Systematic revision of American Glaresidae (Coleoptera: Scarabaeoidea)
Gordon, Robert D.
Hanley, Guy A.
Insecta Mundi
2014
2014-01-10
2014
333
1
91
4NT7
Erichson
1848
[422,535,515,539]
Insecta
Glaresidae
Glaresis
Animalia
Coleoptera
32
31
Arthropoda
genus
1. Mesotibial apex strongly projecting laterally, projection apically acute ( Fig. 34C); metatibia abruptly widened before apex; Mexico, Central America; subgroup C ................................... 12 — Mesotibial apex weakly projecting laterally, projection rounded ( Fig. 29D) (except G. confusa , G. falli ,and G. texanawith apex strongly projecting, apically acute); metatibia gradually widened from base to apex ( Fig. 19G); Mexico, Canada, United States; subgroups A, B ..................... 2 2(1). Metasternal groove shallow, at least inner border of groove tapered ( Fig. 19D); subgroup A .. 3 — Metasternal groove deep, distinct, sides abrupt, ridged ( Fig. 27D); subgroup B ....................... 7 3(2). Outer margin of metatibia with series of small, regular teeth from base to apex, without large posteromedian lateral projection ( Fig. 21F); Texas................................... G. texanaGordon — Outer margin of metatibia with distinct posteromedian lateral projecton ( Fig. 20F); not known from Texas.................................................................................................................................. 4 4(3). Clypeal apex with large, widely spaced tubercles medially ( Fig. 23B); metatibial posteromedian lateral projection large, apically bifid ( Fig. 23F) ............................................... G. falli , n. sp. — Clypeal apex with small, evenly spaced tubercles throughout ( Fig. 22B); metatibial posteromedian lateral projection small, tooth—like ( Fig. 22F) ........................................................................ 5 5(4). Tubercles on head surface small, sparse ( Fig. 22B); apex of mesotibia strongly projecting, apically acute; Alberta, Colorado, Utah................................................................... G. confusaBrown — Tubercles on head large, dense ( Fig. 19B); apex of mesotibia not strongly projecting, apically rounded ....................................................................................................................................... 6 6(5). Posterosuperior metatrochanteral surface with 2 teeth ( Fig. 19C); median lobe of male genitalia wide, apex broadly rounded ( Fig. 19E) ..................................................... G. california , n. sp. —Posteriosuperior metatrochanteral surface with single tooth ( Fig. 20C); median lobe of male genitalia slender, apically acute ( Fig. 20E) ............................................. G. howdeniGordon 7(2). Posterosuperior metatrochanteral surface with 2 teeth ( Fig. 24G) ............................................ 8 — Posterosuperior metatrochanteral surface with single tooth ( Fig. 27G) .................................. 10 8(7). Metasternal sculpture with carinae straight, not irregular, pronotal carinae distinctly spaced, not dense; male genitalia with basal lobe slender, narrowed in basal 1/3, apex rounded ( Fig. 26F); southern California............................................................................ G. bautista, n. sp. — Metasternal sculpture with carinae irregular, dense; pronotal carinae closely spaced; male genitalia not as described for G. bautista; not known from southern California................................... 9 9(8). Metatasternal sculpture irregular, forming reticulate pattern extended from apex to metacoxal cavity of metasternum ( Fig. 24D); apical trochanteral margin with small teeth, teeth often not visible; male genitalia with basal lobe broad, apically rounded ( Fig. 24E); Arizona, California, Mexico............................................................................................................ G. mendicaHorn — Metasternalsculpture irregular, not forming reticulate pattern, not extended to metacoxal cavity ( Fig. 25D); apical trochanteral margin with several distinct teeth; male genitalia with basal lobe narrow, apically acute ( Fig. 25E); California, Idaho, Nevada, Wyoming........................... ............................................................................................................. G. paramendica, n. sp. 10(7). Posteromedian lateral projection of metatibia large, pronounced ( Fig. 29F); surface sculpture of frons and vertex composed of dense raised, irregular ridges coalesced into a reticulate pattern ( Fig. 29B); Baja California Sur................................................................. G. australis. n. sp. — Posteromedian lateral projection of metatibia small, not pronounced ( Fig.20F); surface sculpture of frons and vertex reduced, if ridges present then not forming a reticulate pattern; United States......................................................................................................................................... 11 11(10). Metasternum tumid, protuberant, with heavy, somewhat reticulate surface sculpture laterally ( Fig. 30D); southeast Texas........................................................................... G. tumida, n. sp. — Metasternum flat or medially concave, surface sculpture comparatively light, not forming reticulate pattern; not known from southeast Texas.............................................................................. 12 12(11). Posterosuperiormetafemoral margin with single small tooth ( Fig. 28H); posterior metatrochanteral margin serrate with several small teeth; median lobe of male genitalia wide, not “pinched” medially ( Fig. 28F); New Mexico, west Texas.............. G. caenulenta , n. sp. — Posterosuperiormetafemoral margin without tooth, or occasionally with single, tiny, indistinct tooth; posterior metatrochanteral margin with 3 small teeth ( Fig. 27G); median lobe of male genitalia slender, “pinched” medially ( Fig. 17E); Arizona....................... G. gordoniWarner 13(1). Tubercles in median emargination of clypeal apex large, pronounced, tooth—like ( Fig. 34B); metasternal groove widened from base to apex, apex extremely wide, nearly reaching metipimeron ( Fig. 34D); Costa Rica.......................................................................... G. costaricensis, n. sp. — Tubercles in median emargination of clypeal apex small, tubercles irregular but more or less equal in size across apical margin ( Fig. 33B); metasternal groove not extremely widened, apex not approaching metepimeron ................................................................................................. 14 14(13). Metatibia with posterolateral projection ( Fig. 33H); southern Mexico.................................... 15 — Metatibia without posterolateral projection, lateral flange usually serrate with small teeth ( Fig. 35F); not known from southern Mexico.................................................................................. 16 15(14). Metasternal surface lacking pronounced carinae arranged in reticulate pattern; surface of metafemur finely rugose, appearing smooth ( Fig. 33G); Zacateca, Mexico................................ ...................................................................................................................... G. zacateca, n. sp. — Metasternal surface with pronounced carinae arranged in reticulate pattern ( Fig. 32D); surface of metafemur coarsely microreticulate; Puebla, Mexico.............................. G. donaldi, n. sp. 16(14). Posterior margin of metatrochanter with series of large tubercles, second tubercle to trochanteral margin pronounced, tooth—like ( Fig. 31F); surface of clypeus, apical portion of frons with large, dense tubercles ( Fig. 31B); Guatemala.............................................. G. dentata, n. sp. — Posterior margin of metatrochanter smooth, without tubercles ( Fig. 35F); surface of clypeus and anteromedian portion of frons densely, finely rugose, with small, sparse tubercles ( Fig. 35B); Mexico, Coahuila........................................................................................... G. limbata , n. sp.
Metasternal
33
32
1
[192,1299,739,763]
Posterosuperior
33
32
1
Posterosuperior
33
32
1