The crassipalpis species group of the genus Geodromicus Redtenbacher, 1857 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini) Shavrin, Alexey V. Zootaxa 2019 2019-10-17 4686 4 571 580 8Y3ZQ Shavrin Shavrin 2019 [151,578,837,864] Insecta Staphylinidae Geodromicus Animalia Coleoptera 4 575 Arthropoda species amplissimus sp. nov.  ( Figs. 2, 8–9, 12)   Type material examined:  Holotype ♂: ‘ CHINA(W Sichuan) (4) | Daxue Shan| river valley 5 kmE | Kanding  2500–2800m| 30.03 N/ 102.00 E| 20./  23.V.1997[D.] Wrase’<rectangular label, printed>, ‘ Museumfür Naturkunde| Berlin| Sammlung M. Schülke’<rectangular label, printed>, ‘  HOLOTYPE| Geodromicus| amplissimus sp.n.| Shavrin A.V.des. 2019’ <red rectangulat label, printed> ( CSC).   Paratypes( 56 specimens): 3 ♂♂[two specimens dissected],  2 ♀♀: same data as the holotype ( 1 ♂, 1 ♀: CS; 2 ♂♂, 1 ♀: CSC); 1 ♂[dissected],  1 ♀: ‘ CHINA: Sichuan| Daxue Shan mts., 25 kmSE | Kangding,  3200–3500 m, |  13.VI.–4.VII.2009, | leg. A. Plutenko’<rectangular label, printed>, ‘ Museumfür Naturkunde| Berlin| Sammlung M. Schülke’<rectangular label, printed> ( 1 ♂: CS; 1 ♀: CSC);  1 ♀: ‘ CHINA: Sichuan| Daxue Shan Mts., 25 kmSE | Kangding town,  13. VI.–4.VII.2009| leg. A. Plutenko’<rectangular label, printed> ( CA);  6 ♂♂[two specimens dissected]: ‘ CHINA: W-Sichuan (13) | Daxue Shan, Hailuogou| Glacier Park, Camp1.  2100m| 29.–  31.05.1997, M. Schülke’<rectangular label, printed>, ‘ Museumfür Naturkunde| Berlin| Sammlung M. Schülke’<rectangu- lar label, printed> ( 2 ♂♂: CS; 4 ♂♂: CSC);  2 ♂♂: ‘ CHINA: W-Sichuan (15) | Daxue Shan, Hailuogou Glacier| Park, Camp2, 2550–  2700 m| 29.35.16N, 102.01.53E | 30./  31.05.1997, M. Schülke’<rectangular label, printed>, ‘ Museumfür Naturkunde| Berlin| Sammlung M. Schülke’<rectangular label, printed> ( CSC); 1 ♂[dissected],  3 ♀♀: ‘ CHINA Sichuan, Gongga| Shan, Hailuogou, head of | Glacier1, 2850m  9.VII.[19]96 | 29°35N 102°00EC57’ <rectangular label, printed>, ‘collected by | A. Smetana, J. Farkač| and P. Kabátek’( 1 ♂: CS; 3 ♀♀: NSMT);  1 ♀: ‘ CHINA Sichuan, Gongga| Shan, Hailuogou, above | Camp3, 3050 m  6.VII.[19]96 | 29°35N 102°00EC52’ <rect- angular label, printed> ( NSMT);  1 ♂[without right apical antennomere and left antennomeres 5–11]: ‘ CHINA: Sichuan Gongga| Shan, Hailuogou, in front | of Glacier1, 2850 m | 29°35N 102°00E,  7.VII.| 1998, A. Smetana[C76]’ <rectangular label, printed>, ‘1998 Chinese Expedition| J. Farkač, D. Král, | J. Schneider| & A. Smetana’<rectangular label, printed> ( NSMT); 23 ♂♂( 4 specimensdissected),  11 ♀♀: ‘ CHINA(N-Yunnan) Diqing Tibet| Aut. Pref., Deqin Co., Meili Xue| Shan, E-side,  14 kmW Deqin| 28°27.47’N, 98°46.35’E, 2700 m’ <rectangular label, printed>, ‘small creek valley, glacier outlet | (under stones and gravel along | glacier stream) |  11.VI.2005 D.W.Wrase[09A]’ <rectangular label, printed>, ‘ Museumfür Naturkunde| Berlin| Sammlung M. Schülke’<rect- angular label, printed> ( 6 ♂♂, 2 ♀♀: CS; 17 ♂♂, 9 ♀♀: CSC). All paratypeswith additional red rectangular printed label: ‘ PARATYPE| Geodromicus| amplissimus sp.n.| Shavrin A.V. des. 2019’.   Description.Measurements (n=57): HW: 1.14–1.45; HL: 0.60–0.87; OL: 0.30–0.32; TL: 0.13–0.16; AL ( holotype): 4.05; PL: 0.91–1.25; PWmax: 1.51–2.00; PWmin: 1.37–1.64; ESL: 1.92–2.56; EW: 2.41–2.82; MTbL ( holotype): 1.85; MTrL ( holotype): 0.80 (MTrL 1–4: 0.50; MTrL 5: 0.30); AW: 2.26–2.75; AedL: 1.20–1.50; BL: 5.55–7.50 ( holotype: 7.00). Body reddish-brown to black; mouthparts, antennomeres and legs yellow-brown; apex of apical segment of maxillary palpi, basal portions of antennomeres 1–6, apex of tibia and tarsi yellow. Head with variable, irregular transverse meshes between antennal insertions, fine isodiametric microsculpture on infraorbital ridges, without or with indistinct, irregular transverse microreticulation in interocellar depression; pronotum with very indistinct or distinct, fine, isodiametric or transverse meshes except for median and postero-median portions. Habitus as in Fig. 2. Head 1.6–1.9 times as wide as long, distinctly more convex between ocelli and eyes; anterio-median depression moderately deep and wide, markedly narrowed basad and extending toward wide interocellar depression; temples slightly less than half as long as longitudinal length of eyes. Ocelli distinct; distance between ocelli as long as distance between ocellus and posterior margin of eye, or markedly longer. Punctation fine, irregular, and smooth, markedly denser on infraorbital ridges and sparser in interocellar depression. Apical maxillary palpomere 1.7 times as long as preapical segment. Antennae extending beyond apical fourth of elytra when reclined; lengths × widths of antennomeres ( holotype): 1: 0.42 × 0.15; 2: 0.27 × 0.10; 3: 0.37 × 0.10; 4: 0.35 × 0.12; 5–7: 0.37 × 0.12; 8–9: 0.35 × 0.12; 10: 0.33 × 0.12; 11: 0.50 × 0.12. Pronotum markedly convex, 1.6 times as wide as long, 1.3 times as wide as head, widest in apical third, with rounded anterior angles, distinctly protruded apicad, narrow posterior portion markedly longer than in  G. crassipalpis; median portion without impressions; laterobasal portions distinctly impressed and slightly reflexed. Punctation moderately dense and deep, sparser and finer in anteromedian third, markedly denser laterally, usually with median impunctate area. Elytra slightly to significantly widened posteriad; hind margins straight or rounded. Punctation similar to that of pronotum, but slightly sparser. Metatarsus more than twice as long as metatibia. Abdomen with two distinct transverse tomentose spots in the middle of tergite IV. Male. Posterior margin of tergite VIII straight. Posterior margin of sternite VIII slightly sinuate. Aedeagus ( Fig. 8) with moderately wide median lobe, in apical portion sharply narrowed toward narrowly rounded apex; parameres slightly widened apically, distinctly extending beyond apex of median lobe, with four short apical setae and additional very short setae along entire length of medial edge; internal sac with long narrow flagellum. Lateral aspect of aedeagus as in Fig. 9. Female. Posterior margin tergite VIII straight. Posterior margin of sternite VIII rounded.  Comparative notes.Based on the general shape of the aedeagus,  G. amplissimusShavrin,  sp.n.is most similar to the Himalayan  G. spadiceus, from which it differs by darker elytra, and slightly wider apical portions of median lobe and parameres. The new species differs from all the species of the  crassipalpisgroup by the presence of very short setae on the medial edges of the parameres.   Etymology.The specific epithet is the superlative of the Latin adjective amplus(wide). It refers to the wide pronotum and elytra.   Distribution.The species is know from several locations in Daxue Shanand Gongga Shanranges in Sichuan, and from Meili Xue Shanrange in northern Yunnan( Fig. 12).  Natural History.The specimens were collected from gravel on the banks of a river coming out from Glacier 1 (locality: C57), from flooded gravel banks of a river close to the glacier (locality: C76), by sifting wet moss on large fallen trees in a primary  Abiesforest with rich undergrowth of rhododendron(locality: C52), by sifting dead wood with fungi along a small river in a broadleaved forest with a small clearings and old trees (locality: 13), by sifting wet substrates between fallen trees and dead wood in an old mixed forest with huge old broadleaved and coniferous trees (locality: 15), and in Meili Xue Shanfrom under stones and gravel along a glacier stream (locality: 09A). The elevations range from 2100 to 3500 ma.s.l. 2427264220 1997-05-23 Wrase' & Museum & Naturkunde & Berlin & Sammlung M. Schulke' China 2650 30.03 Kanding 734 102.0 Daxue Shan 4 575 1 1 Sichuan holotype 2427264225 CSC Shavrin A. V. China 4 575 1 Sichuan holotype 2427264233 CS, CSC Shavrin A. V. China 4 575 7 4 3 Sichuan paratype 2427264224 2009-06-13 2009-07-04 2009-06-13 CS, CSC A. Plutenko' & Museum & Naturkunde & Berlin & Sammlung M. Schulke' China 3350 Kangding Daxue Shan mts. 4 575 3 2 1 Sichuan paratype 2427264227 2009-06-13 2009-07-04 2009-06-13 VI, CA A. Plutenko' China Kangding town Daxue Shan Mts. 4 575 1 1 Sichuan paratype 2427264221 1997-05-31 CS, CSC M. Schulke' & Museum & Naturkunde & Berlin & Sammlung M. Schulke' China Daxue Shan 2100 Glacier Park Hailuogou 4 575 12 12 paratype 2427264228 1997-05-31 CSC M. Schulke' & Museum & Naturkunde & Berlin & Sammlung M. Schulke' China Daxue Shan 2700 Park Hailuogou Glacier 4 575 2 2 paratype 2427264219 9. VII. CS, NSMT A. Smetana & J. Farkac & P. Kabatek' China Gongga 29.583334 Hailuogou 1226 102.0 Shan 4 575 7 6 1 Sichuan paratype 2427264226 6. VII. NSMT China Gongga 29.583334 Hailuogou 1226 102.0 Shan 4 575 1 1 Sichuan paratype 2427264222 7. VII. NSMT A. Smetana & Chinese Expedition & J. Farkac & D. Kral & J. Schneider & A. Smetana' China Gongga 29.583334 Hailuogou 1226 102.0 Shan 4 575 1 1 Sichuan paratype 2427264229 2005-06-11 CS, CSC D. W. Wrase & Museum & Naturkunde & Berlin & Sammlung M. Schulke' & All China Deqin Co. 28.457834 Aut. Pref. 12 98.7725 Diqing 4 575 45 22 23 Xizang paratype