Bates, 1892 : 342
Csiki, 1932 : 1217
Trichotichnus birmanicus Bates, 1892
Kirschenhofer, 1992 : 39
Brachypterous ground beetles of the Trichotichnus subgenus Bottchrus Jedlička (Coleoptera, Carabidae) from the Himalaya, with description of fifteen new species
Schmidt, Joachim
Zootaxa
2017
2017-09-22
4323
3
301
358
8LPDD
Bates, 1892
Bates
1892
[151,759,542,568]
Insecta
Carabidae
Trichotichnus
Animalia
Coleoptera
49
350
Arthropoda
species
birmanicus
Bottchrus
Trichotichnus Birmanicus Bates, 1892: 342. Typelocality: “Karin Cheba, alt. 900–1100 m” ( Karen Hills, approximate coordinates 19°13'N 96°35'E), Myanmar. Trichotichnus( Trichotichnus) Batesi Csiki, 1932: 1217, unnecessary replacement name for Trichotichnus birmanicus Bates, 1892. Trichotichnus( Bellogenus) probsti Kirschenhofer, 1992: 39, syn. n. Typelocality: Tatopani, 1100–1400 m, Kali GandakiKhola, Myagdi Distr., Nepal.
Type material examined. Syntype of T. birmanicus: ♂, “Carin Cheba, 900–1100 m, L. Fea, V–XII.88”, “ Trichotichnus BirmanicusBates” (Bates’ handwriting) (MNHN). Holotypeof T. probsti: ♀, “W-Nepal, Dhawalagiri, Myagdi Distr., Kali– Gandaki– Khola, Tatopani, 1100– 1400 m, 14–17.VI.1986, leg. J. Probst”, “ Holotypus, Trichotichnus( Bellogenus) probsti n. sp., det. Kirschenhofer, 91” ( NHMW). Addition material examined. India. 1 ♀, Sikkim, Dzongri, 3000 m, 15.VIII.1977, B. Bhaktaleg. ( NHMB); 1 ♂, 3 ♀, NE India, Meghalaya, 1 kmE of Tura, 550 m, 25°30'N 90°14'E, 10–15.VI.2002, M. Tryzna, P. Bendaleg. (cFCCH, cWR, ZIN). Nepal. 1 ♂, Karnali Prov., Mugu Distr., Mugu Karnali, Mangriup to Taka, 2100–2300 m, 29°34.2'N 82°23.5'E, 28.VI.1999, M. Hartmannleg. ( NME); 1 ♀, BheriProv., Dailekh Distr., Dailekh, S Katia Khola, 800 m, 31.V.1995, A. Weigelleg. ( NME); 1 ♂, Dhaulagiri, Phedito Phalegaon, 28°28'N 83°21'E, 1100– 1800 m, 9.V.2012, J. Schmidtleg. (cJS); 1 ♂, 3 ♀, Annapurna Mts., Ullerisüdl. Ghorepani, ca 2000 m, 16.VI.1993, J. Schmidtleg. (cJS); 1 ♂, Annapurna Mts, Dana bis Tatopani, 1200–1400 m, 13.VI.1993, J. Schmidtleg. (cJS); 1 ♂, Kali GandakiVall., betw. Tatopani and Beni, ca 1000 m, 29.X.1992, J. Schmidtleg. (cJS); 1 ♂, Annapurna, South Himal, 1600 m, GharaW Ghorepani, 29.V.2002, J. Schmidtleg. (cJS); 1 ♂, 3 ♀, Nepal420, Kathmandu Distr., Kathmandu–Baneshwar, 1350 m, 18.IV.1885, Martens& W. Schawallerleg. ( SMNS); 2 ♀, Nepal583, Kathmandu, Baneshwar, 1300 m, 21–25.VI.1998, W. Schawallerleg. ( SMNS); 1 ♀, Nepal477, Kathmandu Distr., Kathmandu–Baneshwar, 1350 m, 2–3.VI.1995, Martens& W. Schawallerleg. ( SMNS); 1 ♀, BagmatiDistr., Kathmandu-Thamel, Wasserreservoir, HF 27°43'N 85°23'E, 1400 m, Grillleg. (cJS); 1 ♂, 1 ♀, Kathmandu, NE Gorkhana Park, Bagmati, 27°43'22"N 85°22'59"E, 1400 m, 26.VI.1997, A. Weigelleg. ( NME); 1 ♂, Kathmandu, nördlicher Stadtrand, ca 1200 m, 17.IX.1992, J. Weipertleg. ( NME); 1 ♂, Kathmandu Vall., 1300 m, 18.VI.1995, Ahrens& Pommeranzleg. (cJS); 1 ♀, Kathmandu Vall., Pharping, 1600 m, 12.VIII.2004, Anand Chaudari leg. (cJS); 1 ♂, same data as preceding, but 2.X.2004(cJS); 2 ♀, same data, but 7.VIII.2004(cJS); 1 ♀, Kathmandu Vall., Bhaktapur, 2004, Anand Chaudari leg. (cJS); 1 ♂, Kathmandu, Thamel, Hotel Norbhu Linka, LF 1300 m, 5.VI.1995, A. Weigelleg. ( NME); 1 ♀, Kathmandu, Bagmati River, 18.X.1987, Winkelmann– Klöckleg. [det. as T.( Bellogenus) probstiKirsch.by E. Kirschenhoferin 1992] (cWR); 1 ♀, Boudha, Mahankal, 20.X.2004(cJS); 1 ♀, S Khumbu-Himal, env. Jiri, Shivalaya, Kimti Khola, 1707 m, 1.V.1993, Kleebergleg. (cJS); 1 ♂, E Nepal, Dhankuta, Arun Valley, SE Makalu, Navagaon – Num, 1900– 700 m, 16.VI.1980, C. Holzschuhleg. [det. as T. glabellusAndr.by E. Kirschenhofer] (cWR), 1 ♂, E Nepal, Arun Valley, Ruhruma –Waleng Iswakhola, 1200 m, 14.VI.1992, J. & J. Probstleg. [det. as T. probstim. by E. Kirschenhoferin 1992] ( NHMW). Bhutan. 1 ♀, Wangdi Phodrang Pr., 40 kmSSE Wangdi Phodrang, Puna Tsang Vall., ca 790 m, 27°12'38"N 90°03'05"E, 25.XI.2005, M. Jächleg. ( NHMW). Burma. 1 ♀, Tenasserim, Malvedaung, 30 kmS of Ye, 300 m, 15–25.XI.1934, R. Malaiseleg. ( SMNHS); 1 ♂, env. Naymyo[= Pyin Oo Lwin], 700–900 m, 12–13.II.1996, S. Kurbatovleg. ( ZIN); 3 ♀, ShanProv., Namhsan, 1600 m, litter, 18.II.1996, S. Kurbatovleg. ( ZIN). China. Yunnan: 1 ♂, Nujiang Lisu Aut. Pref., creek valley, 3 kmSE Gongshan, 27°43'02"N 98°41'27"E, 1450–1500 m, litter/moss, sifted, [20], 5.VI.2007, D.W. Wraseleg. (cWR); 2 ♂, same data as preceding, but A. Pützleg. (cPTZ; cWR); 1 ♀, Taveishan Mts.near Pingpien, 1500 m, 21.VI.1956, Hwang Keleg. ( ZIN); 2 ♂, Lungling, 1600 m, 20.V.1955, O. Kryzhanovskijleg. ( ZIN); 1 ♂, 2 ♀, Baoshan Pref., Gaoligong Shan, 65 kmNNE Tengchong, 1750 m, 25°35'20"N 98°40'21"E, secondary mixed forest, overgrown stone debris, litter / moss sifted, 27.VIII.2009, M. Schülkeleg. [27B] (cWR); 5 ♂, 2 ♀, same data as preceding, but D.W. Wraseleg. [10] (cWR, ZIN); 1 m, 2 ♀, same data as preceding, but pasture, under decaying plant material [10C] (cWR); 1 ♀, Longling, Zhen-an, Bangbie, steamside, 24.81327°N 98.83271°E, 1550 m, 24.V.2005, H.B. Liang, D. Kavanaugh leg. (IOZ); 1 ♀, Baoshan, Longyang, Bawan, Nankang station, 24.82260°N 98.78201°E, 2060 m, 23.V.2005, H.B. Liangleg. (IOZ); 1 ♀, Gongshan Co., Gongshan Township, Dandang Preserve, 1675–1820 m, 27.74323°N 98.66168°Eto 27.74183°N 98.65908°E, 29.VI.2000, Stop#00–16, D. Kavanaughleg. (IOZ); 1 ♀, Gongshan Co., Gongshan, 1500 m, 27.74064°N 98.66557°E, 28.VI.2000, Stop#00– 14, D. Kavanaugh, H.B. Liangleg. (IOZ); 2 ♀, Gongshan Co., Maxidang, along road, 27°52'41"N 98°39'15"E, 1550 m, 19.IV.2002, H.B. Liang, W. Ba, G. Yang, X.Q. Lileg. (IOZ); 1 ♀, Gongshan, Bingzhongluo Vill., 28°01'14"N 98°37'12"E, 1700 m, 13.IV.2002, H.B. Liang, W. Ba leg. (IOZ); 1 ♂, Tengchong, Tengyue, Laifengshan, 25.01734°N 98.47719°E, 1920 m, 1.VI.2006, D. Kavanaugh leg. (IOZ); 1 ♀, Lushui Co., Pianma, Gangfang, Xuetang, 26.12218°N 98.57546°E, 1625 m, 16.V.2005, D. Kavanaugh, D.Z. Dongleg. (IOZ); 1 ♀, Fugong, Pihe, at junction of Nujiang River, 26.53177°N 98.89753°E, 1060 m, 20.IV.2004, D. Kavanaugh, C. Griswoldleg. (IOZ); 1 ♂, 1 ♀, Fugong Co., Shangpa, Guquan Village, roadside, 26.88897°N 98.86558°E, 1230 m, 21.IV.2004, H.B. Liang, X.Y. Lileg. (IOZ); 1 ♀, S Gejiu, 1860 m, 23°17'14"N 103°08'41"E(grassy slope with shrubs near road, litter / soil / moss sifted), 21.VIII.2014, M. Schülkeleg. [17] (cWR); 2 ♀, S Yunnan( Xishuangbanna), 37 NW Jinghong, vic. Guo Men Shan, 22°14'48"N 100°36'22"E, 1000 m, 27.IV.2009, LF, leg. A. Weigel( NME); 1 ♂, same data as preseding, but 22°17.48'N 100°38.85'E, 1080 m, 26.V.2008, LF, A. Weigelleg. ( NME). Sichuan: 1 ♀, S Sichuan, N Lunan Shan, NE of Dechang, W slope of Mt'4282', 1600–2400 m, 27.IV.2001, I. Belousov& A. Korolevleg. ( ZIN). Hainan: 1 ♀, Changjiang County, Bawangling, Dong-er, 19.09901°N 109.17540°E, 1015 m, 8.V.2007, N, H.- B. Liangleg. (IOZ); 1 ♂, 1 ♀, same data as preceding, but 9.V.2007(IOZ); 1 ♂, Qiongzhong, Limushan, Conserve station, light trap, 19.17561°N 109.74246°E, 634 m, 29.III.2007, H.L. Shileg. (IOZ); 1 ♀, Baisha, Nakai Town, on vegetation or ground, 18.9741°N 109.2956°E, 790 m, 13.IV.2010, Lin Meiyngleg. (IOZ). Vietnam. 2 ♂, 4 ♀, Lao Cai Prov., 250 kmfrom Hanoi, Sa Paenv., Hoang Lien Son Nat. Park, 1250 m, 27.VI.–2.VII.1998, A. Napolovleg. ( MPU, ZIN); 1 ♂, same data as preceding, but 2– 15.VI.1998( ZIN); 1 ♀, Lao CaiProv., 6 kmW of SA PA, N slope of Phansipan Mt. Area, 2000–2100 m, near Tramdon (base of Hoang Lien Nat. Park), 22°21'N 103°46'E, V.2005, A.V. Abramovleg. ( Exp. of Russia-Vietnam Tropical Centre) ( ZIN); 11 ♀, Lao CaiProv., 6 kmW of Sa Pa, northern slope of Phansipan Mt. area, May 2008, A.V. Abramovleg. ( ZIN); 1 ♂, Lao CaiProv., Hoang Lien Son Mt. Range, env. Fansipan Mt., Tramton, 1950–2100 m, 15–20.VII.2007, D. Fedorenkoleg. ( ZIN); 1 ♀, Lao CaiProv., Sa Pa env., 22°19.52'N 103°50.35'E, ca 1650 m, 23–27.V.1999, Ahrensleg. (cJS); 1 ♂, Vinh Phu Prov., Tam Dao, 21°27.18'N 105°38.58'E, 1050–1200 m, 2– 6.VI.1999, Ahrensleg. (cJS); 1 ♂, Kon TumProv., Central Highlands, 2–3 kmW of Ngoc Linh Mt., 15°05' N 107°57' E, 1700–1900 m, 25.III.–14.IV.2004, A.V. Abramovleg. ( ZIN); 1 ♀, same data as preceding, but 1700– 2000 m, 20.III.–8.IV.2006, A.V. Abramovleg. ( Exp. of Russia-Vietnam Tropical Centre) ( ZIN); 2 ♂, 2 ♀, N Vietnam, Vinh PhucProv., vic. Tam Dao Town, Tamdao NP, 21°27'N 105°38'E, 700–1000 m, 2–5.V.2013, A. Skaleleg. (cSKAL; cWR); 1 ♂, 1 ♀, same data as preceding, but A. Weigelleg. ( NME); 1 ♂, N Vietnam, Cao BangProv., vic. VinDen, Nui Pia Oac Nat. Res., 22°33'53"N 105°52'53"E, 900–1300 m, 6–10.V.2013, A. Skaleleg. (cWR); 1 ♀, N Vietnam, Cao Bang/ vic. Tinh Tuc, Son Do / Nui Pia Oac NaturalR./ 22°33'53"N 105°52'53"E/ 09–15.V.2014, by light/ 900–1300 m, A. Weigelleg. ( NME); 2 ♂, 1 ♀, S Vietnam, Lam DongProv., Lac Duong Distr., 5 kmNE of Long Lanh Vill., 1400 m, Bi Dup – Nui Ba Nature Reserve, 12°10'44"N 108°40'44"E, IV.2008, A.V. Abramovleg. ( ZIN); 1 ♀, same data as preceding, but V.2009, A.V. Abramovleg. ( ZIN); 1 ♂, 1 ♀, S Vietnam, Lam DongProv., Bi Doup-Nui Ba Reserve, env. Long Lanh, at light, 12°10'44"N 108°40'44"E, 1400–1600 m, 29– 3.III.2008, D. Fedorenkoleg. ( ZIN). Laos. 2 ♀, HuaphanhProv., Ban Saluei env., 1500 m, 30 kmS of Ham Neua, 103°59'33"E 20°13'39"N, Phou PhanMt., 16–17.V.2004, Petr Kreslleg. (cBUL, cWR); 2 ♂, 4 ♀, NE Laos, Pr. Hua PhanBan Saleui, Phou Pan ( Mt.). ca 20°12'N 104°01'E, 1300–1900 m, 11.IV.–15.V.2012, C. Holzschuhleg. ( NME, ZIN); 1 ♂, same data as preceding, but 1–31.V.2011, local collector leg. ( NME). Thailand. 1 ♂, Soppong, 1–9.V.2000, P. Moravecleg. (cWR); 1 ♀, N Thailand, 100 kmNE of Nan, Doi Phu Kha N.P., F. Pavelleg. (cFCCH); 1 ♀, Chiang MaiProv., San Pakia Vill., 19.19°N 98.50°E, 1400 m, 1–15.V.1998, V. Kubáňleg. (cKM); 1 ♂, 1 ♀, same data as preceding, but M. Bednarikleg. (cKM); 2 ♂, same data, but 1–19.V.1998, I. Martinoleg. (cKM, cWR); 1 ♀, NW Thailand, Mae Hong Son, Ban Huai Po, 1600 m, 9–16.V.1991, P. Pacholatkoleg. (cBAS). Malaysia. 1 ♂, 3 ♀, MalaysiaW., Perak, 40 kmSE of Ipoh, 900 m, Banjaran Titi Vangsa Binglet, 29.III.– 29.IV.2004, P. Čechovskyleg. (cBAS, cWR, ZIN); 1 ♂, MalaysiaW, Cammeron Highlands, Ringletenv., 15.IV.2000, M. Snizekleg. (cWR); 3 ♀, KelantanState, 60 kmNE Tanah Rata, Tanah Kerajaan, 1000 m, at light, 12–30.IV.2007, P. Čechovskyleg. (cWR); 1 ♀, MalaysiaW, 25 kmNE of Ipoh, 2100 m, Banjaran Titi Wangsa mts., Korbu mt., 4–13.III.1998, P. Čechovskyleg. (cDOS); 2 ♀, Malaysia, Pahang, Fraser'shill, 17–28.VIII.2010, I. Kamskovleg. ( ISEN, ZIN); 1 ♀, Selangor Prov., Kuala SelangorNature Park, 15 m, 18–19.V.2009, A. Zamesov, E. Zamesovaleg. ( ZIN); 2 ♀, PahangProv., Fraser's Hill., 0 3°42'54"N 101°43'57"E, 1500 m, 17.V.2009, A. Zamesov, E. Zamesovaleg. ( ZIN).
Description(the most distinctive characters only). Habitus as in Fig. 123. Length 5.9–7.4 mm. Body black piceous, slightly iridescent on dorsum; legs yellowish brown, femora not infuscate. Head: Eyes large, markedly convex, wide oval in lateral view. Tempora moderately short, convex, somewhat abruptly sloped to neck. Genae wide, with short setae. Mentumand submentum fused laterally and separated by suture medially. Ligular sclerite narrow, narrowed to apex. Left mandible slightly truncate at tip in front view. Microsculpture strongly obliterate, meshes hardly visible. Antennae slightly surpassing pronotal basal edge, with antennomeres 4–8 elongate. Pronotum: Moderately wide, with sides slightly sinuate in basal third, in some specimens almost straight. Apical margin very shallowly emarginate, apical angles not prominent, widely rounded. Basal margin straight in middle portion, scarcely oblique laterally. Basal angles very slightly obtuse, sharp at tip, without denticle. Lateral gutter slightly widened in apical half. Pronotal disc moderately convex, markedly depressed and flattened laterobasally. Basal foveae usually distinct, shallow, elongate, in most specimens touching basal margin. Punctation in most specimens restricted to the latero-basal areas, variable from very sparse to moderately dense. Microsculpture obliterate on disc, more distinct along margins, consisting of very fine transverse meshes. Elytra: In lateral and caudal view slightly convex, in dorsal view elongate oval. Elytral preapical sinuation comparatively deep. Humerus with a more or less distinct denticle visible from behind. Striae markedly impressed along entire length. Intervals slightly convex, narrowed apically. Parascutellar striole comparatively long, with large pore at base. Interval 3 with one pore behind middle (in the holotypeof T. probsti, two discal pores on left elytron). Microsculpture obliterate, consisting of very fine, indistinct thin lines on disc. Hindwings: Fully developed or reduced to scales. Ventral surface: Metepisternum ( Fig. 146) narrow, markedly longer than wide. Abdominal sternites covered by very short and fine setae medially. Last visible (VII) abdominal sternite with two pairs of marginal setae, in male subtruncate, in female angularly rounded. Legs: Protibia sulcate in basal third ventrally. Profemur with two to five setae on anterio-ventral margin. Tarsomere 5 with two pairs of ventro-lateral setae. Female genitalia: Laterotergite (hemisternite) with two apical setae. Basal palpomere with one apical seta. Apical stylomere moderately curved, with one peg-like seta in both ventral and dorsal margins. Male genitalia: Median lobe of aedeagus ( Fig. 147–150) with apical orifice shifted to the left, arcuate in lateral view, with ventral margin slightly convex in apical portion. Terminal lamella short, markedly narrowed apicad in dorsal view ( Fig. 149). Apical capitulum in lateral view ( Fig. 147) oblique, with a tiny, poorly developed dorsal flange, in caudal view ( Fig. 150) somewhat triangular, narrowly rounded at tip. Internal sac without spines in internal sac. Comparison. Among the Himalayan members of Bottchruscharacterized by a comparatively small body length, T.( B.) birmanicusis recognizable by the following combination of distinctive characters: body less convex, left mandible slightly truncate at tip, eyes larger and more convex, tempora shorter, pronotum markedly depressed and flattened latero-basally, metepisterna ( Fig. 146) noticeably longer and narrower, hindwings fully developed or reduced, elytra with deeper preapical sinuation and with narrower and more convex intervals apically, and median lobe of aedeagus ( Figs 147–150) as described above. In some characters (left mandible slightly truncate at tip, pronotum depressed latero-basally, rather long metepisterna, and protibia sulcate on dorsal side), T.( B.) birmanicusreminds T.( B.) formosusfrom Sikkim, but the latter species is much larger (about 9.5 mm) and has median lobe of aedeagus with longer terminal lamella and much larger apical capitulum. In external morphology and male genitalia, T.( B.) birmanicusis very similar to T.( B.) horni Schauberger, 1935from Borneo, but the latter species differs in slightly larger body (6.8–8.0 mm), pronotum almost impunctate basally, and terminal lamella of the median lobe longer, with apical capitulum more transverse and dorsally slightly more projected.
Remarks. The description of Trichotichnus( B.) probstiis based on a single female collected near Tatopani in the Kali GandakiValley of Central Nepal. It was considered most related to T.( B.) holzschuhibut distinguishing from this taxon by slightly larger, less convex and more elongate body, and by longer elytra with deeper striae and more convex intervals. Re-examination of the holotyperevealed that this taxon is conspecific with T.( B.) birmanicus.
Distribution and geographical variation. Trichotichnus( B.) birmanicusis widely distributed across the Oriental region. Andrewes (1930b)reported this species from North India(Mussoorie in Uttarakhand, and Lebong in Darjeeling, West Bengal), Myanmar(Karen Hills), and Laos. Schauberger (1935)described its distribution as “Himalaya, nördl. Vorderindien, Hinterindien”. Ito (1991)reported T.( B.) birmanicusfrom Thailandand also illustrated its male and female genitalia. Based on the examined material, this species also occurs in Nepal, Sikkim, Bhutan, China( Yunnan, Sichuan, Hainan), Vietnam, Laos, and West Malaysia. In the Himalayan region, the species occurs at altitudes of 550–3000 m. The record of T.( B.) birmanicusfrom Hong Kong( Hua 2002, Aston 2016) most likely refers to another species, Harpaliscus birmanicus Bates, 1892, because it is based on Schauberger (1934: 101)who considered Harpaliscusas a subgenus of Trichotichnusand cited the latter species as Trichotichnus( Harpaliscus) birmanicus. It should be noted that most populations from Nepal, Myanmarand Yunnanare dimorphic, with hindwings either fully developed or reduced to small scales, while the populations from more southern areas seem to be fully winged. Trichotichnus( B.) birmanicusis probably a geographical vicariant of T.( B.) horni Schauberger, 1935described from Borneo, which is also a fully winged species. The status of the specimens examined from West Malaysia(unfortunately, most of them are females) needs further study, because they are, on average, slightly larger ( 6.7–7.5 mm) and robuster than specimens of T.( B.) birmanicusfrom other localities and may be treated in this respect as transition to T.( B.) horni. However, their pronotum is very finely and sparsely punctate in basal foveae and latero-basally, as in other specimens of T.( B.) birmanicus, and thus different of T.( B.) horni.
1655986727
Myanmar
1000
19.216667
Karen Hills
1272
96.583336
49
350
1
holotype
1655986732
Nepal
1250
Myagdi Distr.
49
350
1
Gandaki
holotype
1655986743
[199,1011,943,968]
Kali
Myagdi Distr.
49
350
1
1
Dhawalagiri
holotype
1655986738
1986-06-14
1986-06-17
1986-06-14
NHMW
J. Probst
Kirschenhofer
1250
Tatopani
Khola
49
350
1
Gandaki
holotype
1655986735
[631,1432,1052,1077]
1977-08-15
NHMB
B. Bhakta
3000
Dzongri
49
350
1
1
Sikkim
1655986745
2002-06-10
2002-06-15
2002-06-10
ZIN
M. Tryzna & P. Benda
India
550
25.5
Tura
1246
90.23333
49
350
3
3
Meghalaya
1655986733
1999-06-28
NME
M. Hartmann
Nepal
Karnali Prov.
2200
29.57
Mangri
122
82.39167
Mugu Distr.
49
350
1
1
Karnali
1655986742
1995-05-31
NME
A. Weigel
Nepal
Dailekh Distr.
800
Katia Khola
Dailekh
49
350
1
1
Bheri
1655986726
2012-05-09
J. Schmidt
Nepal
1450
28.466667
Phalegaon
1232
83.35
Phedi
49
350
1
1
Dhaulagiri
1655986734
1993-06-16
J. Schmidt
Nepal
Annapurna Mts.
2000
Ghorepani
Ulleri
49
350
3
3
Dhaulagiri
1655986724
[398,1410,1268,1293]
1993-06-13
J. Schmidt
Nepal
1300
Tatopani
49
350
1
1
Dhaulagiri
1655986740
1992-10-29
J. Schmidt
Nepal
1000
Gandaki
49
350
1
1
Gandaki
1655986752
2002-05-29
J. Schmidt
Nepal
Annapurna
1600
Ghara
South Himal
49
350
1
1
Gandaki
1655986763
1885-04-18
SMNS
Martens & W. Schawaller
Nepal
1350
Kathmandu - Baneshwar
Kathmandu Distr.
49
350
3
3
1655986770
1998-06-21
1998-06-25
1998-06-21
SMNS
W. Schawaller
Nepal
1300
Nepal
49
350
2
2
1655986750
1995-06-02
1995-06-03
1995-06-02
SMNS
Martens & W. Schawaller
Nepal
1350
Kathmandu - Baneshwar
Kathmandu Distr.
49
350
1
1
1655986749
Grill
Nepal
1400
27.716667
Wasserreservoir
1235
85.38333
Kathmandu-Thamel
49
350
1
1
Bagmati
1655986762
1997-06-26
NME
A. Weigel
Nepal
1400
27.722778
Gorkhana Park
20
85.38306
Kathmandu
49
350
1
1
Bagmati
1655986751
1992-09-17
NME
J. Weipert
Nepal
1200
Stadtrand
Kathmandu
49
350
1
1
Bagmati
1655986768
1995-06-18
Ahrens & Pommeranz
Nepal
1300
Kathmandu Vall.
49
350
1
1
Bagmati
1655986757
[225,784,1628,1653]
2004-10-02
Ahrens & Pommeranz
Nepal
1300
Kathmandu Vall.
49
350
1
1
Bagmati
1655986755
[797,1232,1628,1653]
2004-08-07
Ahrens & Pommeranz
Nepal
1300
Kathmandu Vall.
49
350
2
2
Bagmati
1655986765
1995-06-05
NME
A. Weigel
Nepal
Kathmandu
1300
Hotel Norbhu Linka
Thamel
49
350
1
1
Bagmati
1655986753
1987-10-18
Winkelmann & Klock & E. Kirschenhofer
Nepal
Kathmandu
49
350
1
1
Bagmati River
1655986767
1993-05-01
Kleeberg
Nepal
1707
Khumbu-Himal
49
350
1
1
Bagmati River
1655986748
1980-06-16
C. Holzschuh & E. Kirschenhofer
Nepal
700
Navagaon - Num
Makalu
49
350
1
1
1655986761
1992-06-14
NHMW
J. & J. Probst & E. Kirschenhofer
Nepal
1200
Ruhruma - Waleng Iswakhola
Arun Valley
49
350
1
1
1655986758
2005-11-25
NHMW
M. Jach
Bhutan
790
27.210556
Puna Tsang Vall.
20
90.05139
Wangdi Phodrang
49
350
1
1
Wangdi Phodrang
1655986754
1934-11-15
1934-11-25
1934-11-15
SMNHS
R. Malaise
Myanmar
300
Ye
49
350
1
1
1655986759
[283,1307,1988,2013]
1996-02-12
1996-02-13
1996-02-12
ZIN
S. Kurbatov
Myanmar
800
Pyin Oo Lwin
Naymyo
49
350
1
1
1655986766
1996-02-18
ZIN
S. Kurbatov
Myanmar
1600
50
351
Namhsan
49
350
3
3
Shan
1655986760
2007-06-05
D. W. Wrase
China
1475
27.717222
Gongshan
20
98.690834
Nujiang Lisu Aut. Pref.
50
351
1
1
Yunnan
1655986764
[366,1031,223,248]
2007-06-05
A. Putz
China
1475
27.717222
Gongshan
20
98.690834
Nujiang Lisu Aut. Pref.
50
351
2
2
Yunnan
1655986747
1956-06-21
ZIN
Hwang Ke
China
1500
Pingpien
Taveishan Mts.
50
351
1
1
Yunnan
1655986771
[649,1374,260,285]
1955-05-20
ZIN
O. Kryzhanovskij
China
1600
Lungling
50
351
2
2
Yunnan
1655986756
2009-08-27
M. Schulke
China
1750
25.58889
Tengchong
21
98.6725
Baoshan Pref.
50
351
2
2
Shan
1655986769
2009-08-27
ZIN
D. W. Wrase
China
1
25.58889
Tengchong
21
98.6725
Baoshan Pref.
50
351
2
2
Shan
1655986774
2009-08-27
ZIN
D. W. Wrase
China
1
25.58889
Tengchong
21
98.6725
Baoshan Pref.
50
351
2
2
Shan
1655986791
2005-05-23
H. B. Liang
China
Baoshan
2060
24.8226
Bawan
1
98.78201
Longyang
50
351
1
1
Shan
1655986793
2000-06-29
Stop & D. Kavanaugh
China
Gongshan Co.
1748
27.74183
Dandang Preserve
1
98.65908
Gongshan Township
50
351
1
1
Shan
1655986772
2000-06-28
Stop & D. Kavanaugh & H. B. Liang
China
Gongshan Co.
1500
27.74064
Gongshan
1
98.66557
50
351
1
1
Shan
1655986788
2002-04-19
H. B. Liang & W. Ba & G. Yang & X. Q. Li
China
Gongshan Co.
1550
27.878056
Maxidang
20
98.65417
50
351
2
2
Shan
1655986784
2005-05-16
D. Kavanaugh & D. Z. Dong
China
Lushui Co.
1625
26.12218
Gangfang
1
98.57546
Pianma
50
351
1
1
Shan
1655986794
2004-04-20
D. Kavanaugh & C. Griswold
China
1060
26.53177
Nujiang River
1
98.89753
50
351
1
1
Nujiang River
1655986790
2004-04-21
H. B. Liang & X. Y. Li
China
Fugong Co.
1230
26.88897
Guquan Village
1
98.86558
Shangpa
50
351
1
1
Nujiang River
1655986780
2014-08-21
M. Schulke
China
1860
23.287222
Gejiu
21
103.14472
50
351
1
1
Nujiang River
1655986775
2009-04-27
NME
A. Weigel
China
Xishuangbanna
1000
22.246668
Guo Men Shan
21
100.60611
Jinghong
50
351
2
2
Yunnan
1655986787
2008-05-26
NME
A. Weigel
China
Xishuangbanna
1080
22.291334
Guo Men Shan
12
100.6475
Jinghong
50
351
1
1
Yunnan
1655986785
2001-04-27
ZIN
I. Belousov & A. Korolev
China
Lunan Shan
2000
Mt
Dechang
50
351
1
1
Sichuan
1655986779
2007-05-08
B. Liang
China
Changjiang County
1015
19.09901
Dong-er
1
109.1754
Bawangling
50
351
1
1
Hainan
1655986773
2007-05-09
B. Liang
China
Changjiang County
1015
19.09901
Dong-er
1
109.1754
Bawangling
50
351
1
1
Hainan
1655986789
2007-03-29
H. L. Shi
China
Qiongzhong
634
19.17561
Conserve station
1
109.74246
Limushan
50
351
1
1
Hainan
1655986783
2010-04-13
Lin Meiyng
China
790
18.9741
Nakai Town
7
109.2956
Baisha
50
351
1
1
Hainan
1655986795
1998-06-27
1998-07-02
1998-06-27
MPU, ZIN
A. Napolov
China
Lao Cai Prov.
1250
Hoang Lien Son Nat. Park
Sa Pa
50
351
4
4
Hanoi
1655986781
1998-06-02
1998-06-15
1998-06-02
ZIN
A. Napolov
China
Lao Cai Prov.
1250
Hoang Lien Son Nat. Park
Sa Pa
50
351
1
1
Hanoi
1655986792
2005-05
ZIN
A. V. Abramov
China
Phansipan Mt. Area
2050
22.35
Hoang Lien Nat. Park
1260
103.76667
Tram
50
351
1
1
Lao Cai
1655986777
2008-05
ZIN
A. V. Abramov
China
Phansipan Mt. area
50
351
11
11
Lao Cai
1655986776
2007-07-15
2007-07-20
2007-07-15
ZIN
D. Fedorenko
China
2025
Tramton
Fansipan Mt.
50
351
1
1
Lao Cai
1655986796
1999-05-23
1999-05-27
1999-05-23
Ahrens
China
1650
22.325333
Lao Cai
12
103.839165
50
351
1
1
Lao Cai
1655986782
1999-06-02
1999-06-06
1999-06-02
Ahrens
China
1125
21.453
Tam Dao
12
105.643
Vinh Phu Prov.
50
351
1
1
Lao Cai
1655986778
[498,740,1448,1472]
China
Kon Tum
50
351
1
1
Kon Tum
1655986786
2004-03-25
2004-04-14
2004-03-25
ZIN
A. V. Abramov
China
1800
15.083333
Ngoc Linh Mt.
1286
107.95
50
351
1
Central Highlands
1655986797
2006-03-20
2006-04-08
2006-03-20
ZIN
A. V. Abramov
China
1850
15.083333
Exp. of Russia-Vietnam Tropical Centre
1286
107.95
50
351
1
1
Centre
1655986803
2013-05-02
2013-05-05
2013-05-02
A. Skale
Vietnam
850
21.45
Tam
1264
105.63333
Tam Dao Town
50
351
2
2
Vinh Phuc
1655986826
[529,1176,1591,1616]
2013-05-02
2013-05-05
2013-05-02
NME
A. Weigel
Vietnam
850
21.45
Tam
1264
105.63333
Tam Dao Town
50
351
1
1
Vinh Phuc
1655986801
Vietnam
Cao Bang Prov.
50
351
1
1
Cao Bang
1655986809
2013-05-06
2013-05-10
2013-05-06
A. Skale
Vietnam
1100
22.564722
Vin
21
105.88139
50
351
1
Vin
1655986813
2014-05-09
2014-05-15
2014-05-09
NME
A. Weigel
Vietnam
1100
22.564722
Nui Pia Oac Natural
21
105.88139
50
351
1
1
Cao Bang
1655986818
2008-04
ZIN
A. V. Abramov
Vietnam
1400
12.178889
Bi Dup - Nui Ba Nature Reserve
21
108.67889
Long Lanh Vill.
50
351
1
1
Lam Dong
1655986802
[447,1192,1772,1797]
2009-05
ZIN
A. V. Abramov
Vietnam
1400
12.178889
Bi Dup - Nui Ba Nature Reserve
21
108.67889
Long Lanh Vill.
50
351
1
1
Lam Dong
1655986800
2008-03-03
2008-03-29
2008-03-03
ZIN
D. Fedorenko
Vietnam
1500
12.178889
Long Lanh
21
108.67889
50
351
1
1
Lam Dong
1655986799
Laos
1500
20.2275
Ham Neua
21
103.99249
50
351
2
2
Huaphanh
1655986805
[445,1173,1879,1904]
2004-05-16
2004-05-17
2004-05-16
Petr Kresl
Laos
Phou Phan
50
351
2
2
Phou Phan
1655986819
2012-04-11
2012-05-15
2012-04-11
NME, ZIN
C. Holzschuh
Laos
1600
20.2
Mt.
1268
104.01667
50
351
4
4
Pr. Hua Phan
1655986807
[308,1139,1952,1977]
2011-05-01
2011-05-31
2011-05-01
NME
C. Holzschuh
Laos
1600
20.2
Mt.
1268
104.01667
50
351
1
1
Pr. Hua Phan
1655986808
2000-05-01
2000-05-09
2000-05-01
P. Moravec
Thailand
Soppong
50
351
1
1
1655986804
[589,1436,1988,2013]
N. P., F. Pavel
Thailand
Doi Phu Kha
50
351
1
1
Nan
1655986815
[151,1411,151,176]
1998-05-01
1998-05-15
1998-05-01
V. Kuban
1400
19.19
Chiang Mai
763
98.5
51
352
1
1
Chiang Mai
1655986828
1998-05-01
1998-05-15
1998-05-01
M. Bednarik
1400
19.19
Chiang Mai
763
98.5
51
352
2
1
1
Chiang Mai
1655986814
1998-05-01
1998-05-19
1998-05-01
I. Martino
1400
19.19
Chiang Mai
763
98.5
51
352
2
2
Chiang Mai
1655986811
[314,1432,223,248]
1991-05-09
1991-05-16
1991-05-09
P. Pacholatko
Thailand
1600
Ban Huai Po
51
352
1
1
Mae Hong Son
1655986798
2004-03-29
2004-04-29
2004-03-29
ZIN
P. Cechovsky
Malaysia
900
Banjaran Titi Vangsa Binglet
Ipoh
51
352
4
3
1
Perak
1655986812
2000-04-15
M. Snizek
Malaysia
Ringlet
51
352
1
1
Highlands
1655986820
2007-04-12
2007-04-30
2007-04-12
P. Cechovsky
Malaysia
1000
Tanah Kerajaan
Tanah Rata
51
352
3
3
Kelantan
1655986810
1998-03-04
1998-03-13
1998-03-04
P. Cechovsky
Malaysia
2100
Ipoh
51
352
1
1
1655986806
2010-08-17
2010-08-28
2010-08-17
ISEN, ZIN
I. Kamskov
Malaysia
Fraser's
51
352
2
2
Pahang
1655986816
2009-05-18
2009-05-19
2009-05-18
ZIN
A. Zamesov & E. Zamesova
Malaysia
15
Selangor Prov.
51
352
1
1
Selangor
1655986822
2009-05-17
ZIN
A. Zamesov & E. Zamesova
Malaysia
1500
3.7150002
Pahang
21
101.7325
51
352
2
2
Pahang