Sixty new dragonfly and damselfly species from Africa (Odonata) Dijkstra, Klaas-Douwe B. Kipping, Jens Mézière, Nicolas Odonatologica 2015 44 4 447 678 urn:lsid:zoobank.org:act:03A25264-CA05-FFCA-EE87-FABA43BDFD6E Dijkstra, Kipping & Meziere Dijkstra, Kipping & Meziere [165,759,1365,1394] Insecta Chlorocyphidae Chlorocypha GBIF,CoL Animalia Odonata 28 473 Arthropoda species maxima sp. nov.   TaxonomyThis species was initially identified as  C. dahliFraser, 1956that is known with certainty only from the holotypein ZMUCfrom eastern Cameroon, but its character combination is unique compared with photographs of that specimen provided by N.P. Kristensen. It is also genetically notably distinct from other chlorocyphids.    Material studied     Holotype  ♂ . RMNH.INS.508083, Gabon, Ngounie Province, Montagne de Chaillu, 21 kmNE of Moukabou, small sandy stream in dense forest,  641 ma.s.l. ( 1.4816 ° S 11.8009 ° E),  18 -ix- 2013, leg. J. Kipping, RMNH.  Further material.  GABON( Haut-Ogooué Province): 4 ♂,  33 km S of Franceville, Mopia, Bai on Djoumou River, large sandy river,  414 ma.s.l. ( 1.9219 ° S 13.6759 ° E),  11 -i- 2010, leg. K.- D.B. Dijkstra, N. Mézière& C.Vanappelghem, RMNH.  3 ♂, Bakoumba River between Bakoumba-Ville and -Village, sandy river in rainforest,  544 ma.s.l. ( 1.8576 ° S 13.0148 ° E),  02-i- - 2010, leg. K.- D.B. Dijkstra& N. Mézière, RMNH.  2 ♂, La Lékédi, Bakoumba, large sandy river in forest,  591 ma.s.l., ( 1.7594 ° S 12.9501 ° E),  30 -iv- - 2009, leg. C. Vanappelghem& N. Mézière, RMNH.  1 ♂, Moanda, sandy forest stream ( 2–3 mwide),  423 ma.s.l. ( 1.5555 ° S 13.1195 ° E),  07-iii- 2010, leg. N. Mézière, RMNH.  1 ♂, same locality,  28 -ii- 2010, leg. N. Mézière, RMNH.  1 ♂, forest road on right at exit of Moanda, Kounda, sandy stream ( 4 mwide, 50 cmdeep),  423 ma.s.l. ( 1.5555 ° S 13.1195 ° E),  13 -xi- - 2010, leg. N. Mézière, RMNH.  1 ♂, Etéké road, Yeno, sandy and gravelly forest stream,  527 ma.s.l. ( 1.5845 ° S 11.5093 ° E),  23 -iv- 2011, leg. N. Mézière, RMNH.  1 ♂, les eaux claires, Bakoumba, small sandy/gravel river in forest (<  2 m),  680ma.s.l. ( 1.6812 ° S 12.9018 ° E),  30 -iv- 2009, leg. C. Vanappelghem& N. Mézière, RMNH.  1 ♂, Bakoumba road, Landébéngué, sandy forest river ( 4 mwide, 40–80 cmdeep),  404 ma.s.l. ( 1.6234 ° S 13.1330 ° E),  17 -x- - 2010, leg. N. Mézière, RMNH.  1 ♂, Moanda-Bakoumba road, Lemagna, sandy forest stream ( 2–3 mwide),  493 ma.s.l. ( 1.6132 ° S 13.0945 ° E),  26 - ii- 2011, leg. N. Mézière, RMNH.  1 ♂, Bakoumba road, village 3 kmbefore Kounda, Mikouangna (Mouyeugue), sandy river (> 2 mwide),  520 ma.s.l. ( 1.5911 ° S 13.1552 ° E),  26 -v- 2009, leg. N. Mézière, RMNH.  1 ♂, Bakoumba road, Kounda, Mbéressé,sandy forest river, partly open and sunny ( 3 mwide, 50–100 cmdeep),  484 ma.s.l. ( 1.5859 ° S 13.1563 ° E),  06-x- 2012, leg. N. Mézière, RMNH.  1 ♂, Konda,  11 km W of Moanda, sandy stream in dense forest, 423 ma.s.l. ( 1.5555 ° S 13.1195 ° E),  30 -i- 2012, leg. J. Kipping, CJKL.   1 ♂, Mopia,  428 ma.s.l. ( 1.9132 ° S, 13.6347 ° E),  01-ii- 2012, leg. A. Günther, CAGF.  1 ♂, Makokou, broad river with swamps,  494 ma.s.l. ( 0.5577 ° N 12.6141 ° E),  30 -xii- 2011, leg. N. Mézière, CAGF. GABON( Ngounie Province):  1 ♂( RMNH.INS.508836), Parc National des Monts Birougou, sandy forest stream ( 4 mwide, 50 cmdeep) ( Photo 10),  747 ma.s.l. ( 1.9655 ° S 12.2188 ° E),  05-ix- 2013, leg. N. Mézière, RMNH.    Genetics Two unique haplotypes (n = 2) nearest to  C. helenaeLegrand, 1984and highly distinct from other  Chlorocyphaspecies, although  C. dahliwas not available for analysis.    Male morphological diagnosis Similar to  C. dahliby its (a) largely bright occiput, postclypeus, genae and mandibular bases that contrast with the wholly black anteclypeus and labrum and largely black frons; (b) bright ante-humeral stripes that cover most of the mesepisterna; (c) red abdominal dorsum with black markings reduced to a pair of large subapical dots on S 2 and smaller pairs on S 3–4 and sometimes S 5–6 ( Fig. 2); and (d) boldly black and yellow abdomen underside. However, unlike it and most  Chlorocyphahas (1) large size, Hw 25.5 – 28.0 mm (n= 10) rather than 23.0 mm (n = 1); (2) olive-green rather than orange markings on the head and thorax; (3) blue-white apical 75–80 % of the anterior face of the hind tibiae and apical 0–30 % of the mid tibiae, rather than yellow-white apical 80 % of the front and mid tibiae; and (4) yellow dorsum of S 10 contrasting with red on S 2–9.  Chlorocyphaspecies often have white and sometimes yellow streaks on both the mid and hind tibiae, and sometimes on the fore tibiae as well, but both their presence and absence on only the hind pair is unusual, as is their apical rather than basal position.    Etymology Latin “greatest” refers to the species’ exceptional size (feminine adjective).    Range and ecology Calm sandy but often somewhat turbid (i.e., silty) streams in rainforest between 400 and 750 ma.s.l., especially in and around the Chaillu Mountains of southern Gabon.   Pentaphlebiidae 1224547663 2013-09-18 RMNH J. Kipping Gabon 641 m http://www.boldsystems.org/index.php/Public_RecordView?processid=NSPAF436-15 -1.4816 Montagne de Chaillu, 21 km NE of Moukabou 7 11.8009 29 474 RMNH.INS.554425 1 1 Ngounie Province holotype 1224547671 2010-01-11 RMNH D. B. Dijkstra, N. Mézière, C. Vanappelghem Gabon 414 m -1.9219 S of Franceville, Mopia, Bai on Djoumou River 7 13.6759 29 474 4 4 Haut-Ogooué Province 1224547660 2010-01-02 RMNH D. B. Dijkstra, N. Mézière Gabon 544 m -1.8576 Bakoumba River between Bakoumba-Ville and - Village 7 13.0148 29 474 3 3 Haut-ogooue Province 1224547661 2009-04-30 RMNH C. Vanappelghem, N. Mézière Gabon 591 m -1.7594 La Lékédi, Bakoumba 7 12.9501 29 474 2 2 Haut-ogooue Province 1224547677 2010-03-07 RMNH N. Mézière Gabon 423 m -1.5555 Moanda 7 13.1195 29 474 1 1 Haut-ogooue Province 1224547653 2010-02-28 RMNH N. Mézière Gabon 423 m -1.5555 forest road on right at exit of Moanda, Kounda 7 13.1195 29 474 1 1 Haut-ogooue Province 1224547662 2010-11-13 RMNH N. Mézière Gabon 423 m -1.5555 forest road on right at exit of Moanda, Kounda 7 13.1195 29 474 1 1 Haut-ogooue Province 1224547656 2011-04-23 RMNH N. Mézière Gabon 527 m -1.5845 Etéké road, Yeno 7 11.5093 29 474 1 1 Haut-ogooue Province 1224547655 2009-04-30 RMNH C. Vanappelghem, N. Mézière Gabon 680 m -1.6812 les eaux claires, Bakoumba 7 12.9018 les eaux claires, Bakoumba 29 474 1 1 Haut-ogooue Province 1224547674 2010-10-17 RMNH N. Mézière Gabon 404 m -1.6234 Bakoumba road, Landébéngué 7 13.133 29 474 1 1 Haut-ogooue Province 1224547658 2011-02-26 RMNH N. Mézière Gabon 493 -1.6132 Moanda-Bakoumba road, Lemagna 7 13.0945 29 474 1 1 Haut-ogooue Province 1224547669 2009-05-26 RMNH N. Mézière Gabon 520 m -1.5911 Bakoumba road, village 3 km before Kounda, Mikouangna (Mouyeugue) 7 13.1552 29 474 1 1 Haut-ogooue Province 1224547675 2012-10-06 RMNH N. Mézière Gabon 484 m -1.5859 Bakoumba road, Kounda, Mbéressé, 7 13.1563 29 474 1 1 Haut-ogooue Province 1224547672 2012-01-30 CJKL J. Kipping Gabon -1.5555 W of Moanda 7 13.1195 Konda 29 474 1 1 Haut-ogooue Province 1224547670 2012-02-01 CAGF A. Günther Gabon 428 m -1.9132 Mopia 7 13.6347 30 475 1 1 Haut-Ogooué Province 1224547666 2011-12-30 CAGF N. Mézière Gabon 494 m 0.5577 Ngounie Province 7 12.6141 30 475 1 1 Haut-ogooué Province 1224547668 2013-09-05 RMNH N. Mézière Gabon 747 m http://www.boldsystems.org/index.php/Public_RecordView?processid=NSPAF518-15 -1.9655 Parc National des Monts Birougou 7 12.2188 Parc National des Monts Birougou 30 475 RMNH.INS.554425 1 1 Ngounie Province