,  5.–30.v.1997 C. Holzschuhleg. [p] ( ZKDC);  2 ♂♂, LAOS,  24–30.iv.1999,  Louangphrabangpr., 20°33–4′ N 102°14′E, BanSongCha ( 5 kmW),  1200m, Vít Kubáňleg. ( NHMB);  3 ♂♂, LAOS-N,  23.iv.1999  Louangphrabangpr., 20°42′N 102°54′E,  25kmE Muang Ngoy,  1000m, Vít Kubáňleg. [p] ( NHMB);  7 ♂♂, LAOS, Louangnamthapr. 21°09′N 101°19′E, Namtha+ Muang Sing,  5–31. v. 1997, 900–  1200 m Vít Kubáňleg. [p] // Anthelephilasp. det. D.Telnov, 2001 [p] ( NHMB);  2 ♀♀, NW LAOS,  65 kmNW of Luang Nam Tha, Nam Tha NPA,  1050m,  8.–15.iv.2010, S. Murzinlgt. ( ZKDC);  1 ♂, NW LAOS,  5 kmSW Muang Sing, Chiang Tung( Stupa) GH,  750 m,  9.–14.vi.2010, S. Murzinlgt. ( ZKDC);  1 ♂, CHINA, Yunnan, Mengla, Mohan, hillside, 21°11′14.2′′N 101°41′0.2′′E, [p; including locality names in Chinese] //  910 m, swept,  3.VIII.2012, leg. Dávid Rédei[p] ( HNHM).   FIGURES 20–23.  Anthelephila arcana  sp. nov.(holotype ♂). Profemur and tibia (20); abdominal sternum VII (21); sternite VIII, dorsal view (22); apical portion of aedeagus (23). Scale (0.5 mm): A—Fig. 20; (0.3 mm): B—Figs 21–23.   Additional specimens. Laos: 5 ♀♀, same data as holotype ( ZKDC);  1 ♀, same data, except: M. Štrba& R. Hergovitslgt. ( ZKDC);  2 ♀♀, Luang Nam Thaprov., 20 kmNW of Luang Namtha, 21°09.2N 101°18.7E(GPS),  900–1100 m,  5.–30.v.1997, C. Holzschuhlgt. ( ZKDC);  5 ♀♀, Luang Prabangprov., 5 kmW of Ban Song Cha, 20°33–4′ N 102°14′E,  1200 m,  1.–9.v.1999, V. Kubáňleg. ( NHMB);  5 ♂♂, Luang Nam Thaprov., Namtha+ Muang Sing, 21°09′N 101°19′E,  5.–31.v.1997, 900–  1200 m, V. Kubáňlgt. ( NHMB);  1 ♀, Luang Prabangprov, 25 kmE of Muang Ngoy, 20°42′N 102°54′E,  1000 m,  23.iv.1999, V. Kubáňleg. ( NHMB);  3 ♀♀, 65 kmNW of Luang Nam Tha, Nam Tha NPA,  1050 m,  8.–15.iv.2010, S. Murzinlgt. ( ZKDC).   Differential diagnosis.  Anthelephila arcana  sp. nov.belongs to the  A. fossicollisspecies-group. It may resemble  A. davita  sp. nov.by the shape of the male protibiae and sternum VII, but differs clearly in the morphology of the paired prongs (cf. Figs 22 versus48). Its females seem to be well-characterized by distinctly produced terminalia.   Etymology.From the Latin arcanus(secret, withdrawn), refering to the existence of numerous externally similar species.   Distribution. China( Yunnan), Laos. 1586033219 1997-05-05 1997-05-11 1997-05-05 NMPC Jendek & O. Sausa Laos 100 21.153334 Louang Namtha 126 101.31167 10 11 1 1 Louang Namtha holotype 1586033228 [199,774,1554,1579] 1997-05-05 1997-05-11 1997-05-05 ZKDC Jendek & O. Sausa Laos 100 21.153334 Louang Namtha 126 101.31167 10 11 3 3 Louang Namtha paratype 1586033203 1997-05-05 1997-05-11 1997-05-05 ZKDC R. Hergovits Laos 100 21.153334 Strba 126 101.31167 10 11 2 2 Louang Namtha paratype 1586033214 1997-05-13 1997-05-24 1997-05-13 ZKDC NW Louang Namtha, N & Jendek & O. Sausa Laos 100 21.125 Louang Namtha 126 101.35 10 11 2 2 Louang Namtha paratype 1586033229 1997-05-05 1997-05-30 1997-05-05 ZKDC C. Holzschuh Laos 1000 21.153334 Louang Namtha 126 101.31167 10 11 2 2 Louang Namtha paratype 1586033255 1999-04-24 1999-04-30 1999-04-24 NHMB Louangphrabang & Vit Kuban Laos 1200 102.23333 Louangphrabang 946 102.23333 10 11 2 2 Louangphrabang paratype 1586033202 1999-04-23 LAOS-N, NHMB Louangphrabang & Vit Kuban Laos 1000 20.7 25 km E Muang Ngoy 1266 102.9 10 11 3 3 Louangphrabang paratype 1586033254 1997-05-05 1997-05-31 1997-05-05 NHMB Namtha & Muang Sing & Vit Kuban Laos D. Telnov 1200 21.15 Louangnamtha 1265 101.316666 10 11 7 7 Louangnamtha paratype 1586033223 2010-04-08 2010-04-15 2010-04-08 NPA, ZKDC S. Murzin Laos 1050 Nam Tha 10 11 2 2 Luang Nam Tha paratype 1586033206 2010-06-09 2010-06-14 2010-06-09 ZKDC S. Murzin Laos 750 Stupa Chiang Tung 10 11 1 1 paratype 1586033230 2012-08-03 HNHM Chinese & David Redei China 910 21.187277 Mohan 1 101.68339 Mengla 10 11 1 1 Yunnan paratype 1586033193 [199,983,151,176] 1997-05-05 1997-05-11 1997-05-05 ZKDC Jendek & O. Sausa Laos 100 21.153334 Louang Namtha 126 101.31167 12 13 5 5 Louang Namtha 1586033218 1997-05-05 1997-05-11 1997-05-05 ZKDC M. Strba & R. Hergovits Laos 100 21.153334 Louang Namtha 126 101.31167 12 13 1 1 Louang Namtha 1586033247 1997-05-05 1997-05-30 1997-05-05 ZKDC C. Holzschuh Laos 1000 21.153334 Luang Nam Tha prov. 126 101.31167 12 13 2 2 Nam Tha 1586033256 1999-05-01 1999-05-09 1999-05-01 NHMB V. Kuban Laos 1200 102.23333 Ban Song Cha 946 102.23333 12 13 5 5 Luang Prabang 1586033205 1997-05-05 1997-05-31 1997-05-05 NHMB V. Kuban Laos 1200 21.15 Muang Sing 1265 101.316666 Namtha 12 13 5 5 Luang Nam Tha 1586033212 1999-04-23 NHMB V. Kuban Laos 1000 20.7 Muang Ngoy 1266 102.9 12 13 1 1 Luang Prabang 1586033242 2010-04-08 2010-04-15 2010-04-08 NPA, ZKDC S. Murzin Laos 1050 Nam Tha 12 13 3 3 Luang Nam Tha