Dorilas (Eudorylas) excisus Hardy 1949 a : 32 Dorilas (Eudorylas) definitus Hardy 1961: 134 De Meyer and Földvári 2008: 81 Taxonomic revision of the Afrotropical species of the tribe Eudorylini (Diptera, Pipunculidae) FÖLDVÁRI, MIHÁLY Zootaxa 2013 2013-05-22 3656 1 1 121 3C2TN (Hardy, 1949) Hardy 1949 [598,988,1558,1584] Insecta Pipunculidae Eudorylas Animalia Diptera 57 58 Arthropoda species excisus  ( Figs 19G–M, 39G–H, V)      Dorilas (Eudorylas) excisus Hardy 1949 a: 32.     Dorilas (Eudorylas) definitus Hardy 1961: 134- syn.:  De Meyer and Földvári 2008: 81.  Diagnosis: Surstyli turned ventrally, both with two projections at the end; membranous area large, almost reaching upper edge of ST8. Ventrally on the base of SS and on SES ca. 10 hairs on each side; gonopods asymmetrical, both with ridges; phallic guide with membranous parts at the end covered by microtricha; in lateral view there is a hook pointing towards SS; phallus long, coiled, trifid.  Type material:   Malawi: 1♂, 1♀, HT, AT [on the same supporting pin], 5.4.16., Ruo Nyasaland, R.C. Wood, 493, Pres by Com. Inst. Ent., BM 1953-357. ( BMNH).   Democratic Republic of Congo: 1♂[HT of  E. definitus], Congo belge, P.N.G., Miss[ion] H. De Saeger, I/a/3,  7-ii-1950; Réc. H. De Saeger 199;  1♀[AT of  E. definitus], Congobelge P.N.G, Miss[ion] H. De Saeger; Inimvua,  20-v-1952, H. De Saeger, 3488 (both MRAC).   FIGURE 19.Male genitalia of  Eudorylas encerus(A–F) and  E. excisus(G–M).—A: outer surstylus with phallic guide, lateral view, B: dorsal view, C: inner surstylus with epandrial process, lateral view, D: surstyli, dorsal view, E: sperm pump with ejaculatory apodeme, F: ventral view with phallic guide and ejaculatory apodeme (without ST8), G: outer surstylus, lateral view, H: dorsal view, I: inner surstylus, lateral view, J: sperm pump with ejaculatory apodeme, K: phallic guide, gonopods and hypandrium, ventral view, L: phallic guide and phallus lateral view, M: surstyli, dorsal view. Scale bars: 0.1 mm Other material examined:   Angola: 1♂, (A17), Tundavala, c15 mls NW. Sa da Bandeira, c.6600’, 14°43’S 13°28’E,  3.iii.1972, southern Africanexp. BM 1972–1 ( BMNH).   Kenya: 2♂, Karura For., Nairobi,  5500 feet,  9– 13.xii.1970, A.E. Stubbs, B.M. 1972–211 ( BMNH, HNHM);  1♂, Ngong Forest Station  IV-13-17-1968, Paul Spangler( USNM).   Mozambique: 2♂, 1♀, Niassa Prov., Cuamba, Farmof the Catholic University of Mozambique, 17-IV/  2-VI- 2006, 600m, Olmi M.coll. (all MRAC).   Namibia: 1♂, 1♀, West Caprivi Park, Kwando River: Susuwe, 17°45'37''S 23°20'55''E,  28.ix.-02.x.1998, A.H. Kirk-Spriggs, Malaise trap, dry woodland;  1♀, Rundu district, Simanya, Okavango River, 17°33'17''S 18°32'30''E,  23-24.i.1998, Kirk-Spriggs& Marais, malaise traps, riverine forest (all NMNW);  Windhoek, 22°34'S 17°05'E,  8.12.1986, coll. Malaisetrap, J. Irish( HNHM).   South Africa: 1♂, Natal, Richards Bay,  24.10.1994, leg. Roy Danielsson( ZML);  1♂, TsitsikammaN.P., 34°51'17''S 23°53'22''E,  08.iv.1998, A.H. Kirk-Spriggs, sweeping understory veg. in coastal rainforest ( NMNW);  1♂, Barberton, 17.5.13., H.K. Munro;  1♂, Zuurberg Range, north of Addo, E. Cape Prov.,  10 April 1961, collectors B. and P. Stuckenberg;  1♂, Otto’s Bluff, P.M. Burg, 15.X.72, M.E. Irwin;  1♂, Natal, St. Lucia Nature Res., 2832AD,  18–20.xii.1981, Londt& Stuckenberg, Coastalbush & forest;  1♂, Natal Prov., Richmond Dist.Pateni, 30º09’E., 29º56’S.(2930 Cc),  Nov.18, 1971; temp. forest, BE Stuckenberg& ME Irwin(all NMSA);  4♂, Natal#6, Kube Yini Game Res,. 27º48’S: 32º14’E,  360m, Date:  10–14.i. 1994, Natal Museum Expedition(3 NMSA, 1 HNHM);  4♂, Natal Prov., Zululand, 21 mi. S.Ndumu, Game Res. Camp(2732 Aa), Dec, 1, 1971, ME& BJ Irwin, dry scrub forest;  160ft(3 NMSA, 1 HNHM).   Swaziland: 3♂, #45, 13 km N. of Ngogolo, 26º19’S: 31º38’E,  300m, Date  22–24. iv.1991, J Londt& L Schoeman, Panata Ranch/ Bushveld(2 NMSA, 1 HNHM).   Zimbabwe: 1♂, N. Vumba, S. Rhodesia,  25.2.1965, D. Cookson;  1♂, same data,  25.4.1964(both NMSA);  1♂, Rhodesia: Victoria Falls Nat’l. Park,  IV-3-6-1968, Paul Spangler( USNM).  Male  Head. Third antennal segment acuminate; yellow-brown (arista dark brown). Face silvery pollinose. Frons, upper part shining black, lower part silvery pollinose; eyes touching for distance equal to 3 times ocellar triangle. Occiput, greyish pollinose.  Thorax. Humeri yellowish brown. Mesonotum (viewed obliquely from front) black with brown pollinosity, somewhat silvery along anterior margin; brown with greyish pollinosity on the side. Scutellum black with faint brownish-grey pollinosity, with 4–6 short and very weakly developed white hairs. Dorsocentral hairs weakly developed. Halter yellow, base brown. Legs. Trochanters and femora yellow with dark brown ring in the middle 1/3–3/5 of the latter, f3 shining posteroventrally, knees, tibiae yellow. Tarsal segments yellow, last segment brown. Ventroapical row of 0–2 and 4–8 very weakly developed dark spines on first and second femora respectively; no spines on 3rd femur. Subapical (distal) spines on first four tibiae present. No anteromedial hairs on 3rd tibia. Hind trochanter with no hairs or spines on ventral side. Pulvilli shorter than last tarsal segment. Wing. Fourth costal section 0.6–0.8 times as long as third costal section. Cross-vein R-M beyond 2/5 of discal cell. Pterostigma fully coloured. No hairs on tegula.  Abdomen. Viewed obliquely from front tergites brownish dusted, T1 greyish, hind margins brownish dusted, T3,4 with grey patches laterally, distal half of T5 greyish pollinose, sides greyish pollinose. Hairs dispersed, short and weakly developed. Laterally 1–5 dark spines on first tergite. Postabdomen in dorsal view: T6 (or T7) visible; T5 1.4 times as long as ST8. Genitalia without dissection: middle sized ST8, m.a. as an elongated triangle, EP and SS yellow, otherwise black. Genitalia. Surstyli turned ventrally, both with two projections at the end; epandrium elongated; ST8 extending to the right, membranous area large, almost reaching upper edge of ST8 ( Fig. 19H). Ventrally on the base of SS and on SES ca. 10 hairs on each side; gonopods asymmetrical, both with ridges, OG longer; hypandrium with distal part (lobes) turned; phallic guide with membranous parts at the end covered by microtricha; phallus long, coiled, trifid ( Fig. 19K); ejaculatory apodeme flat, fan-shaped (can be linear); sperm pump simple, flat, with two projections at the end ( Fig. 19J). In lateral view PG has a hook pointing towards SS; SES present, although connection to GP is no clear; PHwith supporting structure ( Fig. 19L). Female As male except for the following characters. Frons, eyes separated, silvery pollinose, except shining black narrow line in the middle. Pulvilli and last tarsal segment (brownish) about the same size; claws 1.5 times as long as pulvilli. No ventroapical spines on femora. Hind trochanter covered with very fine white hairs. Fourth costal section equal to third. Cross-vein R-M almost as far as middle of discal cell. Abdominal T1 is not greyish, sides of tergites in general brownish dusted. Female ovipositor as in Figs 39G–H, V.   Distribution— Angola, Democratic Republic of Congo, Malawi, Kenya, Mozambique, Namibia, South Africa, Swaziland, Zimbabwe. 3462316679 [323,574,1901,1926] BMNH Malawi Ruo Nyasaland 57 58 1 1 holotype 3462316387 BMNH Malawi Ruo Nyasaland 57 58 1 1 alotype 3462316675 1950-02-07 MRAC Democratic Republic of the Congo Congo belge 57 58 1 1 Congo belge 3462316589 57 58 3462316549 1972-03-03 BMNH Angola Tundavala -14.716666 southern African 1287 13.466666 Sa da Bandeira 59 60 1 1 3462316341 1970-12-09 1970-12-13 1970-12-09 BMNH, HNHM A. E. Stubbs Kenya 1676 Nairobi Karura For. 59 60 2 2 3462316303 1968-04-13 1968-04-17 1968-04-13 USNM Paul Spangler Kenya Ngong Forest Station 59 60 1 1 3462316670 2006-06-02 MRAC Olmi M. Mozambique Cuamba Catholic University of Mozambique Farm 59 60 3 1 2 Niassa 3462316607 1998-09-28 1998-10-02 1998-09-28 Kirk-Spriggs Namibia West -17.760279 Kwando River 21 23.348612 Park 59 60 2 1 1 Zambezi 3462316562 1998-01-23 1998-01-24 1998-01-23 NMNW Kirk-Spriggs & Marais Namibia Rundu district -17.554722 River 21 18.541666 Simanya 59 60 1 1 Kavango West 3462316516 [556,1432,402,426] 1986-08-12 HNHM Malaise & Irish Namibia -22.566668 Windhoek 1259 17.083334 59 60 1 3462316597 [151,1143,437,462] 1994-10-24 ZML Roy Danielsson South Africa Richards Bay Natal 59 60 1 1 3462316326 1998-04-08 NMNW Kirk-Spriggs South Africa -34.85472 Tsitsikamma 19 23.889444 59 60 1 1 3462316356 [151,597,509,533] H. K. Munro South Africa Barberton 59 60 1 1 3462316653 1961-04-10 Addo, E. & B. and P. Stuckenberg South Africa Zuurberg Range 59 60 1 1 Western Cape 3462316560 [538,1110,545,570] 59 60 3462316579 1981-12-18 1981-12-20 1981-12-18 Londt & Stuckenberg South Africa St. Lucia Nature Res. Natal 59 60 1 1 3462316547 1971-11-18 NMSA BE Stuckenberg & ME Irwin South Africa -29.933332 Richmond Dist. 1224 30.15 Natal Prov. 59 60 1 1 3462316569 1994-01-10 1994-01-14 1994-01-10 NMSA, HNHM Natal Museum Expedition South Africa Natal 360 -27.8 Date 1235 32.233334 Kube Yini Game Res 59 60 4 4 3462316439 NMSA, HNHM ME & BJ Irwin South Africa Natal Prov. 49 Game Res. Camp Zululand 59 60 4 4 3462316652 1991-04-22 1991-04-24 1991-04-22 NMSA, HNHM J Londt & L Schoeman Swaziland 300 -26.316668 Date 1242 31.633333 Ngogolo 59 60 3 3 3462316349 [151,903,794,819] 1965-02-25 D. Cookson Zimbabwe Zimbabwe 59 60 1 1 3462316416 [916,1380,795,819] 1964-04-25 NMSA D. Cookson Zimbabwe Zimbabwe 59 60 1 1 3462316687 1968-04-03 1968-04-06 1968-04-03 USNM Paul Spangler Zimbabwe Victoria Falls Nat'l. Park Rhodesia 59 60 1 1