Maja gibba Griffin, 1976: 200 Sakai, 1976: 239 Griffin & Tranter, 1986: 216 Maja gibba Alcock, 1895 Maja kominatoensis Ng et al., 2008: 123 Paramaja kominatoensis Kubo, 1936 Revision of the spider crab genus Maja Lamarck, 1801 (Crustacea: Brachyura: Majoidea: Majidae), with descriptions of seven new genera and 17 new species from the Atlantic and Indo-West Pacific Ng, Peter K. L. Forges, Bertrand Richer De Raffles Bulletin of Zoology 2015 2015-05-29 63 110 225 4D295 Ng & Forges, 2015 Ng & Forges 2015 [1021,1218,1021,1045] Malacostraca Majidae Paramaja Animalia Decapoda 25 135 Arthropoda species turgida sp. nov.  ( Figs. 6H,  12, 13E, F, 14H, 15N–U, 36J, K, 39K, 43N–P, 48H, I, 53L, 55I, J, 68D–F)     Maja gibba–  Griffin, 1976: 200. – T.  Sakai, 1976: 239(part). –  Griffin & Tranter, 1986: 216(part). (not  Maja gibba Alcock, 1895).    Maja kominatoensis–  Ng et al., 2008: 123, Fig. 90. (not  Paramaja kominatoensis Kubo, 1936).   Material examined.  Holotype: male (74.1 × 66.8 mm) ( NMCR), Balicasag Island, Panglao, Bohol, Philippines,  200–300 m, coll. fishermen with tangle nets,  28 November 2001.  Paratypes:  Philippines– 1 male(71.4 × 64.4 mm) (NSMT-Cr 22329),  1 male(55.4 × 48.4 mm), 1 ovigerous female (62.9 × 57.2 mm) ( AM, ex ZRC 2001.0615), same data as holotype. — 1 young male (26.2 × 21.3 mm) ( ZRC 2001.0616b), Balicasag Island, Panglao, Bohol, coll. fishermen with tangle nets,  28 November 2001. —  1 male(69.2 × 64.0 mm), 1 ovigerous female (47.1 × 41.1 mm) ( ZRC 2013.1185), Balicasag Island, Panglao, Bohol,  200–300 m, coll. fishermen with tangle nets,  June 2002. — 1 ovigerous female (dried) (58.4 × 53.4 mm) ( ZRC 2009.0972), Banacon Island, Bohol, coll. local fisherman, 2000. —  3 females(NSMT-Cr 15388), Balicasag Island, Panglao, Bohol, coll. local fisherman via M. Takeda& H. Komatsu,  February 2003. — 2 males(34.3 × 29.2 mm, 56.4 × 49.7 mm),  1 female(60.1 × 55.6 mm, with anemone) ( ZRC 2013.1239), Balicasag, Panglao, Bohol,  200–300 m, coll. local fishermen with tangle nets,  January 2004. —  1 female(61.3 × 56.6 mm) ( ZRC 2013.1240), Balicasag, Panglao, Bohol,  200–300 m, coll. local fishermen with tangle nets,  February 2004. —  2 males(70.6 × 62.2 mm, 57.8 × 51.5 mm), 2 ovigerous females (60.2 × 53.1 mm, 62.2 × 55.2 mm) ( ZRC 2012.1203), Balicasag, Panglao, Bohol,  200–300 m, coll. local fishermen with tangle nets,  March 2004. —  1 female(61.9 × 58.4 mm) ( ZRC 2013.1235), northwest Panglao, Bohol,  120–160 m, tangle nets, coll. J. Arbasto,  July 2004–  May 2005. —  1 male(70.4 × 63.9 mm) ( ZRC 2012.1201), station CP 2332, off Pamilacan Island, Bohol Sea, 9°38.8’N 123°45.9’E, coll. PANGLAO 2005,  22 May 2005. —  1 male(70.9 × 62.2 mm) ( ZRC 2012.1202), station CP 2372, Bohol/ Sulu Seas, 9°31.4’N 124°00.6’E, PANGLAO 2005,  24 May 2005. —  1 male(70.5 × 63.1 mm) ( ZRC 2012.1204), station CP 2397, off Aligbay Island, Maribojoc Bay, Bohol Sea, 9°34.9’N 123°41.7’E, coll. PANGLAO 2005,  31 May 2005. —  1 male(36.4 × 31.2 mm) ( MNHN), station CP 2406, rock-sand-coral substrate, 9°38.8’N 123°46.5’E– 9°39.7’N 123°46.6’E,  331–334 m, Bohol Sea, coll. PANGLAO 2005,  1 June 2005. —  1 female(59.6 × 54.5 mm) ( MNHN), station CP 2383, sandy substrate, 8°43.1’N 123°19.2’E– 8°44.0’N 123°18.7’E,  253–338 m, Bohol Sea, coll. PANGLAO 2005,  29 May 2005. —  1 male(28.0 × 22.5 mm) ( ZRC 2013.1237), station CP 2331, muddy substrate, 9°37.4’N 123°45.9’E– 9°38.9’N 123°46.9’E,  268.2–273 m, Bohol Sea, coll. PANGLAO 2005,  22 May 2005. —  1 female(31.0 × 26.5 mm) ( ZRC 2013.1231), station CP 2341, sandy-muddy substrate, 9°26.1’N 123°48.4’E– 9°24.9’N 123°49.3’E,  381–544 m, Bohol Sea, coll. PANGLAO 2005,  23 May 2005. —  1 female(62.6 × 57.2 mm) ( ZRC 2013.1241), station CP 2363, sandy substrate, 9°0.1’N 123°26.5’E– 9°0.3’N 123°25.8’E,  439–452 m, Bohol Sea, coll. PANGLAO 2005,  26 May 2005. —  1 female(51.0 × 45.1 mm) ( ZRC 2013.1236), station CP 2392, off Balicasag Island, sandy-muddy substrate, 9°30.1’N 123°43.4’E– 9°29.5’N 123°42.6’E,  242–313 m, Bohol Sea, coll. PANGLAO 2005,  30 May 2005. —  1 female(50.5 × 44.8 mm, with rhizocephalan, photographed) ( ZRC 2013.1236), station CP 2331, muddy substrate, 9°37.4’N 123°45.9’E– 9°38.9’N 123°46.9’E,  268–273 m, Bohol Sea, coll. PANGLAO 2005,  22 May 2005. —  1 female(33.5 × 29.0 mm) ( ZRC 2013.1230), station CP 2396; sandy substrate, 9 °37.1N123°445’E – 9°36.7N 123°42.9’E,  467–609 m, Bohol Sea, coll. PANGLAO 2005,  31 May 2005. —  1 female( ZRC 2013.1234), station CP2405, Maribojoc Bay, Panglao, 9°39.0’N 123°46.1’E,  387–453 m, coll. PANGLAO 2005,  1 June 2005. —  1 male(67.9 × 60.5 mm) ( MNHN), station CC 2743, eastern coast of Luzon Island, coll. AURORA2007,  2 June 2007. —  1 female(42.2 × 37.6 mm) (MNHN- IU-2013-4055), station CP 53, 13°59’N 120°18’E,  215–216 m, coll. RV  Coriolis, MUSORSTOM 2,  27 November 1980. —  1 male(33.7 × 28.4 mm) ( USNM 48508), station 5403, between Leyteand Cebu, 182 fathoms, coll. RV  Albatross,  16 March 1909.  Non-type: Philippines–fragments ( USNM 48509), station 5536, 279 fathoms, coll. RV  Albatross,  19 August 1909.   Solomon Islands– 1 male(25.1 × 19.6 mm) (MNHN-IU-2013-4051), station CP 1851, 10°28’S 162°00’E,  297–350 m, coll. RV  Alis, SALOMON 1,  6 October 2001.   Diagnosis.Adult carapace with branchial and gastric regions ovate; dorsal surface convex, covered by numerous rounded and sharp granules, those on median row larger, tuberculate, forming low crest ( Figs. 12, 13E, F, 14H). Adult pseudorostral spines almost straight, gently diverging, appears dorso-ventrally flattened ( Figs. 12, 13E, F, 36J, K). Supraorbital eave narrow, granulated, with antorbital spine triangular, sharp; intercalated spine irregular, slightly shorter than supraorbital spine; postorbital spine wide, pointing anteriorly, triangular with some granules at base of anterior margin; hepatic area inflated with several granules, largest spiniform, pointing outwards ( Figs. 12, 13E, F, 36J, K). Adult male anterior thoracic sternum relatively narrow ( Fig. 48H, I); lateral margins of anterior edge of male sterno-abdominal cavity straight ( Fig. 48H, I). Distal part of G1 relatively shorter, tip not bent, in line with rest of G1 ( Fig. 15N–U).   Etymology.From the Latin “ turgidus” for swollen, alluding to the carapace appearance of the species.   Remarks.  Paramaja turgida n. sp.is closest to  P. gibba, but can be separated by its proportionately narrower male anterior thoracic sternum ( Fig. 48H, I) (versus relatively broader in  P. gibba, Fig. 48F, G) and the distal part of the G1 is relatively shorter with the tip in line with the rest of the structure ( Fig. 15N–U) (versus relatively longer with the tip gently bent laterally in  P. gibba, Fig. 15K–M). The chelipeds of adult females of  P. turgidaare slender and not inflated, while the ambulatory legs are distinctly shorter and more setose ( Fig. 12B, D). The smallest ovigerous female measured 47.1 by 41.1 mm(ZRC 2013.1185). A young male specimen from the Solomon Islands(25.1 × 19.6 mm, MNHN-IU-2013-4051) ( Fig. 12F) is tentatively referred to  P. turgida. The G1 resembles that of  P. turgida, but at this size, it is not easy to separate from  P. kominatoensis. The less inflated carapace shape of the specimen is closest to  P. turgidaand as such, we provisionally refer it here. Adult males will need to be collected to be certain about their identity as the Solomon Islandsis some distance from the Philippines. See also discussion for  P. gibbaand  P. kominatoensis. 2001-11-28 NMCR Philippines 250 Balicasag Island Bohol 25 135 1 holotype Philippines Philippines 25 135 1 1 paratype 2001-11-28 AM Philippines Philippines 250 Balicasag Island Bohol 25 135 ZRC 2001.0615 1 1 paratype 2001-11-28 ZRC Philippines Panglao Balicasag Island 25 135 ZRC 2001.0616 1 Bohol paratype 2002-06 ZRC Philippines 250 Panglao Balicasag Island 25 135 ZRC 2013.1185 1 1 Bohol paratype ZRC Philippines Banacon Island 25 135 ZRC 2009.0972 1 Bohol paratype 2003-02 M. Takeda & H. Komatsu Philippines Panglao Balicasag Island 25 135 3 3 Bohol paratype 2004-01 ZRC Philippines 250 Panglao Balicasag 25 135 ZRC 2013.1239 1 1 Bohol paratype 2004-02 ZRC Philippines 250 Panglao Balicasag 25 135 ZRC 2013.1240 1 1 Bohol paratype 2004-03 ZRC Philippines 250 34 144 Panglao Balicasag 25 135 ZRC 2012.1203 2 2 Bohol paratype 2004-07 2005-05 2004-07 ZRC J. Arbasto Philippines 140 Panglao 34 144 ZRC 2013.1235 1 1 Bohol paratype 2005-05-22 ZRC Sea Philippines 9.646667 Pamilacan Island 129 123.765 34 144 ZRC 2012.1201 1 1 Bohol paratype 2005-05-24 ZRC Seas Philippines 9.523334 Bohol 129 124.01 34 144 ZRC 2012.1202 1 1 Bohol paratype 2005-05-31 ZRC Sea Philippines 9.581667 Maribojoc Bay 129 123.695 Aligbay Island 34 144 ZRC 2012.1204 1 1 Bohol paratype 2005-06-01 MNHN Philippines 333 9.661667 Sea 129 123.776665 34 144 1 1 Bohol paratype 2005-05-29 MNHN Philippines 296 8.733334 Sea 129 123.31167 34 144 1 1 Bohol paratype 2005-05-22 ZRC Philippines 271 9.648334 Sea 129 123.78167 34 144 ZRC 2013.1237 1 1 Bohol paratype 2005-05-23 ZRC Philippines 463 9.415 Sea 129 123.82166 34 144 ZRC 2013.1231 1 1 Bohol paratype 2005-05-26 ZRC Philippines 446 9.005 Sea 129 123.43 34 144 ZRC 2013.1241 1 1 Bohol paratype 2005-05-30 ZRC Philippines 278 9.491667 Sea 129 123.71 Balicasag Island 34 144 ZRC 2013.1236 1 1 Bohol paratype 2005-05-22 ZRC Philippines 271 9.648334 Sea 129 123.78167 34 144 ZRC 2013.1236 1 1 Bohol paratype 2005-05-31 ZRC Philippines 538 9.611667 Sea 129 123.715 34 144 ZRC 2013.1230 1 1 Bohol paratype 2005-06-01 ZRC Panglao Philippines 420 9.65 Maribojoc Bay 129 123.76833 34 144 ZRC 2013.1234 1 1 paratype 2007-06-02 MNHN Philippines Luzon Island 34 144 CC 2743 1 1 Aurora paratype 1980-11-27 RV Philippines 215 13.983334 Coriolis 1289 120.3 34 144 1 1 paratype 1909-03-16 RV Philippines Albatross 34 144 USNM 48508 1 1 Southern Leyte paratype 1909-08-19 RV Non-type & Philippines Philippines Albatross 34 144 USNM 48509 1 paratype 2001-10-06 RV Solomon Islands 324 -10.466666 Alis 1298 162.0 34 144 1 1 paratype