Falagria nigerrima BERNHAUER, 1908: 38 Revision of the Anaulacaspis species of the Palaearctic region (Coleoptera: Staphylinidae: Aleocharinae) Assing, Volker Beiträge Zur Entomologie = Contributions to Entomology 2016 2016-12-20 66 2 201 255 DJK9 (BERNHAUER, 1908) Bernhauer 1908 [227,674,1075,1102] Insecta Staphylinidae Anaulacaspis Animalia Coleoptera 28 229 Arthropoda species nigerrima  ( Figs 57–61, Map 3)      Falagria nigerrima BERNHAUER, 1908: 38f.   Type material examined:  Lectotype ♂, present designation [dissected prior to present study]: “♂ / Buchara /  nigerrimaBh. Cotypus/ Chicago NHMus, M. Bernhauer Collection / Syntypus  Falagria nigerrimaBernhauer,V.I. Gusarov rev. 2015 /  Anaulacaspis nigerrima(Bh), V.I. Gusarov det. 2015 / Lectotypus ♂  Falagria nigerrimaBernhauer, desig. V. Assing 2016 /  Anaulacaspis nigerrima(Bernhauer), det. V. Assing2016” ( FMNH).  Paralectotypes: 1 ♂: “♂ / 124. / Buchara /  laevigataEpp., Knauth/  nigerrimaBh./  nigerrimaBrh.Typus” ( FMNH); 1 ♂: “♂ / Margelan /  nigerrimaBh. Cotypus” ( FMNH); 1 ♂: “♂ / Sibiria or., Reitter /  nigerrimaBh., det. Bernh.” ( FMNH); 2 ♀♀: “Turcmenien. Reitter. Leder. /  nigerrimaBrh. Cotypus” ( FMNH).All paralectotypesdissected prior to present study and additionally with the following labels: “Chicago NHMus, M.Bernhauer Collection / Syntypus  Falagria nigerrimaBernhauer, V.I. Gusarovrev. 2015 /  Anaulacaspis nigerrima(Bh), V.I. Gusarov det. 2015 / Paralectotypus  Falagria nigerrimaBernhauer, desig. V. Assing 2016 /  Anaulacaspis nigerrima(Bernhauer), det. V. Assing 2016”.   Comment: The original description is based on an unspecified number of syntypesfrom “Buchara, Margelan, Turkmenien, Turkestan (Mts. Ghissar) und Ostsibirien” ( BERNHAUER 1908). Six syntypes, four malesand two femalesare deposited in the Bernhauer collection at the FMNH. One of the males from “Buchara” is designated as the lectotype.  Additional material examined:   Iran:  Razavi Khorasan: 4 ♂♂, 2 ♀♀,  20 kmNW Torbat-e, Heydariyeh, Senobar, 35°26'N, 59°06'E,  1730 m,  28.V.2006, leg. Frisch& Serri( MNB, cAss).     Afghanistan: 1 ♀, N Herat, SW Khouchk, Tchaouni Khoch Robat,  29.VI.1959, leg. Lindberg( NHMW).    Turkmenistan: 10 exs., locality not specified [“Turcmenien”, “ Turkestan”], leg. Reitter( MHNG, MNB, NHMW, cAss).   Map 3: Distributions of  Anaulacaspis formosa(black circles),  A. elegans(white circles),  A. nigrina(black diamonds),  A. eminens(white diamonds), and  A. nigerrima(stars) in the West Palaearctic and Middle Asia, based on revised records.    Kyrgyzstan: 2 ♂♂, 2 ♀♀, Batken, S Ay-Kol, Teo-Jailoo Valley, Tegir-Malik River, 39°43'N, 69°42'E,  2100 m,  28.VI.2012, leg. Frisch( MNB, cAss); 2 ♂♂, 1 ♀, Batken, Isfana – Isfara, WSW Ozgorush, Turkestan range, Lyailek River, 39°44'N, 70°00'E,  1520 m,  22.VI.2012, leg. Frisch( MNB, cAss); 2 ♂♂, 1 ♀, Batken, Isfana – Isfara, WSW Ozgorush, tributary of Lyailek River, 39°44'N, 70°00'E,  1560 m,  23.VI.2012, leg. Frisch( MNB);  1 ex., Ferghansky Alatau, Toskool-Ata,  1.VIII.2002( MNB); 1 ex., Kyrghyzky Alatau, foothills, O.to-Say,  1000 m( MNB).    Uzbekistan: 11 exs., Taškent, leg. Leder, Reiter, Faust, Staudinger( MNB, NHMW, cAss);  3 exs., Buchara( MHNG, MNB).    Tajikistan: 1 ♂, Varzob district, Varzob, Campus‘ Chaika’, 38°46'N, 68°49'E,  1110 m,  7.–8.VII.2012, leg. Barševskis(cAss);  1 ex., Varzob district,  48 kmN Dushanbe, Khodja Obi Gharm,  1900 m,  3.–4.VII.2012, leg. Barševskis(cAss);  37 exs., Hissar-Alai, Adjukcleft near Varzob,  1200 m, stream bank,  1.–3.VII.1990, leg. Schülke, Wrase( MNB, cAss, cWun).  Country not specified:  6 exs., “ Transkaspien”, leg. Reitter( NHMW).   Redescription: External characters as in  A. nigra, except as follows: Whole body black including antennae black, legs blackish brown (old material usually paler, with the pronotum and elytra brown and the legs pale-brown). Head and pronotum with very fine and rather sparse punctation, without distinct sexual dimorphism. Eyes of somewhat variable length, 0.9–1.1 times as long as pronotum. Posterior margin of tergite VIII broadly concave ( Fig. 57). ♂: sternite VIII ( Fig. 58) obtusely pointed posteriorly; median lobe of aedeagus ( Figs 59–61) 0.27–0.30 mmlong, ventral process apically acute (both in ventral and in lateral view), distinctly sinuate, and without membranous median projection at base (lateral view).   Distribution and natural history:  Anaulacaspis nigerrimais widespread and apparently not uncommon in Middle Asia (Northeast Iran, Turkmenistan, Kyrgyzstan, Uzbekistan, Tajikistan, Afghanistan) ( Map 3). One paralectotypewas collected in an unspecified locality in Siberia (“Siberia or.”). The altitudes indicated on the labels range from 1000 to 2100 m. At least some of the specimens were collected on the banks of rivers and streams. 3449483706 FMNH M. Bernhauer Collection & V. I. Gusarov & V. Assing V. Assing Falagria nigerrima Bernhauer Buchara / nigerrima Bh. Cotypus / Chicago 28 229 1 lectotype 3455129309 28 229 paralectotype 3455129323 [831,1436,425,449] 28 229 3455129319 28 229 3455129316 28 229 3449483339 2006-05-28 MNB Frisch & Serri Iran 1730 35.433334 Senobar 1193 59.1 Heydariyeh 28 229 1 Razavi Khorasan 3449483637 1959-06-29 NHMW SW Khouchk & Lindberg Afghanistan Tchaouni Khoch Robat 28 229 1 Herat 3449483347 MHNG, MNB, NHMW, cAss Reitter Turkmenistan Turkestan 28 229 10 3449483661 2002-08-01 2012-06-28 2002-08-01 MNB S Ay-Kol & Frisch Kyrgyzstan Lyailek River 2100 39.733334 Batken 1167 70.0 Lyailek River 29 230 2 3449483496 2002-08-01 2012-06-28 2002-08-01 MNB S Ay-Kol & Frisch Kyrgyzstan Lyailek River 2100 39.733334 Batken 1167 70.0 Lyailek River 29 230 2 3449483455 2002-08-01 2012-06-28 2002-08-01 MNB S Ay-Kol & Frisch Kyrgyzstan Lyailek River 2100 39.733334 Batken 1167 70.0 Lyailek River 29 230 2 3449483557 2002-08-01 2012-06-28 2002-08-01 MNB S Ay-Kol & Frisch Kyrgyzstan Lyailek River 2100 39.733334 Batken 1167 70.0 Lyailek River 29 230 2 3449483731 MNB, NHMW Uzbekistan Uzbekistan 29 230 11 3449483361 MHNG, MNB Buchara Uzbekistan Buchara 29 230 3 3449483578 2012-07-07 2012-07-08 2012-07-07 Barsevskis Tajikistan Varzob district 1110 38.766666 Campus 1173 68.816666 Varzob 29 230 1 3449483705 2012-07-03 2012-07-04 2012-07-03 Barsevskis Tajikistan 1900 Khodja Obi Gharm Varzob district 29 230 1 Dushanbe 3449483409 1990-07-01 1990-07-03 1990-07-01 MNB Schulke & Wrase Tajikistan Hissar-Alai 1200 Varzob Adjuk 29 230 37 3449483406 NHMW Reitter Transkaspien 29 230 6