Systematic revision of American Glaresidae (Coleoptera: Scarabaeoidea) Gordon, Robert D. Hanley, Guy A. Insecta Mundi 2014 2014-01-10 2014 333 1 91 4NT7 Erichson 1848 [422,535,515,539] Insecta Glaresidae Glaresis Animalia Coleoptera 32 31 Arthropoda genus    1. Mesotibial apex strongly projecting laterally, projection apically acute ( Fig. 34C); metatibia abruptly widened before apex; Mexico, Central America; subgroup C ................................... 12  — Mesotibial apex weakly projecting laterally, projection rounded ( Fig. 29D) (except  G. confusa ,  G. falli ,and G. texanawith apex strongly projecting, apically acute); metatibia gradually widened from base to apex ( Fig. 19G); Mexico, Canada, United States; subgroups A, B ..................... 2   2(1). Metasternal groove shallow, at least inner border of groove tapered ( Fig. 19D); subgroup A .. 3  — Metasternal groove deep, distinct, sides abrupt, ridged ( Fig. 27D); subgroup B ....................... 7   3(2). Outer margin of metatibia with series of small, regular teeth from base to apex, without large posteromedian lateral projection ( Fig. 21F); Texas...................................  G.  texanaGordon  — Outer margin of metatibia with distinct posteromedian lateral projecton ( Fig. 20F); not known from Texas.................................................................................................................................. 4   4(3). Clypeal apex with large, widely spaced tubercles medially ( Fig. 23B); metatibial posteromedian lateral projection large, apically bifid ( Fig. 23F) ...............................................  G. falli , n. sp.  — Clypeal apex with small, evenly spaced tubercles throughout ( Fig. 22B); metatibial posteromedian lateral projection small, tooth—like ( Fig. 22F) ........................................................................ 5   5(4). Tubercles on head surface small, sparse ( Fig. 22B); apex of mesotibia strongly projecting, apically acute; Alberta, Colorado, Utah...................................................................   G. confusaBrown  — Tubercles on head large, dense ( Fig. 19B); apex of mesotibia not strongly projecting, apically rounded ....................................................................................................................................... 6   6(5). Posterosuperior metatrochanteral surface with 2 teeth ( Fig. 19C); median lobe of male genitalia wide, apex broadly rounded ( Fig. 19E) .....................................................   G. california , n. sp.   —Posteriosuperior metatrochanteral surface with single tooth ( Fig. 20C); median lobe of male genitalia slender, apically acute ( Fig. 20E) .............................................   G. howdeniGordon   7(2). Posterosuperior metatrochanteral surface with 2 teeth ( Fig. 24G) ............................................ 8  — Posterosuperior metatrochanteral surface with single tooth ( Fig. 27G) .................................. 10   8(7). Metasternal sculpture with carinae straight, not irregular, pronotal carinae distinctly spaced, not dense; male genitalia with basal lobe slender, narrowed in basal 1/3, apex rounded ( Fig. 26F); southern California............................................................................  G. bautista,  n. sp.  — Metasternal sculpture with carinae irregular, dense; pronotal carinae closely spaced; male genitalia not as described for  G. bautista; not known from southern California................................... 9   9(8). Metatasternal sculpture irregular, forming reticulate pattern extended from apex to metacoxal cavity of metasternum ( Fig. 24D); apical trochanteral margin with small teeth, teeth often not visible; male genitalia with basal lobe broad, apically rounded ( Fig. 24E); Arizona, California, Mexico............................................................................................................   G. mendicaHorn  —  Metasternalsculpture irregular, not forming reticulate pattern, not extended to metacoxal cavity ( Fig. 25D); apical trochanteral margin with several distinct teeth; male genitalia with basal lobe narrow, apically acute ( Fig. 25E); California, Idaho, Nevada, Wyoming........................... .............................................................................................................   G. paramendica, n. sp.   10(7). Posteromedian lateral projection of metatibia large, pronounced ( Fig. 29F); surface sculpture of frons and vertex composed of dense raised, irregular ridges coalesced into a reticulate pattern ( Fig. 29B); Baja California Sur.................................................................   G. australis. n. sp.  — Posteromedian lateral projection of metatibia small, not pronounced ( Fig.20F); surface sculpture of frons and vertex reduced, if ridges present then not forming a reticulate pattern; United States......................................................................................................................................... 11   11(10). Metasternum tumid, protuberant, with heavy, somewhat reticulate surface sculpture laterally ( Fig. 30D); southeast Texas...........................................................................   G. tumida, n. sp.  — Metasternum flat or medially concave, surface sculpture comparatively light, not forming reticulate pattern; not known from southeast Texas.............................................................................. 12    12(11). Posterosuperiormetafemoral margin with single small tooth ( Fig. 28H); posterior metatrochanteral margin serrate with several small teeth; median lobe of male genitalia wide, not “pinched” medially ( Fig. 28F); New Mexico, west Texas..............   G. caenulenta , n. sp.  —  Posterosuperiormetafemoral margin without tooth, or occasionally with single, tiny, indistinct tooth; posterior metatrochanteral margin with 3 small teeth ( Fig. 27G); median lobe of male genitalia slender, “pinched” medially ( Fig. 17E); Arizona.......................   G. gordoniWarner   13(1). Tubercles in median emargination of clypeal apex large, pronounced, tooth—like ( Fig. 34B); metasternal groove widened from base to apex, apex extremely wide, nearly reaching metipimeron ( Fig. 34D); Costa Rica..........................................................................   G. costaricensis, n. sp.  — Tubercles in median emargination of clypeal apex small, tubercles irregular but more or less equal in size across apical margin ( Fig. 33B); metasternal groove not extremely widened, apex not approaching metepimeron ................................................................................................. 14   14(13). Metatibia with posterolateral projection ( Fig. 33H); southern Mexico.................................... 15  — Metatibia without posterolateral projection, lateral flange usually serrate with small teeth ( Fig. 35F); not known from southern Mexico.................................................................................. 16   15(14). Metasternal surface lacking pronounced carinae arranged in reticulate pattern; surface of metafemur finely rugose, appearing smooth ( Fig. 33G); Zacateca, Mexico................................ ......................................................................................................................   G. zacateca, n. sp.  — Metasternal surface with pronounced carinae arranged in reticulate pattern ( Fig. 32D); surface of metafemur coarsely microreticulate; Puebla, Mexico..............................   G. donaldi, n. sp.   16(14). Posterior margin of metatrochanter with series of large tubercles, second tubercle to trochanteral margin pronounced, tooth—like ( Fig. 31F); surface of clypeus, apical portion of frons with large, dense tubercles ( Fig. 31B); Guatemala..............................................   G. dentata, n. sp.  — Posterior margin of metatrochanter smooth, without tubercles ( Fig. 35F); surface of clypeus and anteromedian portion of frons densely, finely rugose, with small, sparse tubercles ( Fig. 35B); Mexico, Coahuila...........................................................................................  G. limbata , n. sp. Metasternal 33 32 1 [192,1299,739,763] Posterosuperior 33 32 1 Posterosuperior 33 32 1