New species and records of heterostigmatic mites (Acari: Prostigmata: Heterostigmata) phoretic on scarabaeid dung beetles (Coleoptera: Scarabaeidae) from northwestern Iran Sobhi, Mohammad Hajiqanbar, Hamidreza Mortazavi, Azim Zootaxa 2017 4276 3 427 434 Khaustov, 2008 Khaustov 2008 [238,718,1671,1697] Arachnida Pygmephoridae Pseudopygmephorellus GBIF Animalia Prostigmata 5 432 Arthropoda genus  Type species:  Pseudopygmephorellus reductusKhaustov, 2008, by original designation