Platanthera madagascariensis sensu Schltr., Beihefte zum Botanischen Centralblatt 33: 299 ( Schlechter 1916 ) Benthamia perularioides Schltr ., Repertorium Specierum Novarum Regni Vegetabilis 33: 32 ( Schlechter 1924 ) Revision of the genus Benthamia A. Rich. (Orchidaceae, Orchidioideae, Habenariinae) Hervouet, Jean-Michel Descourvières, Pascal Hermans, Johan Longou, Ludivine Adansonia 2025 3 2025-07-21 47 11 171 249 LHN7 Schltr. Schltr. [990,1279,472,500] Liliopsida Orchidaceae Benthamia Plantae Asparagales 63 232 Tracheophyta species rostrata  ( Fig. 87)    Repertorium Specierum Novarum Regni Vegetabilis33: 33 ( Schlechter 1924). —   Platanthera madagascariensissensu Schltr., Beihefte zum Botanischen Centralblatt33: 299 ( Schlechter 1916), nonKraenzl. ( Kraenzlin 1899: 609).    Benthamia perularioidesSchltr., Repertorium Specierum Novarum Regni Vegetabilis33: 32 ( Schlechter 1924). — Type:  Madagascar• Vakinankaratra, mont Tsiafajavona; 19°19’58”S, 47°16’08”E; alt.  2500 m;  III.1921;  H. Perrier13523; lectotype: P[ P00094567]!, here designated; isolectotype: P[ P00094568]!, syn. nov .    TYPE  MATERIAL. —   Madagascar• Vakinankaratra, mont Ibity au sud d’Antsirabe; alt.  2000 m;  II.1914;  H. Perrier8101; holotype: P[ P00094574]!;  isotype: P[ P00094575]!.   ETYMOLOGY. — From the Latin “ rostratus”, beaked, referring to the midlobe of the lip.   PHENOLOGY. — January to April.  DISTRIBUTION AND ECOLOGY. — Madagascar. Widespread in montane grasslands from 1400 to 2370 m. Can also grow on bare laterite soils ( Fig. 88).   CONSERVATION. — The conservation status of this species is of Least Concern (LC). It has the largest number of specimens of the genus in herbaria and is still widespread in Madagascar. It has however been assessed EN by IUCN in 2015 (https://doi.org/10.2305/IUCN. UK.2015-2.RLTS.T70102386A70124752.en).   ADDITIONAL SPECIMENS EXAMINED. —   Madagascar• sine loc.;  Baron849; P[ P00094569, as  Rolfeella glaberrima, on the same sheet as Dr. Fox 23but different]!, K[ K000415545]! •  sine loc.;  R. Baron5252; P[ P00094570]! •  Ankaratra;  R. Capuron759; P[ P00692268]! •  sine loc.;  A.M. Homolle1112; P[ P00692276]! •  Ihorombe, plaine de la Menarahaka, entre Ihosy et Ivohibe; 22°27’S, 46°15’E;  B. Descoings983; P[ P00692270]! •  Ankaratra;  G.F. Scott-Elliot2096; BM[ BM000034590]! •  Ankaratra;  G.F. Scott-Elliot1982; P[ P00334731]! •  Central Madagascar;  R. Baron2271; K! •  sine loc.;  R. Bonets.n.; P[ P00326135]! •  Analamanga, Tampoketsa d’Ankazobe;  E. Ursch1019; TAN!,  P[ P00692272]! •  Analamanga, Ambatovory; 18°54’S, 47°42’E;  III.1883;  Dr. Fox24; K! ex LDS•  Centre; 1887;  C.M. Le Myre de Vilerss.n.; P[ P00094474]! •  Analamanga, Ambohimanga; 18°45’42”S, 47°33’49”E;  12.VI.1905;  C. d’Alleizettes.n.; L[ L1487720]! •  Analamanga, environs de Nanisana près de Tananarive; 18°54’S, 47°42’E;  20.I.1906;  L. Rotereaus.n.; P[ P06796776]! •  Analamanga, Nanisana; 18°54’S, 47°42’E;  I.1906;  C. d’Alleizette977M; P[ P00334732]! •  Vakinankaratra, Manandona; alt.  1400 m;  IV.1912;  H. Perrier8024; P[ P00094576]! •  Vakinankaratra, mont Ibity; 20°04’03”S, 47°00’04”E; alt.  2000 m;  II.1921;  H. Perrier13581; P[ P00094578]! •  Analamanga, Ambohimanga; 18°45’42”S, 47°33’49”E;  27.III.1921;  R. Decary302; P[ P00334738]! •  Analamanga, Ambohimanga; 18°45’42”S, 47°33’49”E;  IV.1921;  R. Decary52; P[ P00334739]! •  Haute Matsiatra, massif d’Andringitra; alt.  2000 m;  II.1923;  H. Perrier14431; P[ P00094579]! •  Diana, Ankaizina; alt.  1500 m;  19.IV.1923;  R. Decary1918; P[ P00334737]! •  Diana, Ankaizina; alt.  1500 m;  19.IV.1923;  R. Decary1936; P[ P00334736]! •  Diana, Ankaizina; 18°54’S, 47°42’E; alt.  1700 m;  19.IV.1923;  R. Decary1997; P[ P00334735]! •  Vakinankaratra, entre Ambatolampy et Antsirabe; alt.  1600 m;  II.1927;  H. Perrier17906; P[ P00334733, P00334734]! •  Haute Matsiatra, Andringitra;  26.II.1938;  P. Boiteau3135; P[ P00692296, P00692297]! •  Haute Matsiatra, au sommet de l’Ambondrombe; 21°52’30”S, 47°15’37”E; alt.  2000 m;  11.IV.1941;  P. Boiteau4611; P[ P00692437]! •  Analamanga, Ambohitantely; 18°05’S, 47°16’E;  24.II.1945;  A.M. Homolles.n.; P[ P00692274]! the four plants on the left are  Tylostigma nigrescensSchltr.or  Tylostigma perrieriSchltr. •  Analamanga, Tampoketsa d’Ankazobe; 18°13’S, 47°17’E;  7.II.1948;  P. Boiteau6725; TAN! •  Diana, massif de Marivorahona; alt.  2000-2244 m;  18-26.III.1951;  H. Humbertet al. 25753; P[ P02115322plant on the right only, P02115321, plant on the right only]! •  Alaotra-Mangoro, Ambatosoratra, Atozaka, RN3; 17°38’30”S, 48°50’00”E;  20.III.1953;  ResNat6055; P[ P00692314]! •  Haute Matsiatra, RN 5, Canton Sendrisoa, District Ambalavao;  27.II.1954;  R. Rakoto6484; P[ P00692315]! •  Vakinankaratra, route d’Ambatolampy, Faratsiho; 19°23’S, 47°16’E; alt. 1800;  VIII.1955;  J.Bosser10880; P[ P00692227]! •  Vakinankaratra, Ambohimandroro, Ampatolampy;  XI.1955;  J. Bosser9206; TAN! •  Vakinankaratra, Nanokely, Ankaratra; 19°32’S, 47°03’E; alt.  2000 m;  II.1957;  J. Bosser10878; P[ P00692226]! •  Vakinankaratra, Nanokely(district de Faratsiho); 19°32’S, 47°03’E;  26.II.1960;  M. Peltier1915; P[ P00692290]! •  Analamanga, Soavina PK 22 route de Tamatave; 18°52’S, 47°40’E;  III.1960;  J. Bosser13532; P[ P00692228]! •  Amoron’i Mania, Fiadanana, Ilaka, Ambositra; 20°34’30”S, 46°57’30”E;  20.III.1960;  M. Peltier2149; P[ P00692292]! •  Analamanga, vallée de Soavina, PK 23 route de Tamatave;  II.1961;  J. Bosser14863; TAN! •  Vakinankaratra, Manjakatompo; 19°21’25”S, 47°18’28”E;  III.1961;  J. Bosser15024; P[ P00692229]! •  Analamanga, lake Mantasoa; 19°00’45”S, 47°50’17”E;  IX.1962;  J. Bosser16144; P[ P00692230]! •  Vakinankaratra, Nanokely, Ankaratra; 19°32’S, 47°03’E;  II.1963;  J. Bosser17573; P[ P00692241]! •  Route de Majunga;  III.1963;  J. Bosser16783; P[ P00692235]! •  Vakinankaratra, route d’Ambatolampy; 19°23’S, 47°16’E;  III.1963;  J. Bosser16797; P[ P00692236]! •  Vakinankaratra, route Ambatolampy-Faratsiho;  II.1964;  J. Bosser19264; P[ P00692246, P00692247]! •  Haute Matsiatra, sentier du pic Boby, Andringitra; 22°11’S, 46°53’E;  IV.1964;  J. Bosser19478; P[ P00692248, P00692249]! •  Haute Matsiatra, Andringitra; 22°13’S, 46°55’E;  IV.1964;  J.Bosser19492; P[ P00692251]! •  Haute Matsiatra, Andringitra;  IV.1964;  J. Bosser19506; P[ P00692252, P00692253]! •  Vakinankaratra, Col des Tapias,  45 kmavant Ambositra; 20°14’34”S, 47°06’04”E;  IV.1964;  J. Bosser19598; P[ P00692259]! •  sine loc.;  28.III.1969; anonym; HEID[ HEID250340]! •  Vakinankaratra, Mt. Ibity western slope; 20°05’44”S, 47°00’12”E; alt.  1900 m;  III.1972;  G. Cremers2037; TAN! •  Vakinankaratra, south of Antsirabe, Mt. Ibity; 20°10’S, 47°03’E; alt.  1300-1800 m;  III.1985;  L.C. Barnettet al. 491; TAN!,  MO[ MO3022638]! •  Haute Matsiatra, Andringitra Forest Reserve, along path to Pic Boby; 22°11’S, 46°55’E; alt.  1900-2050 m;  III.1989; P. Goldblattet al. 8964; P[ P00692435]!,  MO[ MO4314431]!,  TAN!,  WAG[ WAG.1130546]! •  Vakinankaratra, Ankaratra Massif, W of Ambatolampy; 19°20’S, 47°16’E; alt.  2350 m;  III.1989; P. Goldblattet al. 9012; MO[ MO4314448]!,  TAN! •  Amoron’i Mania, on hillside near Talavina, PK 204 from Antananarivo to Ambositra; 20°10’42”S, 47°05’24”E; alt.  1230- 1330 m;  V.1993;  C.H. Jongkindet al. 829; K!,  MO[ MO3022616]!,  P[ P00692179]!,  TAN! •  Vakinankaratra, Col Tapia,  36 kmS of Antsirabe; 20°10’41”S, 47°05’16”E; alt.  1400 m;  IV.1995;  D. DuPuyet al. M906; K!,  P[ P00059605]!,  MO!,  WAG!, TAN! •  Analamanga, Tampoketsa d’Ankazobe; 17°52’07”S, 47°03’39”E;  18.III.1999;  J.N. Labat2951; P[ P00160016, P00160017]!,  G!,  K!,  MO!, TAN! •  Between Antananarivo and Ambositra, Tapia forest; 2000;  J. Hermans4393; K! •  Vakinankaratra, between Antsirabe and Ambositra, on RN7, about  3 kmbefore Col Tapia; alt.  1440 m;  IV.2001;  J. Hermans5251; K! •  Vakinankaratra, between Antsirabe and Ambositra, on RN7, about at Col Tapia; 20°14’25”S, 47°06’12”E; alt.  1740 m;  V.2001;  J. Hermans5205; K! with spirit•  Amoron’i Mania, Ambositra, Ivato, road to Antoetra; 20°45’43”S, 47°19’31”E; alt.  1701 m;  25.I.2007;  A. SiederFS4188and  J. Hermans8086; WU[ WU0063773, spirit material]! •  Amoron’i Mania, Ambatofinandrahana,  25 kmW of Itremo village; 20°34’12”S, 46°34’55”E; alt.  1647 m;  21.II.2010;  L. Rajavelonaet al. RJL148; K[ K664548]!,  TAN! •  Amoron’i Mania, Antoetra, on inselberg; 20°46’27”S, 47°19’16”E;  II.2015; alt.  1676 m;  A. & C. Siederet al. 6801; WU! spirit material• N 4, 126 km north of Tana, inselberg;  II.2017,  A.& C. Siederet al. 7309; WU! flower in alcohol•  Vakinankaratra, between Antsirabe and Ambositra, on RN7, Col Tapia;  IV.2018;  J. Hermans8241; K! •  Analamanga, Andakana RN4; 18°15’58”S, 47°10’51”E;  19.III.2019; alt.  1461 m;  A.& C. Siederet al. 7792; WU! •  Analamanga, RN4, c.  12 kmafter Ankazobe; alt.  1461 m;  III.2019;  Sieder A.& C.et al., L7787; WU!.   NOTES A very widespread species, with a variable spur length. Any differences between  Benthamia rostrataand  B. perularioidesdo not hold when a large number of specimens are examined. Moreover, there are no distinct distribution zones as suggested by Perrier ( Perrier 1939: 24, 25), with  Benthamia perularioidesnorth of Ankaratra and  Benthamia rostratasouth of Ankaratra. This species was also often misidentified as  Benthamia bathieana, which added to the confusion.   Benthamia perularioidesand  B. rostratawere described in the same publication by Schlechter. The rules of nomenclature allow a choice of which name to prioritise. We have chosen  B. rostratabecause this name is more widely used. The type indicated by Schlechter for his  Platanthera madagascariensisis “Perrier de la Bathie XXI, β“. He copied this on specimen P00094574 which has handwritten XXI and B together with the number 8101. Schlechter (1924: 33)gave “ H. Perrier 8101(XXI B)” when he coined the name  Benthamia rostrata. The specimen P00094574 was thus sent to Berlin and returned to Paris, so there is no ambiguity that it is the holotype. For  Benthamia perularioidesSchlechtercited Perrier 13523 for the type, but there are in P two specimenswith this collection number. We have chosenP00094567 as lectotypebecause Perrier indicated on it that it was sent to Schlechter in April 1921. This species, which has an elongated spur, is different from the typeof the genus  Benthamiaand other species with a scrotiform or short spur. It is perhaps misplaced in  Benthamia. As a matter of fact, specimens identified as  Benthamia perularioidesin ( Ngugi et al.2020) were found to be embedded in  Cynorkis. At the beginning of the flowering time the spur is enveloped by the bracts, as is also observed in some African  Habenaria. See also: Schlechter (1930: t.12 n°45, as  Benthamia perularioides), Perrier (1939: 24, as  Benthamia perularioides), Perrier (1939: 25), Cribb & Hermans (2009: 42, as  Benthamia perularioides), Cribb & Hermans (2009: 43), Bosser & Lecoufle (2011: 180), Hervouet (2018: 172).   DESCRIPTION Erect slender terrestrial herb, 22-70 cmtall. Tubers 4-11, clustered, fusiform, slender and white, 5-8 mmin diameter, 20-60 mmlong, roots up to 8, wiry, white, 1-2 mmin diameter. Stem 2-4 mmin diameter, with the base covered by sheaths 25-30 mmlong, with above slightly fleshy leaves and higher up 7-10 lanceolate sheaths, gradually turning into floral bracts. Leaves 2-3, more or less narrowly oblong, 7-20 × 1.5- 4 cm, shortly acuminate or mucronate. Rachis rigid, erect, 16-25 × 1.2 cm, densely many-flowered, somewhat unilateral with 30-50 flowers. Floral bracts lanceolate, very acuminate, erectly spreading, 5-15 × 1.5-3 mm, embedding the ovary, the lower ones longer than the ovary. Ovary twisted, 5-10 mmlong, diameter 0.6-1.6 mm, the basal part fused to the base of the floral bract. Flowers resupinate, c. 7.5 mmand 4 mmdeep, olive green turning yellowish; sepals narrowly oblong, obtuse, slightly navicular, the dorsal obscurely 3-veined, erect, 1.8-3.2 × 0.7-1.6 mm, the laterals single-veined, spreading, 2.3-4 × 0.7-1.9 mm; petals narrowly oval-obtuse, obscurely 2-veined, a little expanded at the anterior margin towards the base, 1.7-3.2 × 0.9-1.6 mm, forming a hood with the dorsal sepal; lip a little longer than the sepals, 2.8-4 × 1-2.3 mm, rhomboid in outline, tri-lobed, sometimes geniculate between the spur and the lobes, the central lobe tongue-shaped with apex curving up, obtuse, 1.5 × 0.5 mm, the lateral lobes triangular, obtuse, 0.5 × 0.7 mm; spur subcylindrical, trumpet-shaped at first then becoming thinner, pendent or trapped into the bract, narrowed from the base to the obtuse apex, whitish green, as long as the sepals, 2.3-6 × 0.3-0.6 mm. Anther not apiculate, suborbicular, 1-2.2 × 0.8-1.3 mm, with two distinct chambers; auricles very rounded, thick, almost a quarter the length of the anther; lateral lobes of the rostellum obsolete, the middle one obtuse, pointing backwards. Sweetly scented in the evening. 5898709058 1914-02 P H. Perrier Madagascar 2000 mont Ibity au sud d'Antsirabe Vakinankaratra 63 232 P00094574 1 holotype 5898708401 [976,1205,902,926] 1914-02 P H. Perrier Madagascar 2000 Ibity Vakinankaratra 63 232 P00094575 1 isotype 5898708799 P, K Baron Madagascar Madagascar 63 232 P00094569, K000415545 1 5898708868 P R. Baron 63 232 P00094570 1 5898708662 P R. Capuron Ankaratra Ankaratra 63 232 P00692268 1 5898708767 [989,1433,1443,1467] P A. M. Homolle 63 232 P00692276 1 5898708812 P B. Descoings Ihorombe -22.45 plaine de la Menarahaka 1260 46.25 63 232 P00692270 1 5898708724 BM G. F. Scott-Elliot Ankaratra 63 232 BM000034590 1 5898708354 P G. F. Scott-Elliot Ankaratra 63 232 P00334731 1 5898708831 [983,1356,1549,1574] K R. Baron Central Madagascar Central Madagascar 63 232 1 5898708665 P R. Bonet 63 232 P00326135 1 5898708924 TAN E. Ursch Tampoketsa d'Ankazobe Analamanga 63 232 1 5898708820 [1025,1171,1603,1627] P 63 232 P00692272 1 5898709042 1883-03 K, LDS Dr. Fox -18.9 Ambatovory 1273 47.7 Analamanga 63 232 1 5898708628 P C. M. Le Myre de Vilers Centre Centre 63 232 P00094474 1 5898709024 1905-06-12 L C. d'Alleizette -18.761667 Ambohimanga 21 47.56361 Analamanga 63 232 L1487720 1 5898708439 1906-01-20 P L. Rotereau -18.9 Analamanga 1273 47.7 63 232 P06796776 1 5898708919 1906-01 P C. d'Alleizette -18.9 Nanisana 1273 47.7 Analamanga 63 232 P00334732 1 5898708408 1912-04 P H. Perrier 1400 Manandona Vakinankaratra 63 232 P00094576 1 5898709125 1921-02 P H. Perrier 2000 -20.067501 mont Ibity 21 47.00111 Vakinankaratra 63 232 P00094578 1 5898709062 1921-03-27 P R. Decary -18.761667 Ambohimanga 21 47.56361 Analamanga 63 232 P00334738 1 5898708994 1921-04 P R. Decary -18.761667 Ambohimanga 21 47.56361 Analamanga 63 232 P00334739 1 5898708315 1923-02 P H. Perrier 2000 massif d'Andringitra Haute Matsiatra 63 232 P00094579 1 5898708375 1923-04-19 P R. Decary 1500 Ankaizina Diana 63 232 P00334737 1 5898708377 1923-04-19 P R. Decary 1500 64 233 Ankaizina Diana 63 232 P00334736 1 5898708636 1923-04-19 P R. Decary 1700 -18.9 Ankaizina 1273 47.7 Diana 64 233 P00334735 1 5898708786 1927-02 P H. Perrier Vakinankaratra 1600 entre Ambatolampy et Antsirabe 64 233 P00334733, P00334734 1 5898709043 1938-02-26 P P. Boiteau Andringitra Haute Matsiatra 64 233 P00692296, P00692297 1 5898708971 1941-04-11 P P. Boiteau 2000 -21.875 au sommet de l'Ambondrombe 21 47.260277 Haute Matsiatra 64 233 P00692437 1 5898708461 1945-02-24 P A. M. Homolle -18.083334 Ambohitantely 1276 47.266666 Analamanga 64 233 P00692274 1 5898708965 1948-02-07 TAN P. Boiteau -18.216667 Tampoketsa d'Ankazobe 1276 47.283333 Analamanga 64 233 1 5898708321 1951-03-18 1951-03-26 1951-03-18 P H. Humbert 2122 Diana 64 233 P02115322, P02115321 1 5898709128 1953-03-20 P ResNat -17.641666 Ambatosoratra 21 48.833332 Alaotra-Mangoro 64 233 P00692314 1 5898708890 1954-02-27 P R. Rakoto District Ambalavao Haute Matsiatra Canton Sendrisoa 64 233 P00692315 1 5898708580 1955-08 P J. Bosser Vakinankaratra -19.383333 Faratsiho 1272 47.266666 route d'Ambatolampy 64 233 P00692227 1 5898708888 1955-11 TAN J. Bosser Vakinankaratra Ampatolampy Ambohimandroro 64 233 1 5898708651 1957-02 P J. Bosser Vakinankaratra 2000 -19.533333 Ankaratra 1271 47.05 Nanokely 64 233 P00692226 1 5898708677 1960-02-26 P M. Peltier Vakinankaratra -19.533333 Nanokely 1271 47.05 Nanokely 64 233 P00692290 1 5898708606 1960-03 P J. Bosser -18.866667 Analamanga 1273 47.666668 64 233 P00692228 1 5898708657 1960-03-20 P M. Peltier Amoron'i Mania -20.575 Ilaka 21 46.958336 Fiadanana 64 233 P00692292 1 5898708477 1961-02 TAN J. Bosser Analamanga 64 233 1 5898708849 1961-03 P J. Bosser -19.356945 Manjakatompo 21 47.307777 Vakinankaratra 64 233 P00692229 1 5898709073 1962-09 P J. Bosser -19.0125 lake Mantasoa 21 47.838055 Analamanga 64 233 P00692230 1 5898708979 1963-02 P J. Bosser Vakinankaratra -19.533333 Ankaratra 1271 47.05 Nanokely 64 233 P00692241 1 5898708930 1963-03 P J. Bosser Route de Majunga Route de Majunga 64 233 P00692235 1 5898708499 1963-03 P J. Bosser -19.383333 Vakinankaratra 1272 47.266666 Vakinankaratra 64 233 P00692236 1 5898708828 1964-02 P J. Bosser route Ambatolampy-Faratsiho Vakinankaratra 64 233 P00692246, P00692247 1 5898708391 1964-04 P J. Bosser Haute Matsiatra -22.183332 sentier du pic Boby 1261 46.883335 64 233 P00692248, P00692249 1 5898708471 1964-04 P J. Bosser -22.216667 Andringitra 1260 46.916668 Haute Matsiatra 64 233 P00692251 1 5898708669 1964-04 P J. Bosser Andringitra Haute Matsiatra 64 233 P00692252, P00692253 1 5898708492 1964-04 P J. Bosser Vakinankaratra -20.242779 Col des Tapias 21 47.10111 Col des Tapias 64 233 P00692259 1 5898708823 1969-03-28 HEID 64 233 HEID250340 1 5898708782 1972-03 TAN G. Cremers 1900 -20.095556 Mt. Ibity western slope 21 47.003334 Vakinankaratra 64 233 1 5898708952 1985-03 TAN L. C. Barnett Vakinankaratra 1550 -20.166666 Mt. Ibity 1268 47.05 south of Antsirabe 64 233 1 5898708324 [304,498,1496,1520] MO 1268 64 233 MO3022638 1 5898708936 1989-03 P Goldblatt Haute Matsiatra 1975 -22.183332 Andringitra Forest Reserve 1261 46.916668 64 233 P00692435 1 5898708522 [132,341,1576,1600] MO 1261 64 233 MO4314431 1 5898708647 [348,412,1576,1600] TAN 1261 64 233 1 5898708639 [420,652,1576,1600] WAG 1261 64 233 WAG.1130546 1 5898708676 1989-03 MO Goldblatt 2350 -19.333334 Ankaratra Massif 1272 47.266666 Vakinankaratra 64 233 MO4314448 1 5898708349 [131,184,1656,1680] TAN 1272 64 233 1 5898709089 1993-05 K C. H. Jongkind Amoron'i Mania 1280 -20.178333 on hillside near Talavina 21 47.09 64 233 1 Antananarivo 5898708937 [572,775,1710,1734] MO 21 64 233 MO3022616 1 Antananarivo 5898708317 [131,287,1736,1761] P 21 64 233 P00692179 1 Antananarivo 5898708781 [291,344,1737,1761] TAN 21 64 233 1 Antananarivo 5898708310 1995-04 K D. DuPuy 1400 -20.178055 Col Tapia 21 47.087776 Vakinankaratra 64 233 M906 1 Antananarivo 5898708458 [284,440,1790,1814] P 21 64 233 P00059605 1 Antananarivo 5898708346 [444,499,1790,1814] MO 21 64 233 1 Antananarivo 5898708569 [503,627,1790,1814] WAG, TAN 21 64 233 1 Antananarivo 5898708931 1999-03-18 P J. N. Labat -17.86861 Tampoketsa d'Ankazobe 21 47.060833 Analamanga 64 233 P00160016, P00160017 1 Antananarivo 5898709006 [561,592,1843,1867] G 21 64 233 1 Antananarivo 5898709015 [600,629,1843,1867] K 21 64 233 1 Antananarivo 5898708563 [637,754,1843,1867] MO, TAN 21 64 233 1 Antananarivo 5898709037 K J. Hermans Between Antananarivo and Ambositra 64 233 1 Antananarivo 5898708381 2001-04 K J. Hermans Vakinankaratra 1440 between Antsirabe and Ambositra 64 233 1 Antananarivo 5898709055 2001-05 K J. Hermans Vakinankaratra 1740 -20.240278 between Antsirabe and Ambositra 21 47.103333 64 233 1 Antananarivo 5898708455 2007-01-25 WU A. Sieder & J. Hermans Amoron'i Mania 1701 -20.761944 Ivato 21 47.325275 Ambositra 64 233 WU0063773 1 Antananarivo 5898708446 2010-02-21 K L. Rajavelona 1647 -20.570002 Ambatofinandrahana 21 46.581944 Amoron'i Mania 64 233 RJL148, K664548 1 Antananarivo 5898708600 [813,868,1459,1483] TAN 21 64 233 1 Antananarivo 5898708416 2015-02 WU A. & C. Sieder 1676 -20.774166 Antoetra 21 47.32111 Amoron'i Mania 64 233 1 Antananarivo 5898708634 2017-02 WU A. & C. Sieder inselberg 64 233 1 Antananarivo 5898708355 2018-04 K J. Hermans Vakinankaratra between Antsirabe and Ambositra 64 233 1 Antananarivo 5898708742 2019-03-19 WU A. & C. Sieder 1461 -18.266111 Analamanga 21 47.180836 64 233 1 Antananarivo 5898708652 2019-03 WU A. & C. Sieder 1461 Analamanga 64 233 L 1 Antananarivo